1. Family: Boraginaceae Juss.
    1. Genus: Cordia L.
      1. Cordia subcordata Lam.

        This species is accepted, and its native range is S. Somalia to N. Mozambique, W. Indian Ocean to Pacific.

    [FZ]

    Boraginaceae, E. S. Martins (Trichodesma by R. K. Brummit). Flora Zambesiaca 7:4. 1990

    Habit
    Shrub up to 4 m. high (or a tree up to 15 m. high outside the Flora Zambesiaca area); branches angular when young, glabrous or subglabrous.
    Leaves
    Leaves alternate; petiole 2.5–7.0 cm. long, subglabrous; lamina 6–20 x 5–16 cm., ovate or widely ovate to circular, glabrous or with some minute appressed not bulbous-based hairs, older ones frequently dotted with groups of cystoliths above, more or less pubescent along the midrib and some secondary nerves and usually minutely tomentose in the axils of the principal secondary and tertiary nerves below, obtuse to rounded and often shortly acuminate at apex, obtuse to truncate, rarely subcordate at the base, with entire or slightly repand margins, papyraceous, with 6 pairs of secondary nerves.
    Inflorescences
    Inflorescence cymose, loosely branched, short, 6–20-flowered, terminal, often apparently opposite-leaved.
    Flowers
    Flowers hermaphrodite, heterostylous, on pedicels 2–8 mm. long joined near the apex.
    Calyx
    Calyx 12–15 mm. long and 4–5 mm. in diam. at the mouth, cylindric or slightly conical, coriaceous, usually with 3–5 rounded teeth up to 4 mm. long, glabrous or very sparsely strigose outside and inside but densely strigose inside on the teeth.
    Corolla
    Corolla 30–45 mm. long, infundibuliform, orange, glabrous or nearly so; limb nearly as broad as the length of the corolla; lobes 5–7, broad, rounded, sometimes with crenate margin, spreading.
    Stamens
    Stamens inserted at c. 20 mm. from the base of the corolla tube; filaments in the short-styled flowers c. 8 mm. long, in the long-styled flowers 3–5 mm. long, glabrous; anthers 2.5–3.5 mm. long.
    Pistil
    Ovary c. 3 mm. long, conical, glabrous; style in the short-styled flowers c. 22 mm. long, first-branched at c. 17 mm. and with stigmatic branches c. 1.5 mm. long, in the long-styled flowers c. 30 mm. long, first-branched at 21–23 mm. and with stigmatic branches 2.5–3.0 mm. long, flattened.
    Fruits
    Fruit a nut, c. 25 x 22 mm., subglobose, smooth, shining, completely enclosed in the enlarged urceolate calyx; mesocarp suberose; pyrene angular, much ridged and scabrous, containing 1–2 seeds.
    [FTEA]

    Boraginaceae, B. Verdcourt. Flora of Tropical East Africa. 1991

    Habit
    Shrub or tree, 1.5–15 m. tall with spreading branches; trunk thick, up to 60 cm. diameter; stems pale grey, ± ridged; bark brown, ± rough.
    Leaves
    Leaf-blades broadly ovate to almost round, 4–34 cm. long, 3–17 cm. wide, rounded to acute or subacute or shortly acuminate at the apex, rounded at the base or sometimes ± subcordate or obtuse or quite tapering, slightly succulent, very shortly pubescent above when young but later ± glabrous above but often with cystolith spots, velvety to ± tomentose along edges of costa and nervation beneath or ± glabrous; petiole 1.5–11 cm. long.
    Inflorescences
    Inflorescence axillary or pseudo-terminal 6–20-flowered cymes, 10–14 cm. long, pubescent when young; peduncles 0.5–2 cm. long; pedicels 2–5(–10) mm. long; flowers heterostylous.
    Calyx
    Calyx brownish, cylindric-funnel-shaped, 1.1–2 cm. long, 3–5-valvate or -lobed, the lobes 2–5 mm. long, finally accrescent and completely enveloping the fruit, glabrous or slightly pubescent outside.
    Corolla
    Corolla white or usually apricot, orange or red, funnel-shaped; tube 2–3 cm. long, 3 mm. diameter at base, 1–2 cm. wide at the throat; lobes 5–7, rounded to obovate-spathulate, 1.5–2.5 cm. long and wide, spreading, usually crinkly-edged.
    Filaments
    Filaments inserted just below the middle of the corolla-tube, the anthers level with the throat.
    Style
    Style 2–2.5 cm. long (short) or ± 3 cm. long (long); primary branches 2–7 mm. long, the 4 stigmatic lobes 0.5–2 mm. long, flattened.
    Fruits
    Fruit yellow to red, becoming blackish, ovoid, obovoid or subglobose, 1.5–4 cm. long, 2–3 cm. wide, sometimes ± beaked, the enveloping calyx becoming hard; endocarp very hard, angular and furrowed, 4-locular but only 2 seeds developing.
    Figures
    Fig. 1/1,2, p. 8.
    Habitat
    Shore above high tide mark; inland fringe of mangrove swamps; littoral savanna; 0–1.5 (–75)m.
    Distribution
    K7 P T3 T6 T8 Z

    Images

    Distribution

    Native to:

    Aldabra, Andaman Is., Bangladesh, Bismarck Archipelago, Cambodia, Chagos Archipelago, Christmas I., Cocos (Keeling) Is., Comoros, Cook Is., Fiji, Gilbert Is., Hainan, Howland-Baker Is., India, Kenya, Laccadive Is., Lesser Sunda Is., Line Is., Madagascar, Malaya, Maldives, Mozambique, Mozambique Channel I, Nauru, New Caledonia, New Guinea, Nicobar Is., Niue, Northern Territory, Phoenix Is., Pitcairn Is., Queensland, Samoa, Seychelles, Solomon Is., Somalia, South China Sea, Tanzania, Thailand, Tokelau-Manihiki, Tonga, Tuvalu, Vanuatu, Vietnam, Wake I., Wallis-Futuna Is., Western Australia

    Introduced into:

    Mauritius, Society Is.

    Cordia subcordata Lam. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Jul 1, 2013 I Made Maduarta (Pung) [IMM 185], Lesser Sunda Is. K000734531
    Jan 1, 1988 Forster, G. [s.n.], Society Is. K000998028
    Drummond, R.B. [3988], Kenya 16376.000
    Jeffrey, C. [533], Seychelles 22734.000
    Hunt, P.F. [RSS 2770], Solomon Is. 6049.185
    Renvoize, S.A. [736], Seychelles 7317.000
    Wood, D., Seychelles 38053.000
    Wood, D., Seychelles 38054.000
    Wheatley, J.I. [578], Vanuatu 54987.000
    Forster, G. [s.n.], Society Is. K000998029
    s.coll. [Cat. no. 9063] K001132158
    s.coll. [Cat. no. 9063] K001132159
    s.coll. [Cat. no. 9063] K001132160
    s.coll. [Cat. no. 9063] K001132161
    s.coll. [Cat. no. 9063] K001132162
    s.coll. [Cat. no. 9063] K001132163
    s.coll. [Cat. no. 9063] K001132164
    s.coll. [Cat. no. 9063] K001132165
    s.coll. [Cat. no. 9063], Thailand K001132166

    First published in Tabl. Encycl.: n.º 1899 (1792)

    Accepted by

    • Girmansyah, D. & al. (eds.) (2013). Flora of Bali an annotated checklist: 1-158. Herbarium Bogorensis, Indonesia.
    • Mostaph, M.K. & Uddin, S.B. (2013). Dictionary of plant names of Bangladesh, Vasc. Pl.: 1-434. Janokalyan Prokashani, Chittagong, Bangladesh.
    • Gardner, R.O. (2011). Trees and shrubs of Niue: an identification guide to the island's indigenous and naturalised woody plants: 1-244. R.O. Gardner, Auckland, New Zealand.
    • Suzuki, M., Taufatofua, P. & Koyama, T. (2007). New records of plants from Tonga Makinoa, n.s., 6: 25-62.
    • Thulin, M. (ed.) (2006). Flora of Somalia 3: 1-626. The Royal Botanic Gardens, Kew.
    • Autrey, J.C., Bosser, J. & Ferguson, I.K. (eds.) (2005). Flore des Mascareignes 121-126: 1. IRD Éditions, MSIRI, RBG-Kew, Paris.
    • Turner, I.M., Xing, F. & Corlett, R.T. (2000). An annotated check-list of the vascular plants of the South China Sea and its shores The Raffles bulletin of Zoology, Suppl. 8: 23-116.
    • Govaerts, R. (1999). World Checklist of Seed Plants 3(1, 2a & 2b): 1-1532. MIM, Deurne.
    • Fosberg, F.R. & Stoddard, D.R. (1994). Flora of the Phoenix Islands, Central Pacific Atoll Research Bulletin 393: 1-60.
    • Thaman, R.R., Fosberg, F.R., Manner, H.I. & Hassall, D.C. (1994). The Flora of Nauru Atoll Research Bulletin 392: 1-223.
    • George, A.S., Orchard, A.E. & Hewson, H.J. (eds.) (1993). Oceanic islands 2 Flora of Australia 50: 1-606. Australian Government Publishing Service, Canberra.
    • Verdcourt, B. (1991). Flora of Tropical East Africa, Boraginaceae: 1-124.
    • Launert, E. & Pope, G.V. (eds.) (1990). Flora Zambesiaca 7(4): 1-114. Royal Botanic Gardens, Kew.
    • Hancock, I.R. & Henderson, C.P. (1988). Flora of the Solomon Islands Research Bulletin Dodo Creek Research Station 7: 1-203.
    • Fosberg, F.R. & Sachet, M.-H. (1987). Flora of the Gilbert Island, Kiribati, Checklist Atoll Research Bulletin 295: 1-33.
    • Woodroffe, C.D. (1985). Vegetation and flora of Nui Atoll, Tuvalu Atoll Research Bulletin 283: 1-18.
    • Banks, R.C. (ed.) (1982). Wildlife and wildlife habitat of American Samoa 2: 1-151. U.S. Fisch and Wildlife services, Washington.
    • Heine, H. (1976). Flore de la Nouvelle-Calédonie et Dépendances 7: 1-212. Muséum National d'Histoire Naturelle, Paris.
    • Fosberg, F.R. (1975). Vascular plants of Aitutaki Atoll Research Bulletin 190: 73-84.
    • Parham, B.E.V. (1971). The Vegetation of the Tokelau Islands with special reference to the plants of Nukunonu Atoll New Zealand Journal of Botany 9: 576-609.

    Literature

    Kew Backbone Distributions
    • Girmansyah, D. & al. (eds.) (2013). Flora of Bali an annotated checklist: 1-158. Herbarium Bogorensis, Indonesia.
    • Mostaph, M.K. & Uddin, S.B. (2013). Dictionary of plant names of Bangladesh, Vasc. Pl.: 1-434. Janokalyan Prokashani, Chittagong, Bangladesh.
    • Gardner, R.O. (2011). Trees and shrubs of Niue: an identification guide to the island's indigenous and naturalised woody plants: 1-244. R.O. Gardner, Auckland, New Zealand.
    • Pandey, R.P. & Dilwakar, P.G. (2008). An integrated check-list flora of Andaman and Nicobar islands, India Journal of Economic and Taxonomic Botany 32: 403-500.
    • Autrey, J.C., Bosser, J. & Ferguson, I.K. (eds.) (2005). Flore des Mascareignes 121-126: 1. IRD Éditions, MSIRI, RBG-Kew, Paris.
    • Sheppard, C.R.C. & Seaward, M.R.D. (eds.) (1999). Ecology of the Chagos archipelago: 1-350. Westbury Academic & Scientific Publishing, Otley.
    • Ananda Rao, T. & Ellis, J.L. (1995). Flora of Lakshadweep islands off the Malabar coast, peninsular India, with emphasis on phytogeographical distribution of plants Journal of Economic and Taxonomic Botany 19: 235-250.
    • Fosberg, F.R. & Stoddard, D.R. (1994). Flora of the Phoenix Islands, Central Pacific Atoll Research Bulletin 393: 1-60.
    • Thaman, R.R., Fosberg, F.R., Manner, H.I. & Hassall, D.C. (1994). The Flora of Nauru Atoll Research Bulletin 392: 1-223.
    • George, A.S., Orchard, A.E. & Hewson, H.J. (eds.) (1993). Oceanic islands 2 Flora of Australia 50: 1-606. Australian Government Publishing Service, Canberra.
    • Hancock, I.R. & Henderson, C.P. (1988). Flora of the Solomon Islands Research Bulletin Dodo Creek Research Station 7: 1-203.
    • Fosberg, F.R. & Sachet, M.-H. (1987). Flora of the Gilbert Island, Kiribati, Checklist Atoll Research Bulletin 295: 1-33.
    • Woodroffe, C.D. (1985). Vegetation and flora of Nui Atoll, Tuvalu Atoll Research Bulletin 283: 1-18.
    • Morat, P. & Veillon, J.-M. (1985). Contributions à la conaissance de la végétation et de la flore de Wallis et Futuna Bulletin du Muséum National d'Histoire Naturelle. Section B, Adansonia 7: 259-329.
    • Fosberg, F.R. (1975). Vascular plants of Aitutaki Atoll Research Bulletin 190: 73-84.
    • Parham, B.E.V. (1971). The Vegetation of the Tokelau Islands with special reference to the plants of Nukunonu Atoll New Zealand Journal of Botany 9: 576-609.
    • Fosberg, F.R. (1957). The Maldive islands, Indian Ocean Atoll Research Bulletin 58: 1-37.
    • Christopherson, E. (1935). Flowering Plants of Samoa Bernice P. Bishop Museum Bulletin 128: 1-221.
    Flora of Tropical East Africa
    • Martins in Flora Zambesiaca 7(4): 62 (1990).
    • Nowicke, Miller & Bittner in Journ. Palynol. 23–24: 59–64 (1988).
    • Warfa, Acta Univ. Upsal. 174: 51, fig. 2/C, 4/B (1988).
    • Fosberg & Renvoize, Fl. Aldabra: 194, fig. 31/1–3 (1980).
    • Heine, Fl. Nouv.-Caléd. 7: 104, t. 23 (1976).
    • Cufod., Enumeratio Plantarum Aethiopiae Spermatophyta (Supplement in Bull. Jard. Bot. Brux.) p. 769 (1961).
    • Dale & Greenway, Kenya Trees and Shrubs p. 71 (1961).
    • I.M. Johnston in Journ. Arn. Arb. 32: 3 (1951).
    • J.P.M. Brenan, Check-lists of the Forest Trees and Shrubs of the British Empire no. 5, part II, Tanganyika Territory p. 76 (1949).
    • R. O. Williams, Useful and Ornamental Plants in Zanzibar and Pemba p. 212 (1949).
    • Bak. & Wright in Flora of Tropical Africa 4(2): 10 (1905).
    • C.B.Cl. in Fl. Brit. Ind. 4: 140 (1883).
    • Vatke in Linnaea 43: 314 (1882).
    • A.DC. in DC., Prodr. 9: 477 (1845).
    • Poir. in Encycl. Méth. Bot. 7: 41 (1806).
    • Lam., Tab. Encycl. 1: 421 (1792).

    Sources

    Art and Illustrations in Digifolia
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2018. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2018. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0