1. Family: Poaceae Barnhart
    1. Genus: Bothriochloa Kuntze
      1. Bothriochloa ischaemum (L.) Keng

        This species is accepted, and its native range is Temp. Eurasia, NW. Africa.

    [GB]
    Habit
    Perennial; caespitose. Rootstock evident. Culms erect, or geniculately ascending; 40-80 cm long. Leaves mostly basal. Ligule a ciliolate membrane. Leaf-blades 3-15 cm long; 2-4 mm wide; glaucous. Leaf-blade surface scaberulous. Leaf-blade margins tuberculate-ciliate; hairy at base.
    Inflorescences
    Inflorescence composed of racemes. Racemes 5-15; digitate; 4-6 cm long. Rhachis fragile at the nodes; flattened; villous on margins. Rhachis internodes linear. Rhachis internode tip transverse. Spikelets in pairs. Fertile spikelets sessile; 1 in the cluster. Companion sterile spikelets pedicelled; 1 in the cluster. Pedicels linear; flattened; with a translucent median line (also present in internodes); villous.
    Sterile
    Companion sterile spikelets well-developed; containing empty lemmas, or male; lanceolate; 3.5-5 mm long; as long as fertile. Companion sterile spikelet glumes without depressions; glabrous. Companion sterile spikelet lemmas enclosed by glumes.
    Spikelets
    Spikelets comprising 1 basal sterile florets; 1 fertile florets; without rhachilla extension. Spikelets lanceolate, or oblong; dorsally compressed; 3.5-5 mm long; falling entire; deciduous with accessory branch structures. Spikelet callus pilose; base obtuse; attached transversely. Companion sterile spikelets well-developed; containing empty lemmas, or male; lanceolate; 3.5-5 mm long; as long as fertile. Companion sterile spikelet glumes without depressions; glabrous. Companion sterile spikelet lemmas enclosed by glumes.
    Fertile
    Spikelets comprising 1 basal sterile florets; 1 fertile florets; without rhachilla extension. Spikelets lanceolate, or oblong; dorsally compressed; 3.5-5 mm long; falling entire; deciduous with accessory branch structures. Spikelet callus pilose; base obtuse; attached transversely.
    Glume
    Glumes dissimilar; exceeding apex of florets; firmer than fertile lemma. Lower glume elliptic; 1 length of spikelet; chartaceous; without keels; keel-less except near apex. Lower glume surface flat; without pits; pilose; hairy below. Lower glume apex acute. Upper glume lanceolate; 1-keeled.
    Florets
    Basal sterile florets barren; without significant palea. Lemma of lower sterile floret oblong; hyaline. Fertile lemma linear; hyaline; without keel. Lemma apex entire; awned; 1 -awned. Principal lemma awn apical; geniculate; 12-15 mm long overall; with twisted column. Column of lemma awn glabrous. Palea absent or minute.
    Flowers
    Anthers 3.
    Distribution
    Europe: central, southwestern, southeastern, and eastern. Africa: north and Macaronesia. Asia-temperate: Siberia, Soviet Middle Asia, Caucasus, western Asia, China, and eastern Asia. Asia-tropical: India, Malesia, and Papuasia. Pacific: south-central. North America: north-central USA, south-central USA, southeast USA, and Mexico. South America: Caribbean, northern South America, and western South America.
    Reference
    Andropogoneae. Fl Pak 1993.

    Images

    Distribution

    Native to:

    Afghanistan, Algeria, Austria, Bulgaria, China North-Central, China South-Central, China Southeast, Corse, Czechoslovakia, East Himalaya, France, Germany, Greece, Hainan, Hungary, India, Inner Mongolia, Iran, Iraq, Italy, Kazakhstan, Kirgizstan, Korea, Krym, Lebanon-Syria, Manchuria, Mongolia, Morocco, Nepal, North Caucasus, Oman, Pakistan, Poland, Portugal, Qinghai, Romania, South European Russi, Spain, Switzerland, Tadzhikistan, Taiwan, Thailand, Tibet, Transcaucasus, Tunisia, Turkey, Turkey-in-Europe, Turkmenistan, Ukraine, Uzbekistan, West Himalaya, Western Sahara, Xinjiang, Yugoslavia

    Introduced into:

    Alabama, Argentina Northeast, Arizona, Arkansas, Aruba, Bolivia, Brazil Northeast, Brazil Southeast, California, Canary Is., Colombia, Colorado, Costa Rica, Cuba, Dominican Republic, Easter Is., Florida, Haiti, Jamaica, Jawa, Kansas, Leeward Is., Lesser Sunda Is., Louisiana, Mexico Central, Mexico Northeast, Mexico Northwest, Mexico Southeast, Mexico Southwest, Mississippi, Netherlands Antilles, Netherlands Antilles, New Guinea, New Mexico, New York, Ogasawara-shoto, Ogasawara-shoto, Oklahoma, Philippines, Puerto Rico, South Carolina, Tennessee, Texas, Utah, Venezuela

    Bothriochloa ischaemum (L.) Keng appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Schrenk, A.G. von [s.n.] K001057432

    First published in Contr. Biol. Lab. Sci. Soc. China, Bot. Ser. 10: 201 (1936)

    Accepted by

    • Chang, C.S., Kim, H. & Chang, K.S. (2014). Provisional checklist of vascular plants for the Korea peninsula flora (KPF): 1-660. DESIGNPOST.
    • Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies Smithsonian Contributions to Botany 98: 1-1192.
    • Verloove, Fl. & Reyes-Betancort, J.A. (2011). Additions to the flora of Tenerife (Canary islands, Spain) Collectanea Botanica 30: 63-78.
    • Giraldo-Cañas, D. (2011). Catálogo de la familia Poaceae en Colombia Darwiniana 49: 139-247.
    • Clayton, W.D. & Snow, N. (2010). A key to Pacific Grasses: 1-107. Kew Publishing, Royal Botanic Gardens, Kew.
    • Kandwal, M.K. & Gupta, B.K. (2009). An update on grass flora of Uttarkhand Indian Journal of Forestry 32: 657-668.
    • Hokche, O., Berry, P.E. & Huber, O. (eds.) (2008). Nuevo Catálogo de la Flora Vascular de Venezuela: 1-859. Fundación Instituto Botánico de Venezuela.
    • Wu, Z. & Raven, P.H. (eds.) (2006). Poaceae Flora of China 22: 1-733. Missouri Botanical Garden Press, St. Louis.
    • Clayton, W.D., Harman, K.T. & Williamson, H. (2006). World Grass Species - Synonymy database The Board of Trustees of the Royal Botanic Gardens, Kew.
    • Dávila, P., Mejia-Saulés, M.T., Gómez-Sánchez, N., Valdés-Reyna, J., Ortíz, J.J., Morín, C., Castrejón, J. & Ocampo, A. (2006). Catálogo de las Gramíneas de México: 1-671. CONABIO, México city.
    • Takhtajan, A.L. (ed.) (2006). Conspectus Florae Caucasi 2: 1-466. Editio Universitatis Petropolitanae.
    • Hammel, B.E. & al. (2003). Manual de Plantas de Costa Rica 3: 1-884. Missouri Botanical Garden Press, St. Louis.
    • Flora of North America Editorial Committee (2003). Flora of North America North of Mexico 25: 1-781. Oxford University Press, New York, Oxford.
    • Noltie, H.J. (2000). Flora of Bhutan 3(2): 457-883. Royal Botanic Gardens, Edinburgh.
    • Press, J.R. et al. (2000). Annotated Checklist of the Flowering Plants of Nepal: i-x, 1-430. Natural History Museum, London.
    • Fedorov, A.A. (ed.) (1999). Flora of Russia. The European part and bordering regions 1: 1-546. A.A. Balkema, Rotterdam, Broekfield.
    • Renvoize, S.A. (1998). Gramineas de Bolivia: 1-644. The Royal Botanic Gardens, Kew.
    • Lee, W.T. (1996). Lineamenta Florae Koreae: 1-1688. Soul T'ukpyolsi: Ak'ademi Sojok.
    • Ghazanfar, S.A. (1992). An Annotated Catalogue of the Vascular Plants of Oman and their Vernacular names Scripta Botanica Belgica 2: 1-153.
    • Karthikeyan, S., Jain, S.K., Nayar, M.P. & Sanjappa, M. (1989). Florae Indicae Enumeratio: Monocotyledonae: 1-435. Botanical Survey of India, Calcutta.
    • Kobayashi, S. & Ono, M. (1987). A Revised List of Vascular Plants Indigenous and Introduced to the Bonin (Ogasawara) and the Volcano (Kazan) Islands Ogasawara Research 13: 1-55.
    • Koyama, T. (1987). Grasses of Japan and its neighboring regions: an identification manual: 1-570. Kodansha, Tokyo, Japan.
    • Davis, P.H. (ed.) (1985). Flora of Turkey and the East Aegean Islands 9: 1-724. Edinburgh University Press, Edinburgh.
    • Cope, T.A. (1982). Flora of Pakistan 143: 1-678. Department of Botany, University of Karachi, Karachi.
    • Bor, N.L. (1970). Flora Iranica 70: 1-573. Naturhistorisches Museums Wien.
    • Henty, E.E. (1969). A manual of the grasses of New Guinea: 1-215. Division of Botany, Dpt. of Forests, LAE, New Guinea.
    • Bor, N.L. (1968). Flora of Iraq 9: 1-588. Ministry of Agriculture & Agrarian Reform, Baghdad.

    Literature

    Kew Backbone Distributions
    • Forzza, R.C. & al. (2013). Lista de Espécies da Flora do Brasil http://floradobrasil.jbrj.gov.br/2010/.
    • Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies Smithsonian Contributions to Botany 98: 1-1192.
    • Verloove, Fl. & Reyes-Betancort, J.A. (2011). Additions to the flora of Tenerife (Canary islands, Spain) Collectanea Botanica 30: 63-78.
    • Giraldo-Cañas, D. (2011). Catálogo de la familia Poaceae en Colombia Darwiniana 49: 139-247.
    • Dobignard, D. & Chatelain, C. (2010). Index synonymique de la flore d'Afrique du nord 1: 1-455. Éditions des conservatoire et jardin botaniques, Genève.
    • Clayton, W.D. & Snow, N. (2010). A key to Pacific Grasses: 1-107. Kew Publishing, Royal Botanic Gardens, Kew.
    • Kandwal, M.K. & Gupta, B.K. (2009). An update on grass flora of Uttarkhand Indian Journal of Forestry 32: 657-668.
    • Hokche, O., Berry, P.E. & Huber, O. (eds.) (2008). Nuevo Catálogo de la Flora Vascular de Venezuela: 1-859. Fundación Instituto Botánico de Venezuela.
    • Wu, Z. & Raven, P.H. (eds.) (2006). Poaceae Flora of China 22: 1-733. Missouri Botanical Garden Press, St. Louis.
    • Dávila, P., Mejia-Saulés, M.T., Gómez-Sánchez, N., Valdés-Reyna, J., Ortíz, J.J., Morín, C., Castrejón, J. & Ocampo, A. (2006). Catálogo de las Gramíneas de México: 1-671. CONABIO, México city.
    • Takhtajan, A.L. (ed.) (2006). Conspectus Florae Caucasi 2: 1-466. Editio Universitatis Petropolitanae.
    • Hammel, B.E. & al. (2003). Manual de Plantas de Costa Rica 3: 1-884. Missouri Botanical Garden Press, St. Louis.
    • Flora of North America Editorial Committee (2003). Flora of North America North of Mexico 25: 1-781. Oxford University Press, New York, Oxford.
    • Noltie, H.J. (2000). Flora of Bhutan 3(2): 457-883. Royal Botanic Gardens, Edinburgh.
    • Press, J.R. et al. (2000). Annotated Checklist of the Flowering Plants of Nepal: i-x, 1-430. Natural History Museum, London.
    • Renvoize, S.A. (1998). Gramineas de Bolivia: 1-644. The Royal Botanic Gardens, Kew.
    • Lee, W.T. (1996). Lineamenta Florae Koreae: 1-1688. Soul T'ukpyolsi: Ak'ademi Sojok.
    • Ghazanfar, S.A. (1992). An Annotated Catalogue of the Vascular Plants of Oman and their Vernacular names Scripta Botanica Belgica 2: 1-153.
    • Karthikeyan, S., Jain, S.K., Nayar, M.P. & Sanjappa, M. (1989). Florae Indicae Enumeratio: Monocotyledonae: 1-435. Botanical Survey of India, Calcutta.
    • Kobayashi, S. & Ono, M. (1987). A Revised List of Vascular Plants Indigenous and Introduced to the Bonin (Ogasawara) and the Volcano (Kazan) Islands Ogasawara Research 13: 1-55.
    • Davis, P.H. (ed.) (1985). Flora of Turkey and the East Aegean Islands 9: 1-724. Edinburgh University Press, Edinburgh.
    • Tutin, T.G. & al. (eds.) (1980). Flora Europaea 5: 1-452. Cambridge University Press.
    • Bor, N.L. (1970). Flora Iranica 70: 1-573. Naturhistorisches Museums Wien.
    • Henty, E.E. (1969). A manual of the grasses of New Guinea: 1-215. Division of Botany, Dpt. of Forests, LAE, New Guinea.
    • Bor, N.L. (1968). Flora of Iraq 9: 1-588. Ministry of Agriculture & Agrarian Reform, Baghdad.
    • Ovczinnikov, P.N. (ed.) (1957). Flora Tadzhikskoi SSR 1: 1-547. Izd-vo Akademii nauk SSSR, Moskva.
    • Pavlov, N.V. (ed.) (1956). Flora Kazakhstana 1: 1-354. Alma-Ata, Izd-vo Akademii nauk Kazakhskoi SSR.
    • Roshevitz, R.J. & al. (eds.) (1950). Flora Kirgizskoi SSR 2: 1-315. Frunze : Izd-vo KirgizFAN SSSR.
    • Guinea, E. (1948). Catálogo razonado de las plantas del Sáhara Español Anales del Jardin Botanico de Madrid 8: 357-429.
    • Merkodovich, N.A. (ed.) (1941). Flora Uzbekistana 1: 1-566. Izd-va Akademii nauk Uzbekskoi SSR, Tashkent.
    • Fedtschenko, B.A. & al. (1932). Flora Turkmenii 1: 1-340. Turkmenskoe gosudarstvennoe izd., Ashkhabad.

    Sources

    GrassBase - The Online World Grass Flora
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2018. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2018. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0