1. Family: Poaceae Barnhart
    1. Genus: Zoysia Willd.
      1. Zoysia matrella (L.) Merr.

        This species is accepted, and its native range is Tropical & Subtropical Asia to NW. Pacific.

    [GB]
    Habit
    Perennial; mat forming. Rhizomes elongated. Stolons present. Culms prostrate; 5-25 cm long; wiry. Leaf-sheaths longer than adjacent culm internode; glabrous on surface. Ligule a ciliate membrane. Leaf-blades filiform, or linear; flat, or involute; 2-10 cm long; 1-3 mm wide.
    Inflorescences
    Inflorescence composed of racemes. Racemes 1; single; multilateral; 0.6-3.5 cm long. Spikelets ascending; solitary. Fertile spikelets pedicelled. Pedicels 0.3-1 mm long.
    Spikelets
    Spikelets comprising 1 fertile florets; without rhachilla extension. Spikelets lanceolate to ovate; laterally compressed; 2-3.5 mm long; falling entire.
    Fertile
    Spikelets comprising 1 fertile florets; without rhachilla extension. Spikelets lanceolate to ovate; laterally compressed; 2-3.5 mm long; falling entire.
    Glume
    Glumes one the lower absent or obscure; exceeding apex of florets; firmer than fertile lemma; shiny. Upper glume lanceolate to ovate; 1 length of spikelet; coriaceous; without keels; 3 -veined. Upper glume apex acute; muticous, or awned; 1 -awned. Upper glume awn 0-1 mm long.
    Florets
    Fertile lemma oblong; 2-2.5 mm long; hyaline; without keel; 1 -veined. Lemma apex emarginate. Palea 0.5 length of lemma; hyaline; without keels.
    Flowers
    Anthers 3; 1.4 mm long.
    Distribution
    Africa: west tropical and western Indian ocean. Asia-temperate: China and eastern Asia. Asia-tropical: India, Indo-China, Malesia, and Papuasia. Australasia: Australia. Pacific: southwestern, south-central, northwestern, and north-central. North America: Mexico. South America: Mesoamericana, Caribbean, western South America, and Brazil.
    Reference
    Cynodonteae. WDC 1996.

    Images

    Distribution

    Native to:

    Bangladesh, Borneo, Caroline Is., China Southeast, Cocos (Keeling) Is., Hainan, India, Japan, Jawa, Kazan-retto, Korea, Laos, Lesser Sunda Is., Malaya, Maluku, Marianas, Marshall Is., Myanmar, Nansei-shoto, New Guinea, Nicobar Is., Ogasawara-shoto, Philippines, Queensland, Solomon Is., South China Sea, Sri Lanka, Sulawesi, Sumatera, Taiwan, Thailand, Vietnam

    Introduced into:

    Alabama, Bahamas, Benin, Bermuda, Brazil Southeast, Brazil West-Central, Burkina, Cayman Is., Christmas I., Colombia, Costa Rica, Cuba, Dominican Republic, Ecuador, El Salvador, Fiji, Florida, Gabon, Galápagos, Georgia, Ghana, Gulf of Guinea Is., Haiti, Hawaii, Honduras, Ivory Coast, Leeward Is., Louisiana, Madagascar, Maldives, Marquesas, Mauritius, Mexico Northeast, Mexico Southwest, New Mexico, Nicaragua, Nigeria, North Carolina, Puerto Rico, Rodrigues, Réunion, Samoa, Society Is., Togo, Trinidad-Tobago, Tubuai Is., West Himalaya

    Zoysia matrella (L.) Merr. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Mar 24, 2009 Kerr, A.F.G. [10722], Thailand K000648131
    Mar 24, 2009 Kerr, A.F.G. [7945A], Thailand K000648109
    Mar 24, 2009 Sorensen, Th. [2470], Thailand K000648110
    Mar 24, 2009 Wichian [495], Thailand K000648111
    Mar 24, 2009 Ryves, T.B. [KS95/004], Thailand K000648132
    Jun 1, 2000 Bor, N.L. [S.57], Malaysia K000290610 isotype
    Kerr, A.F.G. [4393], Thailand K000621619
    Kerr, A.F.G. [4293], Thailand K000621620
    Kerr, A.F.G. [6981], Thailand K000621621
    Kerr, A.F.G. [7945], Thailand K000621622
    Kerr, A.F.G. [10695], Thailand K000621623
    Kerr, A.F.G. [10722], Thailand K000621624
    Kerr, A.F.G. [10967], Thailand K000621625
    Kerr, A.F.G. [10967], Thailand K000621626
    Kerr, A.F.G. [13464], Thailand K000621628
    Kerr, A.F.G. [13464], Thailand K000621629
    Kerr, A.F.G. [15099], Thailand K000621630
    Kerr, A.F.G. [15099], Thailand K000621631
    Kerr, A.F.G. [19092], Thailand K000621632
    Sorensen, Th. [1710], Thailand K000682688
    Larsen, K. [1279], Thailand K000648130
    Smitinand, T. [8916], Thailand K000682687
    Marcan, A. [2258], Thailand K000621633
    Marcan, A. [2263], Thailand K000621634
    Marcan, A. [2263], Thailand K000621635
    Lima, J.M. [315], Brazil K001101764

    First published in Philipp. J. Sci., C 7: 230 (1912)

    Accepted by

    • Chang, C.S., Kim, H. & Chang, K.S. (2014). Provisional checklist of vascular plants for the Korea peninsula flora (KPF): 1-660. DESIGNPOST.
    • Thiombiano, A., Schmidt, M., Dressler, S., Ouédraogo, A., Hahn, K. & Zizka, G. (2012). Catalogue des plantes vasculaires du Burkina Faso Boissiera 65: 1-391.
    • Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies Smithsonian Contributions to Botany 98: 1-1192.
    • Figueiredo, E., Paiva, J., Stévart, T., Oliveira, F. & Smith, G.F. (2011). Annotated catalogue of the flowering plants of São Tomé and Príncipe Bothalia 41: 41-82.
    • Giraldo-Cañas, D. (2011). Catálogo de la familia Poaceae en Colombia Darwiniana 49: 139-247.
    • Clayton, W.D. & Snow, N. (2010). A key to Pacific Grasses: 1-107. Kew Publishing, Royal Botanic Gardens, Kew.
    • Ahmed, Z.U. (ed.) (2008). Encyclopedia of Flora and Fauna of Bangladesh 12: 1-505. Asiatic Society of Bangladesh.
    • Pandey, R.P. & Dilwakar, P.G. (2008). An integrated check-list flora of Andaman and Nicobar islands, India Journal of Economic and Taxonomic Botany 32: 403-500.
    • Newman, M., Ketphanh, S., Svengsuksa, B., Thomas, P., Sengdala, K., Lamxay, V. & Armstrong, K. (2007). A checklist of the vascular plants of Lao PDR: 1-394. Royal Botanic Gardens, Edinburgh.
    • Wu, Z. & Raven, P.H. (eds.) (2006). Poaceae Flora of China 22: 1-733. Missouri Botanical Garden Press, St. Louis.
    • Clayton, W.D., Harman, K.T. & Williamson, H. (2006). World Grass Species - Synonymy database The Board of Trustees of the Royal Botanic Gardens, Kew.
    • Dávila, P., Mejia-Saulés, M.T., Gómez-Sánchez, N., Valdés-Reyna, J., Ortíz, J.J., Morín, C., Castrejón, J. & Ocampo, A. (2006). Catálogo de las Gramíneas de México: 1-671. CONABIO, México city.
    • Mallett, K. (ed.) (2005). Flora of Australia 44B: 1-486. Australian Government Publishing Service, Canberra.
    • Kress, W.J., DeFilipps, R.A., Farr, E. & Kyi, D.Y.Y. (2003). A Checklist of the Trees, Shrubs, Herbs and Climbers of Myanmar Contributions from the United States National Herbarium 45: 1-590.
    • Flora of North America Editorial Committee (2003). Flora of North America North of Mexico 25: 1-781. Oxford University Press, New York, Oxford.
    • Turner, I.M. (1995 publ. 1997). A catalogue of the Vascular Plants of Malaya Gardens' Bulletin Singapore 47(2): 347-655.
    • Davidse, G. & al. (eds.) (1994). Flora Mesoamericana 6: 1-543. Universidad Nacional Autónoma de México, México, D.F.
    • Dassanayake (ed.) (1994). A Revised Handbook to the Flora of Ceylon 8: 1-458. Oxford & IBH Publishing Co. PVT. LTD., New Delhi, Calcutta.
    • George, A.S., Orchard, A.E. & Hewson, H.J. (eds.) (1993). Oceanic islands 2 Flora of Australia 50: 1-606. Australian Government Publishing Service, Canberra.
    • Karthikeyan, S., Jain, S.K., Nayar, M.P. & Sanjappa, M. (1989). Florae Indicae Enumeratio: Monocotyledonae: 1-435. Botanical Survey of India, Calcutta.
    • Kobayashi, S. & Ono, M. (1987). A Revised List of Vascular Plants Indigenous and Introduced to the Bonin (Ogasawara) and the Volcano (Kazan) Islands Ogasawara Research 13: 1-55.
    • Koyama, T. (1987). Grasses of Japan and its neighboring regions: an identification manual: 1-570. Kodansha, Tokyo, Japan.
    • Walker, E.H. (1976). Flora of Okinawa and the southern Ryukyu islands: 1-1159. Smithsonian Institution Press, Washington D.C., U.S.A.
    • Hepper, F.N. (ed.) (1972). Flora of West Tropical Africa, ed. 2, 3(2): 277-574.
    • Gilliland, H.B. (1971). A revised flora of Malaya 3: 1-319. The Royal Botanic Gardens, Singapore.
    • Bosser, J. (1969). Gramiées des pasturages et des cultures a Madagascar: 1-440. ORSTOM, Paris.
    • Henty, E.E. (1969). A manual of the grasses of New Guinea: 1-215. Division of Botany, Dpt. of Forests, LAE, New Guinea.

    Literature

    Kew Backbone Distributions
    • Jaramillo Díaz, P. & Guézou, A. (2017). CDF Checklist of Galapagos Vascular Plants - FCD Lista de especies de Plantas Vasculares de Galápagos http://www.darwinfoundation.org/datazone/checklists/vascular-plants/.
    • Valdés-Reyna, J. Villaseñor, J.L., Encina-Domínguez, J.A. & Oritz, E. (2015). The grass family (Poaceae) in Coahuila, Mexico: diversity and distribution Botanical Sciences 93: 119-129.
    • Chang, C.S., Kim, H. & Chang, K.S. (2014). Provisional checklist of vascular plants for the Korea peninsula flora (KPF): 1-660. DESIGNPOST.
    • Forzza, R.C. & al. (2013). Lista de Espécies da Flora do Brasil http://floradobrasil.jbrj.gov.br/2010/.
    • Thiombiano, A., Schmidt, M., Dressler, S., Ouédraogo, A., Hahn, K. & Zizka, G. (2012). Catalogue des plantes vasculaires du Burkina Faso Boissiera 65: 1-391.
    • Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies Smithsonian Contributions to Botany 98: 1-1192.
    • Figueiredo, E., Paiva, J., Stévart, T., Oliveira, F. & Smith, G.F. (2011). Annotated catalogue of the flowering plants of São Tomé and Príncipe Bothalia 41: 41-82.
    • Giraldo-Cañas, D. (2011). Catálogo de la familia Poaceae en Colombia Darwiniana 49: 139-247.
    • Clayton, W.D. & Snow, N. (2010). A key to Pacific Grasses: 1-107. Kew Publishing, Royal Botanic Gardens, Kew.
    • Kandwal, M.K. & Gupta, B.K. (2009). An update on grass flora of Uttarkhand Indian Journal of Forestry 32: 657-668.
    • Ahmed, Z.U. (ed.) (2008). Encyclopedia of Flora and Fauna of Bangladesh 12: 1-505. Asiatic Society of Bangladesh.
    • Pandey, R.P. & Dilwakar, P.G. (2008). An integrated check-list flora of Andaman and Nicobar islands, India Journal of Economic and Taxonomic Botany 32: 403-500.
    • Nelson Sutherland, C.H. (2008). Catálogo de las plantes vasculares de Honduras. Espermatofitas: 1-1576. SERNA/Guaymuras, Tegucigalpa, Honduras.
    • Newman, M., Ketphanh, S., Svengsuksa, B., Thomas, P., Sengdala, K., Lamxay, V. & Armstrong, K. (2007). A checklist of the vascular plants of Lao PDR: 1-394. Royal Botanic Gardens, Edinburgh.
    • Wu, Z. & Raven, P.H. (eds.) (2006). Poaceae Flora of China 22: 1-733. Missouri Botanical Garden Press, St. Louis.
    • Sosef, M.S.M. & al. (2006). Check-list des plantes vasculaires du Gabon Scripta Botanica Belgica 35: 1-438.
    • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
    • Dávila, P., Mejia-Saulés, M.T., Gómez-Sánchez, N., Valdés-Reyna, J., Ortíz, J.J., Morín, C., Castrejón, J. & Ocampo, A. (2006). Catálogo de las Gramíneas de México: 1-671. CONABIO, México city.
    • Mallett, K. (ed.) (2005). Flora of Australia 44B: 1-486. Australian Government Publishing Service, Canberra.
    • Kress, W.J., DeFilipps, R.A., Farr, E. & Kyi, D.Y.Y. (2003). A Checklist of the Trees, Shrubs, Herbs and Climbers of Myanmar Contributions from the United States National Herbarium 45: 1-590.
    • Flora of North America Editorial Committee (2003). Flora of North America North of Mexico 25: 1-781. Oxford University Press, New York, Oxford.
    • Turner, I.M., Xing, F. & Corlett, R.T. (2000). An annotated check-list of the vascular plants of the South China Sea and its shores The Raffles bulletin of Zoology, Suppl. 8: 23-116.
    • Jørgensen, P.M. & León-Yánez, S. (eds.) (1999). Catalogue of the Vascular Plants of Ecuador Monographs in Systematic Botany from the Missouri Botanical Garden 75: i-viii, 1-1181.
    • Davidse, G. & al. (eds.) (1994). Flora Mesoamericana 6: 1-543. Universidad Nacional Autónoma de México, México, D.F.
    • Dassanayake (ed.) (1994). A Revised Handbook to the Flora of Ceylon 8: 1-458. Oxford & IBH Publishing Co. PVT. LTD., New Delhi, Calcutta.
    • George, A.S., Orchard, A.E. & Hewson, H.J. (eds.) (1993). Oceanic islands 2 Flora of Australia 50: 1-606. Australian Government Publishing Service, Canberra.
    • Karthikeyan, S., Jain, S.K., Nayar, M.P. & Sanjappa, M. (1989). Florae Indicae Enumeratio: Monocotyledonae: 1-435. Botanical Survey of India, Calcutta.
    • Kobayashi, S. & Ono, M. (1987). A Revised List of Vascular Plants Indigenous and Introduced to the Bonin (Ogasawara) and the Volcano (Kazan) Islands Ogasawara Research 13: 1-55.
    • Koyama, T. (1987). Grasses of Japan and its neighboring regions: an identification manual: 1-570. Kodansha, Tokyo, Japan.
    • Walker, E.H. (1976). Flora of Okinawa and the southern Ryukyu islands: 1-1159. Smithsonian Institution Press, Washington D.C., U.S.A.
    • Hepper, F.N. (ed.) (1972). Flora of West Tropical Africa, ed. 2, 3(2): 277-574.
    • Gilliland, H.B. (1971). A revised flora of Malaya 3: 1-319. The Royal Botanic Gardens, Singapore.
    • Bosser, J. (1969). Gramiées des pasturages et des cultures a Madagascar: 1-440. ORSTOM, Paris.
    • Henty, E.E. (1969). A manual of the grasses of New Guinea: 1-215. Division of Botany, Dpt. of Forests, LAE, New Guinea.
    • Fosberg, F.R. (1957). The Maldive islands, Indian Ocean Atoll Research Bulletin 58: 1-37.
    • Masamune, G. (1942). Enumeratio phanerogamarum Bornearum: 1-739. Taihoku, Formosa.
    • Hubbard, C.E. & Vaughan, R.E. (1940). Grasses of Mauritius and Rodriguez: 1-128. Crown Agents for the Colonies, London.

    Sources

    GrassBase - The Online World Grass Flora
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2018. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2018. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0