1. Family: Lamiaceae Martinov
    1. Genus: Leonotis (Pers.) R.Br.
      1. Leonotis nepetifolia (L.) R.Br.

        This species is accepted, and its native range is Africa, Indian Subcontinent.

    [FTEA]

    Lamiaceae (Labiatae), A.J. Paton, G. Bramley, O. Ryding, R.M. Polhill, Y.B. Harvey, M. Iwarsson, F. Willis, P.B. Phillipson, K. Balkwill, C.W. Lukhoba, D.F. Otieno, & R.M. Harley. Leonotis, M Iwarsson & YB Harve. Flora of Tropical East Africa. 2009

    Type
    Lectotype, chosen by Iwarsson, loc. cit. (1985): plate on p. 117 of Hermann, Horti Academi Lugduno-Batavi Catalogus (1687), figured from a plant raised from seeds originally from Surinam
    Habit
    Annual or short-lived perennial herb, (0.3–)1–3(–4.5) m high, with a single stem from a small (less than 10 cm long) taproot
    Stem
    Stem with internodes 2–15 cm long, usually repeatedly branching at the nodes below the inflorescence, green (brown at base), with short antrorse or retrorse hairs and sessile glands; usually with a tuft of 1–4 mm long, white or yellowish hairs on the upper half of the leaf nodes, occasionally extending a few cm above the node
    Leaves
    Leaves petiolate; blade ovate, (2.5–)5–20(–23) × (2–)4–15 cm, deeply crenate with 25–51 teeth from base to apex, apex acute to acuminate, base cuneate-attenuate to cordate, both surfaces laxly to densely covered with short hairs and sessile glands, lower surface more densely pubescent (especially on veins) and consequently more greyish green, with 6–10 lateral veins on either side, venation yellowish white; petiole (5–)30–80(–125) mm long, covered with appressed antrorse or retrorse hairs and sessile glands
    Inflorescences
    Inflorescence of (1–)2–5(–7) verticils per branch, usually spaced by prolonged internodes (50–)70–280 mm long; bracts narrowly lanceolate or absent, 25–90(–150) × 4–13 mm, acute, base attenuate, hairy like the leaves; bracteoles linear, (3–)7–15(–20) × 0.5–1.5 mm, acuminate, apically white, hairy like the leaves; verticils spherical to cylindrical, 30–67 mm in diameter, with ± 20–28 branches 2–16 mm long, with 9–18 buds per branch; pedicels 0.5–2 mm long (slightly extended in fruit), shortly pubescent
    Calyx
    Calyx 14–25 mm long at anthesis, slightly enlarged in fruit, 4–5.6 mm in diameter at mouth, 8(–10)-toothed, basally yellowish white and the distal half green, covered with short hairs, occasionally with a tuft of long, spreading hairs on the distal third, subglabrous in fruit, with sessile glands; mouth oblique; tube slightly bent dorsally; teeth deltate to narrowly triangular, apiculate; dorsal tooth 4–7(–11) mm long; lateral teeth 4, 1–3 mm long; ventral teeth 3, united at base forming a recurved anterior lip 4–7 mm long (including apiculate teeth 0.5–3 mm long), to 5 mm wide
    Corolla
    Corolla 19–38 mm long, with orange or whitish yellow hairs; tube 9–20 mm long, widening at mouth, with 3 distinct hair fringes within (near base); posterior lip 10–20 mm long, with white (or occasionally orange) woolly hair enclosing anthers and stigma well developed; anterior lip 3-lobed, (3–)5–7 mm long, deflexed on the tube (but shrivelled and twisted), subglabrous, eglandular or with sessile glands
    Fruits
    Nutlets with a grey and brown marble-like, nonglossy, slightly waxy surface, 2.9–4.3 × 1.1–1.9 mm.

    Images

    Distribution

    Native to:

    Angola, Bangladesh, Benin, Botswana, Burkina, Burundi, Cameroon, Central African Repu, Chad, Congo, Eritrea, Ethiopia, Gabon, Gambia, Ghana, Guinea, Guinea-Bissau, Gulf of Guinea Is., India, Ivory Coast, Kenya, Liberia, Madagascar, Malawi, Mali, Mauritius, Mozambique, Namibia, Nepal, Niger, Nigeria, Northern Provinces, Rwanda, Senegal, Sierra Leone, Sri Lanka, Sudan, Swaziland, Tanzania, Togo, Uganda, Zambia, Zaïre, Zimbabwe

    Introduced into:

    Alabama, Aldabra, Argentina Northeast, Bahamas, Bermuda, Bolivia, Brazil North, Brazil Northeast, Brazil South, Brazil Southeast, Brazil West-Central, Cambodia, Canary Is., China South-Central, Colombia, Costa Rica, Cuba, Dominican Republic, Ecuador, El Salvador, French Guiana, Guatemala, Guyana, Haiti, Honduras, Jamaica, Jawa, Leeward Is., Malaya, Mexico Central, Mexico Gulf, Mexico Northeast, Mexico Northwest, Mexico Southeast, Mexico Southwest, Morocco, Netherlands Antilles, New Caledonia, Nicaragua, Paraguay, Peru, Puerto Rico, Seychelles, Society Is., Southwest Caribbean, St.Helena, Sumatera, Suriname, Trinidad-Tobago, Turks-Caicos Is., Venezuela, Venezuelan Antilles, Vietnam, Windward Is.

    Leonotis nepetifolia (L.) R.Br. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Jan 1, 2009 David, B. [CL857], Cambodia K000855737
    Dec 1, 1994 Rosevear, D. R. [87/37], Cameroon K000025462
    Jan 25, 1994 Garrett, H.B.G. [902], Thailand K000855740
    Jan 25, 1994 Thorel [s.n.], Vietnam K000855743
    Jan 1, 1994 Maitland, T.D. [0s.n.95], Cameroon K000025460
    Jan 1, 1920 Bakhuizen van den Brink, R.C. [4840], Indonesia K000735304
    Smitinand, T. [4878], Thailand K000855742
    Koorders, S.H. [31849], Indonesia K000735303
    Robinson, C.B. [1202], Vietnam K000855739
    Pierre, L. [521A], Vietnam K000855738
    Kerr, A.F.G. [911], Thailand K000855744
    Teruya, Z. [1564], Indonesia K000735302
    De Silva, F. [Cat. no. 2042] K001114967
    Marran, A. [2279], Thailand K000855741
    s.coll. [Cat. no. 2042] K001114968
    s.coll. [Cat. no. 2042] K001114969
    s.coll. [Cat. no. 2042] K001114970
    s.coll. [Cat. no. 2042] K001114971
    s.coll. [Cat. no. 2042] K001114972
    s.coll. [Cat. no. 2042] K001114973

    First published in W.T.Aiton, Hortus Kew. 3: 409 (1811)

    Accepted by

    • Velayos, M., Barberá, P., Cabezas, F.J., de la Estrella, M., Fero, M. & Aedo, C. (2014). Checklist of the vascular plants of Annobón (Equatorial Guinea) Phytotaxa 171: 1-78.
    • Verloove, F. (2013). New xenophytes from Gran Canaria (Canary islands, Spain), with emphasis on naturalised and (potentially) invasive species Collectanea Botanica 32: 59-82.
    • Paton, A.J., Bramley, G., Ryding, O., Polhill, R.M., Harvey, Y.B., Iwarsson, M., Otieno, D., Balkwill, K., Phillipson, P.B., Harley, R.M. & Willis, F. (2013). Flora Zambesiaca 8(8): 1-346. Royal Botanic Gardens, Kew.
    • Leti, M., Hul, S., Fouché, J.-G., Cheng, S.K. & David, B. (2013). Flore photographique du Cambodge: 1-589. Éditions Privat, Toulouse.
    • Davidse, G. & al. (eds.) (2012). Flora Mesoamericana 4(2): 1-533. Universidad Nacional Autónoma de México, México, D.F.
    • López Patiño, E.J., Szeszko, D.R., Rascala Pérez, J. & Beltrán Retis, A.S. (2012). The flora of the Tenacingo-Malinalco-Zumpahuacán protected natural area, state of Mexico, Mexico Harvard Papers in Botany 17: 65-167.
    • Dobignard, A. & Chatelain, C. (2012). Index synonymique de la flore d'Afrique du nord 4: 1-431. Éditions des conservatoire et jardin botaniques, Genève.
    • Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies Smithsonian Contributions to Botany 98: 1-1192.
    • Garcia-Mendoza, A.J. & Meave, J.A. (eds.) (2012). Diversidad florística de Oaxaca: de musgos a angiospermas (colecciones y listas de especies), ed. 2: 1-351. Instituto de Biología, Universidad Nacional Autónoma de México.
    • Morales, R. (2011). Les Labiadas (Lamiaceae) de Guinea Ecuatorial Anales del Jardin Botanico de Madrid 68: 199-223.
    • Idárraga-Piedrahita, A., Ortiz, R.D.C., Callejas Posada, R. & Merello, M. (eds.) (2011). Flora de Antioquia: Catálogo de las Plantas Vasculares 2: 1-939. Universidad de Antioquia, Medellín.
    • Kral, R., Diamond, A.R., Ginzbarg, S.L., Hansen, C.J., Haynes, R.R., Keener, B.R., Lelong, M.G., Spaulding, D.D. & Woods, M. (2011). Annotated checklist of the vascular plants of Alabama: 1-112. Botanical reseach institute of Texas.
    • Paton, A.J., Bramley, G., Ryding, O., Polhill, R., Harvey, Y., Iwarsson, M., Willis, F., Phillipson, P., Balkwill, K., Lukhoba, C., Otiend, D & Harley (2009). Lamiaceae (Labiatae) Flora of Tropical East Africa: 1-430.
    • Khanam, M. & Hassan, M.A. (2008). Lamiaceae Flora of Bangladesh 58: 1-161. Bangladesh National Herbarium, Dhaka.
    • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
    • Hokche, O., Berry, P.E. & Huber, O. (eds.) (2008). Nuevo Catálogo de la Flora Vascular de Venezuela: 1-859. Fundación Instituto Botánico de Venezuela.
    • Nelson Sutherland, C.H. (2008). Catálogo de las plantes vasculares de Honduras. Espermatofitas: 1-1576. SERNA/Guaymuras, Tegucigalpa, Honduras.
    • Sosef, M.S.M. & al. (2006). Check-list des plantes vasculaires du Gabon Scripta Botanica Belgica 35: 1-438.
    • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
    • Mabberley, D.J. & De Kok, R.P.J. (2004). Labiatae Flore de la Nouvelle-Calédonie et Dépendances 25: 20-141. Muséum National d'Histoire Naturelle, Paris.
    • Calane da Silva, M., Izdine, S. & Amuse, A.B. (2004). A Preliminary Checklist of the Vascular Plants of Mozambique: 1-184. SABONET, Pretoria.
    • Govaerts, R. (2003). World Checklist of Selected Plant Families Database in ACCESS: 1-216203. The Board of Trustees of the Royal Botanic Gardens, Kew.
    • Welsh, S.L. (1998). Flora Societensis: 1-420. E.P.S. Inc. Utah.
    • Gonzalez, F., Nelson Diaz, J. & Lowry, P. (1995). Flora Illustrada de San Andrés y Providencia: 1-281. Universidad Nacional de Colombia, Instituto de Ciencias Naturales, Colombia.
    • Friedmann, F. (1994). Flore des Seychelles Dicotylédones: 1-663. ORSTOM éditions.
    • MacKee, H.S. (1994). Catalogue des plantes introduites et cultivées en Nouvelle-Calédonie, ed. 2: 1-164. Museum national d'histoire naturelle, Paris.
    • Boulvert, Y. (1977). Catalogue de la Flore de Centrafrique 2(1): 1-85. ORSTOM, Bangui.
    • Baker, R.E.D. in Williams, R.O. (1955). Labiatae Flora of Trinidad and Tobago 2(6): 421-435. Government Printing Office, Port-of-Spain.

    Literature

    Kew Backbone Distributions
    • Leti, M., Hul, S., Fouché, J.-G., Cheng, S.K. & David, B. (2013). Flore photographique du Cambodge: 1-589. Éditions Privat, Toulouse.
    • López Patiño, E.J., Szeszko, D.R., Rascala Pérez, J. & Beltrán Retis, A.S. (2012). The flora of the Tenacingo-Malinalco-Zumpahuacán protected natural area, state of Mexico, Mexico Harvard Papers in Botany 17: 65-167.
    • Dobignard, A. & Chatelain, C. (2012). Index synonymique de la flore d'Afrique du nord 4: 1-431. Éditions des conservatoire et jardin botaniques, Genève.
    • Kral, R., Diamond, A.R., Ginzbarg, S.L., Hansen, C.J., Haynes, R.R., Keener, B.R., Lelong, M.G., Spaulding, D.D. & Woods, M. (2011). Annotated checklist of the vascular plants of Alabama: 1-112. Botanical reseach institute of Texas.
    • Khanam, M. & Hassan, M.A. (2008). Lamiaceae Flora of Bangladesh 58: 1-161. Bangladesh National Herbarium, Dhaka.
    • Hokche, O., Berry, P.E. & Huber, O. (eds.) (2008). Nuevo Catálogo de la Flora Vascular de Venezuela: 1-859. Fundación Instituto Botánico de Venezuela.
    • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
    • Mabberley, D.J. & De Kok, R.P.J. (2004). Labiatae Flore de la Nouvelle-Calédonie et Dépendances 25: 20-141. Muséum National d'Histoire Naturelle, Paris.
    • Iwarsson, M. & Harvey, Y. (2003). Monograph of the genus Leonotis (Pers.) R.Br. (Lamiaceae) Kew Bulletin 58: 597-645.
    • Welsh, S.L. (1998). Flora Societensis: 1-420. E.P.S. Inc. Utah.
    • Gonzalez, F., Nelson Diaz, J. & Lowry, P. (1995). Flora Illustrada de San Andrés y Providencia: 1-281. Universidad Nacional de Colombia, Instituto de Ciencias Naturales, Colombia.
    • Friedmann, F. (1994). Flore des Seychelles Dicotylédones: 1-663. ORSTOM éditions.
    • Brown, L.C. (1982). The Flora and Fauna of St Helena: 1-88. Land Resources Development Centre, Surbiton, England.
    • Correll, D.S. & Correll, H.B. (1982). Flora of the Bahama Archipelago: 1-1692. J.Cramer, Vaduz.
    • Baker, R.E.D. in Williams, R.O. (1955). Labiatae Flora of Trinidad and Tobago 2(6): 421-435. Government Printing Office, Port-of-Spain.
    • Britton, N. (1918). Flora of Bermuda: 1-585. Charles Scribner's Sons, New York.
    Flora of Tropical East Africa
    • Fl. Eth. 5: 549 (2006).
    • K.B. 58: 632 (2003)
    • K.T.S.L.: 628, fig. (1994)
    • U.K.W.F. ed. 2: 285 (1994)
    • Blundell, Wild Fl. E. Afr., ed. 2: 403, t. 432 (1992)
    • F.S.A. 28 (4): 37 (1985)
    • F.W.T.A., ed. 2, 2: 470, fig. (1963)
    • F.P.U.: 152, fig. 96 (1962)
    • F.P.S. 3: 212 (1956)
    • U.O.P.Z.: 327, t. 329 (1949)
    • F.P.N.A. 2: 155 (1947)
    • F.T.A. 5: 491 (1900)
    • E.P. IV. 3a: 246 (1896)
    • Bot. Mag. 65, t. 3700 (1839)
    • Aiton, Hort. Kew., ed. 2, 3: 409 (1811)

    Sources

    Art and Illustrations in Digifolia
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2018. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2018. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Plants and People Africa
    Common Names from Plants and People Africa http://www.plantsandpeopleafrica.com/
    © Plants and People Africa http://www.plantsandpeopleafrica.com http://creativecommons.org/licenses/by/3.0/