Skip to main content
This species is accepted, and its native range is Africa, Tropical & Subtropical Asia to N. Australia.
A specimen from Kew's Herbarium

[FZ]

Sphenocleaceae, M. Thulin. Flora Zambesiaca 7:1. 1983

Morphology General Habit
Herb, up to 1·5 m. tall.
Morphology Stem
Stem hollow, often much branched.
Morphology Leaves
Leaf–blade lanceolate to narrowly ovate or elliptic, up to 12 x 5 cm., attenuate at the base, acute or subacute at the apex; petiole up to 1·5 cm. long.
Morphology Reproductive morphology Inflorescences
Spikes cylindrical–conical, up to 12 cm. long but usually much shorter, and c. 1 cm. in diameter, with only a few flowers open at a time; peduncle up to 8 cm. long; bracts and bracteoles oblanceolate–spathulate, the tips arched over the flowers except at anthesis.
Morphology Reproductive morphology Flowers
Flowers sessile, attached longitudinally to the rhachis by a linear base.
Morphology Reproductive morphology Flowers Calyx
Calyx–lobes broadly triangular, rounded at apex, 1–1·5 mm. long.
Morphology Reproductive morphology Flowers Corolla
Corolla white to greenish white or pink, 2–3 mm. long; lobes ovate–triangular, united about half–way.
Morphology Reproductive morphology Flowers Androecium Stamens Filaments
Filaments slightly dilated at base; anthers c. 0·5 mm. long.
Morphology Reproductive morphology Flowers Gynoecium Ovary
Ovary obovoid, 1–5–2 mm. long.
Morphology Reproductive morphology Fruits
Capsule 4–5 mm. in diameter, dehiscing below the calyx–lobes which fall with the lid, leaving the scarious base persistent on the rhachis.
Morphology Reproductive morphology Seeds
Seeds yellowish brown, oblong, c. 0·5 mm. long.

[CPLC]

Bernal, R., Gradstein, S.R. & Celis, M. (eds.). 2015. Catálogo de plantas y líquenes de Colombia. Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá. http://catalogoplantasdecolombia.unal.edu.co

Distribution
Nativa en Colombia; Alt. 0 - 900 m.; Amazonia, Guayana y Serranía de La Macarena, Llanura del Caribe, Orinoquia, Valle del Cauca, Valle del Magdalena.
Morphology General Habit
Hierba
Conservation
Preocupación Menor

[FTEA]

Sphenocleaceae, H. K. Airy Shaw. Flora of Tropical East Africa. 1968

Morphology General Habit
Roots numerous, long, cord-like.
Morphology Stem
Stem hollow, up to 150 cm. tall, often much branched.
Morphology Leaves
Leaf-blade oblong to lanceolate-oblong, attenuate at both ends, acute or obtuse, 2.5–12.5 cm. long; petiole up to 3 cm. long.
Morphology Reproductive morphology Inflorescences
Spikes cylindric, up to 7.5 cm. long, narrowed at apex; peduncle up to 8 cm. long.
Morphology Reproductive morphology Inflorescences Bracts
Bracts and bracteoles spatulate, the tips arched over the flowers except during anthesis.
Morphology Reproductive morphology Flowers
Flowers densely crowded, though characteristically only 1 or 2 open at a time, rhomboid or hexagonal by compression, sessile, wedge-shaped below, attached longitudinally to the rhachis by a linear base.
Morphology Reproductive morphology Flowers Calyx
Calyx-segments deltoid-semicircular, obtuse, ultimately slightly accrescent and connivent.
Morphology Reproductive morphology Flowers Corolla
Corolla whitish, pinkish or purplish, 2.5–4 mm. long; segments ovate-triangular, obtuse or acute, united about half-way, connivent.
Morphology Reproductive morphology Flowers Androecium Stamens Filaments
Filaments slightly dilated at base.
Morphology Reproductive morphology Flowers Gynoecium Ovary
Ovary obovoid, 2.5 mm. long, apex broad, free, truncate.
Morphology Reproductive morphology Fruits
Capsule 4–5 mm. diameter, dehiscing below the calyx-segments which fall with the lid, leaving the scarious base persistent on the rhachis.
Morphology Reproductive morphology Seeds
Seeds yellowish-brown, 0.5 mm. long.
Figures
Fig. 1.
Habitat
In and near pools, swamps, streamsides, periodically inundated depressions, ditches, irrigation channels, and wet places generally; 0–1250 m.
Distribution
widespread in tropical Africa (excluding the NE. Horn), extending south to the Transvaalalso in Madagascarwidespread (but probably introduced) in tropical Asia and America P T1 T3 T5 T6 U3 Z

[FWTA]

Sphenocleaceae, F. N. Hepper. Flora of West Tropical Africa 2. 1963

Morphology General Habit
An erect glabrous herb with spongy stems 1-4 ft. high
Morphology Reproductive morphology Flowers
Flowers greenish-white in congested spikes
Ecology
In rice-fields, beside rivers and open swamps, often gregarious in tidal creeks.

Doubtfully present in:

Angola

Native to:

Afghanistan, Andaman Is., Assam, Bangladesh, Benin, Botswana, Burkina, Burundi, Cambodia, Cameroon, Central African Repu, Chad, China South-Central, China Southeast, Congo, Egypt, Ethiopia, Gabon, Gambia, Ghana, Guinea, Guinea-Bissau, Hainan, India, Iran, Iraq, Ivory Coast, Jawa, KwaZulu-Natal, Lesser Sunda Is., Madagascar, Malawi, Malaya, Mali, Mauritania, Mozambique, Myanmar, Namibia, Nepal, Niger, Nigeria, Northern Provinces, Northern Territory, Pakistan, Philippines, Rwanda, Senegal, Sierra Leone, Solomon Is., Somalia, Sri Lanka, Sudan, Sulawesi, Swaziland, Tadzhikistan, Taiwan, Tanzania, Thailand, Togo, Uganda, Uzbekistan, Vietnam, Zambia, Zaïre, Zimbabwe

Introduced into:

Alabama, Argentina Northeast, Argentina Northwest, Arkansas, Belize, Bolivia, Brazil North, Brazil Northeast, Brazil Southeast, Brazil West-Central, Colombia, Costa Rica, Cuba, Dominican Republic, Ecuador, Florida, Georgia, Guatemala, Haiti, Honduras, Japan, Louisiana, Mexico Southwest, Mississippi, Missouri, Nicaragua, North Carolina, Oklahoma, Paraguay, Puerto Rico, South Carolina, Texas, Trinidad-Tobago, Venezuela

Sphenoclea zeylanica Gaertn. appears in other Kew resources:

Date Reference Identified As Barcode Type Status
Jones, A.P.D. [18808], Nigeria 16965.000
Deighton, F.C., Sierra Leone 16966.000
Meikle, R.D., Nigeria 16967.000
Meikle, R.D. [995], Nigeria 17394.000
Hepper, F.N. [1261], Cameroon 18122.000
Harrison, S.G. [899], Guyana 18931.000
Colombia 46100.000
Bidgood, S. [6061], Tanzania K000468364
Wallich, N. [Cat. no. 1516], Malaysia K001113307
s.coll. [Cat. no. 1516] K001113311
s.coll. [Cat. no. 1516] K001113312
s.coll. [Cat. no. 1516] K001113316
De Silva, F. [Cat. no. 1516] K001113306
s.coll. [Cat. no. 1516] K001113313
s.coll. [Cat. no. 1516], India K001113314
s.coll. [Cat. no. 1516] K001113315
s.coll. [Cat. no. 1516] K001113308
Gomez, W. [Cat. no. 1516], Myanmar K001113310
s.coll. [Cat. no. 1516] K001113317
s.coll. [Cat. no. 1516] K001113309

First published in Fruct. Sem. Pl. 1: 113 (1788)

Accepted by

  • Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies Smithsonian Contributions to Botany 98: 1-1192.
  • Airy Shaw, H.K. (1968). Flora of Tropical East Africa, Sphenocleaceae: 1-3.
  • Balick, M. J., M. H. Nee & D.E. Atha (2000). Checklist of the vascular plants of Belize Memoirs of the New York Botanical Garden 85: 1-246.
  • Barooah, C. & Ahmed, I. (2014). Plant diversity of Assam. A checklist of Angiosperms and Gymnosperms: 1-599. Assam science technology and environment council, India.
  • Barry, J. P. & Celles, J.S. (1991). Flore de Mauritanie 1: 1-359. Centre Regional de Documentation Pedagogique, Nice.
  • Boudet, G., Lebrun, J.P. & Demange, R. (1986). Catalogue des plantes vasculaires du Mali: 1-465. Etudes d'Elevage et de Médecine Vétérinaire des Pays Tropicaux.
  • Boulos, L. (2002). Flora of Egypt 3: 1-373. Al Hadara Publishing, Cairo.
  • Boulvert, Y. (1977). Catalogue de la Flore de Centrafrique 2(1): 1-85. ORSTROM, Bangui.
  • Brunel, J.F., Hiepo, P. & Scholz, H. (eds.) (1984). Flore Analytique du Togo Phanérogames: 1-751. GTZ, Eschborn.
  • Catarino, L., Martins, E.S., Diniz, M.A. & Pinto-Basto, M.F. (2006). Check-list da flora vascular do parque natural das Lagos de Cufada (Guiné-Bissau) Garcia de Orta, Série de Botânica 17: 97-141.
  • Catarino, L., Sampaio Martins, E., Pinto-Basto, M.F. & Diniz, M.A. (2006). Plantas Vasculares e Briófitos da Guiné-Bissau: 1-298. Instituto de investigação científica tropical, Instituto Português de apoio ao desenvolvimento.
  • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
  • Dassanayake (ed.) (1983). A Revised Handbook to the Flora of Ceylon 4: 1-532. Oxford & IBH Publishing Co. PVT. LTD., New Delhi, Calcutta.
  • Dy Phon, P. (2000). Dictionnaire des plantes utilisées au Cambodge: 1-915. chez l'auteur, Phnom Penh, Cambodia.
  • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
  • Garcia-Mendoza, A.J. & Meave, J.A. (eds.) (2012). Diversidad florística de Oaxaca: de musgos a angiospermas (colecciones y listas de especies), ed. 2: 1-351. Instituto de Biología, Universidad Nacional Autónoma de México.
  • Germishuizen, G. & Meyer, N.L. (eds.) (2003). Plants of Southern Africa an annotated checklist Strelitzia 14: 1-1231. National Botanical Institute, Pretoria.
  • Ghazanfar, S.A. & Edmandson, J.R. (eds.) (2016). Flora of Iraq 5(1): 1-284. Ministry of Agriculture & Agrarian Reform, Baghdad.
  • Girmansyah, D. & al. (eds.) (2013). Flora of Bali an annotated checklist: 1-158. Herbarium Bogorensis, Indonesia.
  • Hammel, B.E., Grayum, M.H., Herrera, C. & Zamora, N. (eds.) (2015). Manual de Plantas de Costa Rica. Volumen VIII. Dicotyledóneas (Sabiaceae-Zygophyllaceae) Monographs in Systematic Botany from the Missouri Botanical Garden 131: 1-657. Missouri Botanical Garden.
  • Hancock, I.R. & Henderson, C.P. (1988). Flora of the Solomon Islands Research Bulletin Dodo Creek Research Station 7: 1-203.
  • Hedberg, I., Kelbessa, E., Edwards, S., Demissew, S. & Persson, E. (eds.) (2006). Flora of Ethiopia and Eritrea 5: 1-690. The National Herbarium, Addis Ababa University, Ethiopia & The Department of Systematic Botany, Upps.
  • Hokche, O., Berry, P.E. & Huber, O. (eds.) (2008). Nuevo Catálogo de la Flora Vascular de Venezuela: 1-859. Fundación Instituto Botánico de Venezuela.
  • Hutchinson, J., Dalziel, J.M. & Keay, R.W.J. (1954-1958). Flora of West Tropical Africa, ed. 2, 1: 1-828.
  • Idárraga-Piedrahita, A., Ortiz, R.D.C., Callejas Posada, R. & Merello, M. (eds.) (2011). Flora de Antioquia: Catálogo de las Plantas Vasculares 2: 1-939. Universidad de Antioquia, Medellín.
  • Jørgensen, P.M. & León-Yánez, S. (eds.) (1999). Catalogue of the Vascular Plants of Ecuador Monographs in Systematic Botany from the Missouri Botanical Garden 75: i-viii, 1-1181. Missouri Botanical Garden.
  • Jørgensen, P.M., Nee, M.H. & Beck., S.G. (eds.) (2013). Catálogo de las plantas vasculares de Bolivia Monographs in Systematic Botany from the Missouri Botanical Garden 127: 1-1741. Missouri Botanical Garden.
  • Kral, R., Diamond, A.R., Ginzbarg, S.L., Hansen, C.J., Haynes, R.R., Keener, B.R., Lelong, M.G., Spaulding, D.D. & Woods, M. (2011). Annotated checklist of the vascular plants of Alabama: 1-112. Botanical reseach institute of Texas.
  • Kress, W.J., DeFilipps, R.A., Farr, E. & Kyi, D.Y.Y. (2003). A Checklist of the Trees, Shrubs, Herbs and Climbers of Myanmar Contributions from the United States National Herbarium 45: 1-590. Smithsonian Institution.
  • Launert, E. (ed.) (1983). Flora Zambesiaca 7(1): 1-394. Royal Botanic Gardens, Kew.
  • Lebrun, J.-P., Audru, J., Gaston, A. & Mosnier, M. (1972). Catalogue des Plantes Vasculaires du Tchad Méridional: 1-289. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
  • Lebrun, J.p., Toutain, B., Gaston, A. & Boudet, G. (1991). Catalogue des Plantes Vasculaires du Burkina Faso: 1-341. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
  • Lejoy, J., Ndjele, M.-B. & Geerinck, D. (2010). Catalogue-flore des plantes vasculaires des districts de Kisangani et de la Tshopo (RD Congo) Taxonomania 30: 1-307.
  • Lewalle, J. (1970). Liste floristique et répartition altitudinale de la flore du Burundi occidental: 1-84. Université officielle de Bujumbura.
  • Lisowski, S. (2009). Flore (Angiospermes) de la République de Guinée Scripta Botanica Belgica 41: 1-517.
  • Lê, T.C. (2005). Danh l?c các loài th?c v?t Vi?t Nam 3: 1-1248. Hà N?i : Nhà xu?t b?n Nông nghi?p.
  • Mostaph, M.K. & Uddin, S.B. (2013). Dictionary of plant names of Bangladesh, Vasc. Pl.: 1-434. Janokalyan Prokashani, Chittagong, Bangladesh.
  • Nelson Sutherland, C.H. (2008). Catálogo de las plantes vasculares de Honduras. Espermatofitas: 1-1576. SERNA/Guaymuras, Tegucigalpa, Honduras.
  • Pandey, R.P. & Dilwakar, P.G. (2008). An integrated check-list flora of Andaman and Nicobar islands, India Journal of Economic and Taxonomic Botany 32: 403-500.
  • Peyre de Fabregues, B. & Lebrun, J.-P. (1976). Catalogue des Plantes Vascularies du Niger: 1-433. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
  • Sennikov, A.N. (ed.) (2017). Flora of Uzbekistan 2: 1-200. Navro'z Publishers, Toshkent.
  • Sikarwar, R.L.S. (2014). Angiosperm diversity assessment of Chitrakootthe legendary place of Vindhyan range, India Journal of Economic and Taxonomic Botany 38: 563-619.
  • Singh, M. & Ali, S.J. (2008). A study on aquatic Angiosperms of district Mau (East U.P.) Journal of East African Natural History 32: 877-890.
  • Sita, P. & Moutsambote, J.-M. (2005). Catalogue des plantes vasculaires du Congo, ed. sept. 2005: 1-158. ORSTOM, Centre de Brazzaville.
  • Sosef, M.S.M. & al. (2006). Check-list des plantes vasculaires du Gabon Scripta Botanica Belgica 35: 1-438.
  • Stevens, W.D., Ulloa U., C., Pool, A. & Montiel, O.M. (2001). Flora de Nicaragua Monographs in Systematic Botany from the Missouri Botanical Garden 85: i-xlii, 1-2666. Missouri Botanical Garden.
  • Thulin, M. (ed.) (2006). Flora of Somalia 3: 1-626. The Royal Botanic Gardens, Kew.
  • Van Steenis, C.G.G.J. (ed.) (1955-1958). Flora Malesiana 5: 1-595. Noordhoff-Kolff N.V., Djakarta.
  • Wu, Z. & Raven, P.H. (eds.) (2011). Flora of China 19: 1-884. Science Press (Beijing) & Missouri Botanical Garden Press (St. Louis).
  • Zuloaga, F.O., Morrone, O. , Belgrano, M.J., Marticorena, C. & Marchesi, E. (eds.) (2008). Catálogo de las Plantas Vasculares del Cono Sur Monographs in Systematic Botany from the Missouri Botanical Garden 107: 1-3348. Missouri Botanical Garden.
  • de Moura Júnior, E.G. & al. (2015). Updated checklist of aquatic macrophytes from Northern Brazil Acta Amazonica 45: 111-132.

Literature

Catálogo de Plantas y Líquenes de Colombia

  • ColPlantA (2021). "ColPlantA. Facilitated by the Royal Botanic Gardens, Kew. Published on the Internet; http://www.colplanta.org/"

Flora of West Tropical Africa

  • Airy Shaw in Fl. Males., ser. 1, 4: 27.
  • Berhaut Fl. Sén. 197
  • Chev. Bot. 382
  • F.T.A. 3: 481
  • Fruct. 1: 113, t. 24, fig. 5 (1788)

Kew Backbone Distributions

  • ( 2021-continuously updated). Natural Resources Conservation Services Plant Database http://plants.usda.gov/cgi_bin/topics.cgi?earl=checklist.html.
  • Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies Smithsonian Contributions to Botany 98: 1-1192.
  • Airy Shaw, H.K. (1968). Flora of Tropical East Africa, Sphenocleaceae: 1-3.
  • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
  • Balick, M. J., M. H. Nee & D.E. Atha (2000). Checklist of the vascular plants of Belize Memoirs of the New York Botanical Garden 85: 1-246.
  • Barooah, C. & Ahmed, I. (2014). Plant diversity of Assam. A checklist of Angiosperms and Gymnosperms: 1-599. Assam science technology and environment council, India.
  • Barry, J. P. & Celles, J.S. (1991). Flore de Mauritanie 1: 1-359. Centre Regional de Documentation Pedagogique, Nice.
  • Boudet, G., Lebrun, J.P. & Demange, R. (1986). Catalogue des plantes vasculaires du Mali: 1-465. Etudes d'Elevage et de Médecine Vétérinaire des Pays Tropicaux.
  • Boulos, L. (2002). Flora of Egypt 3: 1-373. Al Hadara Publishing, Cairo.
  • Boulvert, Y. (1977). Catalogue de la Flore de Centrafrique 2(1): 1-85. ORSTROM, Bangui.
  • Brunel, J.F., Hiepo, P. & Scholz, H. (eds.) (1984). Flore Analytique du Togo Phanérogames: 1-751. GTZ, Eschborn.
  • Catarino, L., Martins, E.S., Diniz, M.A. & Pinto-Basto, M.F. (2006). Check-list da flora vascular do parque natural das Lagos de Cufada (Guiné-Bissau) Garcia de Orta, Série de Botânica 17: 97-141.
  • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
  • Dassanayake (ed.) (1983). A Revised Handbook to the Flora of Ceylon 4: 1-532. Oxford & IBH Publishing Co. PVT. LTD., New Delhi, Calcutta.
  • Dy Phon, P. (2000). Dictionnaire des plantes utilisées au Cambodge: 1-915. chez l'auteur, Phnom Penh, Cambodia.
  • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
  • Forzza, R.C., Zappi, D. & Souza, V.C. (2016-continuously updated). Flora do Brasil 2020 em construção http://reflora.jbrj.gov.br/reflora/listaBrasil/ConsultaPublicaUC/ResultadoDaConsultaNovaConsulta.do.
  • GBIF (2008-2020). Global Biodiversity Information Facility http://www.gbif.org/.
  • Garcia-Mendoza, A.J. & Meave, J.A. (eds.) (2012). Diversidad florística de Oaxaca: de musgos a angiospermas (colecciones y listas de especies), ed. 2: 1-351. Instituto de Biología, Universidad Nacional Autónoma de México.
  • Germishuizen, G. & Meyer, N.L. (eds.) (2003). Plants of Southern Africa an annotated checklist Strelitzia 14: 1-1231. National Botanical Institute, Pretoria.
  • Ghazanfar, S.A. & Edmandson, J.R. (eds.) (2016). Flora of Iraq 5(1): 1-284. Ministry of Agriculture & Agrarian Reform, Baghdad.
  • Girmansyah, D. & al. (eds.) (2013). Flora of Bali an annotated checklist: 1-158. Herbarium Bogorensis, Indonesia.
  • Hammel, B.E., Grayum, M.H., Herrera, C. & Zamora, N. (eds.) (2015). Manual de Plantas de Costa Rica. Volumen VIII. Dicotyledóneas (Sabiaceae-Zygophyllaceae) Monographs in Systematic Botany from the Missouri Botanical Garden 131: 1-657. Missouri Botanical Garden.
  • Hancock, I.R. & Henderson, C.P. (1988). Flora of the Solomon Islands Research Bulletin Dodo Creek Research Station 7: 1-203.
  • Hedberg, I., Kelbessa, E., Edwards, S., Demissew, S. & Persson, E. (eds.) (2006). Flora of Ethiopia and Eritrea 5: 1-690. The National Herbarium, Addis Ababa University, Ethiopia & The Department of Systematic Botany, Upps.
  • Hepper, F.N. (ed.) (1963). Flora of West Tropical Africa, ed. 2, 2: 1-544.
  • Hokche, O., Berry, P.E. & Huber, O. (eds.) (2008). Nuevo Catálogo de la Flora Vascular de Venezuela: 1-859. Fundación Instituto Botánico de Venezuela.
  • Hutchinson, J., Dalziel, J.M. & Keay, R.W.J. (1954-1958). Flora of West Tropical Africa, ed. 2, 1: 1-828.
  • Idárraga-Piedrahita, A., Ortiz, R.D.C., Callejas Posada, R. & Merello, M. (eds.) (2011). Flora de Antioquia: Catálogo de las Plantas Vasculares 2: 1-939. Universidad de Antioquia, Medellín.
  • Jørgensen, P.M. & León-Yánez, S. (eds.) (1999). Catalogue of the Vascular Plants of Ecuador Monographs in Systematic Botany from the Missouri Botanical Garden 75: i-viii, 1-1181. Missouri Botanical Garden.
  • Jørgensen, P.M., Nee, M.H. & Beck., S.G. (eds.) (2013). Catálogo de las plantas vasculares de Bolivia Monographs in Systematic Botany from the Missouri Botanical Garden 127: 1-1741. Missouri Botanical Garden.
  • Kral, R., Diamond, A.R., Ginzbarg, S.L., Hansen, C.J., Haynes, R.R., Keener, B.R., Lelong, M.G., Spaulding, D.D. & Woods, M. (2011). Annotated checklist of the vascular plants of Alabama: 1-112. Botanical reseach institute of Texas.
  • Kress, W.J., DeFilipps, R.A., Farr, E. & Kyi, D.Y.Y. (2003). A Checklist of the Trees, Shrubs, Herbs and Climbers of Myanmar Contributions from the United States National Herbarium 45: 1-590. Smithsonian Institution.
  • Launert, E. (ed.) (1983). Flora Zambesiaca 7(1): 1-394. Royal Botanic Gardens, Kew.
  • Lebrun, J.-P., Audru, J., Gaston, A. & Mosnier, M. (1972). Catalogue des Plantes Vasculaires du Tchad Méridional: 1-289. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
  • Lebrun, J.p., Toutain, B., Gaston, A. & Boudet, G. (1991). Catalogue des Plantes Vasculaires du Burkina Faso: 1-341. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
  • Lejoy, J., Ndjele, M.-B. & Geerinck, D. (2010). Catalogue-flore des plantes vasculaires des districts de Kisangani et de la Tshopo (RD Congo) Taxonomania 30: 1-307.
  • Lewalle, J. (1970). Liste floristique et répartition altitudinale de la flore du Burundi occidental: 1-84. Université officielle de Bujumbura.
  • Lisowski, S. (2009). Flore (Angiospermes) de la République de Guinée Scripta Botanica Belgica 41: 1-517.
  • Lê, T.C. (2005). Danh l?c các loài th?c v?t Vi?t Nam 3: 1-1248. Hà N?i : Nhà xu?t b?n Nông nghi?p.
  • Mostaph, M.K. & Uddin, S.B. (2013). Dictionary of plant names of Bangladesh, Vasc. Pl.: 1-434. Janokalyan Prokashani, Chittagong, Bangladesh.
  • Nasir, E. & Ali, S.I. (eds.) (1970-1995). Flora of West Pakistan 1-131.
  • Nelson Sutherland, C.H. (2008). Catálogo de las plantes vasculares de Honduras. Espermatofitas: 1-1576. SERNA/Guaymuras, Tegucigalpa, Honduras.
  • Ovchinnikov, P.N. (ed.) (1988). Flora Tadzhikskoi SSR 9: 1-568. Izd-vo Akademii nauk SSSR, Moskva.
  • Pandey, R.P. & Dilwakar, P.G. (2008). An integrated check-list flora of Andaman and Nicobar islands, India Journal of Economic and Taxonomic Botany 32: 403-500.
  • Peyre de Fabregues, B. & Lebrun, J.-P. (1976). Catalogue des Plantes Vascularies du Niger: 1-433. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
  • Sennikov, A.N. (ed.) (2017). Flora of Uzbekistan 2: 1-200. Navro'z Publishers, Toshkent.
  • Sikarwar, R.L.S. (2014). Angiosperm diversity assessment of Chitrakootthe legendary place of Vindhyan range, India Journal of Economic and Taxonomic Botany 38: 563-619.
  • Sita, P. & Moutsambote, J.-M. (2005). Catalogue des plantes vasculaires du Congo, ed. sept. 2005: 1-158. ORSTOM, Centre de Brazzaville.
  • Sosef, M.S.M. & al. (2006). Check-list des plantes vasculaires du Gabon Scripta Botanica Belgica 35: 1-438.
  • Stevens, W.D., Ulloa U., C., Pool, A. & Montiel, O.M. (2001). Flora de Nicaragua Monographs in Systematic Botany from the Missouri Botanical Garden 85: i-xlii, 1-2666. Missouri Botanical Garden.
  • Thulin, M. (ed.) (2006). Flora of Somalia 3: 1-626. The Royal Botanic Gardens, Kew.
  • Van Steenis, C.G.G.J. (ed.) (1955-1958). Flora Malesiana 5: 1-595. Noordhoff-Kolff N.V., Djakarta.
  • Van Steenis, C.G.G.J. (ed.) (1960-1972). Flora Malesiana 6: 1-1023. Noordhoff-Kolff N.V., Djakarta.
  • Wu, Z. & Raven, P.H. (eds.) (2011). Flora of China 19: 1-884. Science Press (Beijing) & Missouri Botanical Garden Press (St. Louis).
  • Zuloaga, F.O., Morrone, O. , Belgrano, M.J., Marticorena, C. & Marchesi, E. (eds.) (2008). Catálogo de las Plantas Vasculares del Cono Sur Monographs in Systematic Botany from the Missouri Botanical Garden 107: 1-3348. Missouri Botanical Garden.
  • de Moura Júnior, E.G. & al. (2015). Updated checklist of aquatic macrophytes from Northern Brazil Acta Amazonica 45: 111-132.

Flora of Tropical East Africa

  • Airy Shaw in Fl. Males., ser. 1, 4: 27, fig. 1 (1948).
  • Engl. in Die Pflanzenwelt Ost-Afrikas und der Nachbargebiete, Theile C: 400 (1895).
  • F. W. Andr., The Flowering Plants of the Sudan 3: 71, fig. 13 (1956).
  • Gaertn., De Fruct. 1: 113, t. 24/5 (1788).
  • Hepper in Flora of West Tropical Africa, ed. 2, 2: 307, fig. 272 (1963).
  • Robyns, Flore des Spermatophytes du Parc National Albert 2: 405 (1947).
  • Roessler in Prodr. Fl. Südwestafr., 138, Sphenocleaceae (1966).
  • W. B. Hemsl. in Flora of Tropical Africa 3: 481 (1877).

Art and Illustrations in Digifolia
Digital Image © Board of Trustees, RBG Kew

Catálogo de Plantas y Líquenes de Colombia
http://creativecommons.org/licenses/by-nc-sa/3.0

Colombian resources for Plants made Accessible
http://creativecommons.org/licenses/by/3.0/

Flora Zambesiaca
Flora Zambesiaca
http://creativecommons.org/licenses/by-nc-sa/3.0

Flora of Tropical East Africa
Flora of Tropical East Africa
http://creativecommons.org/licenses/by-nc-sa/3.0

Flora of West Tropical Africa
Flora of West Tropical Africa
http://creativecommons.org/licenses/by-nc-sa/3.0

Herbarium Catalogue Specimens
Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

Kew Backbone Distributions
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

Kew Names and Taxonomic Backbone
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0