1. Family: Capparaceae Juss.
    1. Genus: Maerua Forssk.
      1. Maerua angolensis DC.

        This species is accepted, and its native range is Tropical & S. Africa.

    [FZ]

    Capparidaceae, H. Wild. Flora Zambesiaca 1:1. 1960

    Habit
    Shrub or small tree up to c. 8 m. tall; young branches pubescent or glabrescent, becoming densely lenticelled.
    Leaves
    Leaves simple; lamina 2.5–7 x 1.3–5.5 cm., lanceolate, broadly elliptic or ovate to broadly ovate, apex rounded or emarginate, mucronulate, base broadly cuneate or rounded, glabrous or pubescent on both sides, nerves in 5–6 pairs, slightly prominent on both sides; petioles up to 3 cm. long, often slightly swollen just below the lamina.
    Inflorescences
    Inflorescence of short, terminal, corymbose racemes on the main branches or on short side branches or flower solitary in the axils of the upper leaves; pedicels up to 2.5 cm. long, glabrous or pubescent; bracts subulate, trifid with the longer middle lobe up to 3 mm. long or similar to the leaves but smaller.
    Receptacle
    Receptacle 8–15 mm. long, 3 mm. wide, cylindric, slightly wider at the mouth, glabrous or pubescent outside, with a fimbriately lobed disk, c. 2.5 mm. long at its margin within.
    Calyx
    Sepals 1–1.7 x 0.5–0.7 cm., narrowly oblong-obovate or elliptic, apex rounded to a shortly apiculate apex, pubescent on both sides or glabrous on both sides except at the margins.
    Corolla
    Petals 0.
    Androgynophore
    Androgynophore c. the same length as the receptacle.
    Stamens
    Stamens c. 50; filaments c. 4 cm. long; anthers 2 x 0.75 mm., narrowly oblong, arcuate.
    Ovary
    Ovary on a gynophore up to 4 cm. long, narrowly oblong; ovules c. 80 on 2 placentas.
    Fruits
    Fruit 2–16 x 1 cm., narrowly cylindric, torulose.
    Seeds
    Seeds from several to numerous, c. 6 mm. in diam., subglobose, with a smooth, pale brown testa.
    [FWTA]

    Capparidaceae, Hutchinson and Dalziel. Flora of West Tropical Africa 1:1. 1954

    Habit
    Shrub, or tree to 30 ft. high or more, in savannah
    Flowers
    Flowers white.
    [FTEA]

    Capparidaceae, J. Elffers, R. A. Graham & G. P. Dewolf. Flora of Tropical East Africa. 1964

    Habit
    Shrub or small tree up to about 10 m. tall, often rather rambling, with spreading or drooping branches.
    Stem
    Young twigs pubescent or subglabrous.
    Leaves
    Leaves petiolate, simple; blade ovate, ovate-lanceolate or elliptic, more rarely obovate to suborbicular, 3–10 cm. long, 1.5–5 cm. wide, apically obtuse, emarginate or rounded, sometimes mucronate, basally usually rounded to cordate, glabrous or more rarely pubescent; petiole 1–3 cm. long.
    Inflorescences
    Inflorescence a short corymbose raceme, terminal or on short lateral branches; pedicels 8–12 mm. long.
    Calyx
    Sepals white or pale green, marginally tomentose; receptacle narrowly cylindrical, usually rather long, 5–25 mm. long, often fleshy, ribbed.
    Corolla
    Petals 0.
    Stamens
    Stamens many, variable in number, 15–40 mm. long, yellow.
    Ovary
    Ovary narrowly cylindrical, glabrous.
    Fruits
    Fruits cylindrical, up to 22 cm. long, markedly torulose, green, verrucose, glabrous.
    Habitat
    Deciduous woodland and bushland, scattered-tree grassland and edges of upland rain-forest, also coral-strands; 0–1500 m.
    Distribution
    K1 K4 K5 K6 K7 T1 T2 T3 T4 T5 T6 T7 T8 U1 from Somali Republic (N.) to South Africa (Natal) and west to Senegal

    Images

    Distribution

    Native to:

    Angola, Benin, Botswana, Burkina, Burundi, Cameroon, Caprivi Strip, Central African Repu, Chad, Djibouti, Eritrea, Ethiopia, Gambia, Ghana, Guinea, Kenya, KwaZulu-Natal, Malawi, Mali, Mauritania, Mozambique, Niger, Nigeria, Northern Provinces, Rwanda, Senegal, Somalia, Sudan, Swaziland, Tanzania, Uganda, Zambia, Zaïre, Zimbabwe

    Maerua angolensis DC. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Apr 1, 1992 Eden Foundation [90], Niger K000354980
    Jan 1, 1964 Oldeman, R.A.A. [937], Côte d'Ivoire K000354978
    Jan 1, 1963 Morton, J.K. [4215], Ghana K000354976
    Jan 1, 1957 Miller, O.B. [328], Botswana K000511088
    Jan 1, 1957 Lugard, E.J. [26], Botswana K000511089
    Jan 1, 1957 Lugard, E.J. [28], Botswana K000511090
    Jan 1, 1914 s.coll. [s.n.], Angola K000230721
    Milne-Redhead, E. [5015], Nigeria 16179.000
    Latilo, M.G. [69405], Nigeria K000354979
    Dalziel, J.M. [8122], Gambia K000354975
    Akpabla, G.K. [544], Ghana K000354977
    Pope, G.V. [334], Botswana K000511084
    Hagerup, O. [64], Mali K000354972
    Heudelot, D. [s.n.], Senegal K000354970
    Heudelot, D. [s.n.], Senegal K000354971
    Biegel, H. [4104], Botswana K000511087
    Smith, P.A. [2570], Botswana K000511086
    Smith, P.A. [736], Botswana K000511093
    Hansen, O.J. [3198], Botswana K000511091
    Morwe, F.O. [38], Botswana K000511092
    Harbor, C.C. [6890], Botswana K000511085
    Chevalier, A.J.B. [24], Mali K000354973
    Chevalier, A.J.B. [24], Mali K000354974
    Laferrere, M. [104], Mali K000354969
    Jan 1, 1914 Thomson, T. [s.n.], Yemen Maerua thomsonii K000643966
    Schweinfurth, G. [457], Socotra Maerua thomsonii K000230757

    First published in Prodr. 1: 254 (1824)

    Accepted by

    • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
    • Kalema, J. & Beentje, H. (2012). Conservation checklist of the trees of Uganda: 1-235. Kew Publishing, Royal Botanic Gardens, Kew.
    • Dobignard, A. & Chatelain, C. (2011). Index synonymique de la flore d'Afrique du nord 3: 1-449. Éditions des conservatoire et jardin botaniques, Genève.
    • Lisowski, S. (2009). Flore (Angiospermes) de la République de Guinée Scripta Botanica Belgica 41: 1-517.
    • Timberlake, J.R., Bayliss, J., Alves, T., Francisco, J., Harris, T., Nangoma, D. & de Sousa, C. (2009). Biodiversity and Conservation of Mchese Mountain, Malawi. Report produced under the Darwin Initiative Award 15/036: 1-71. Royal Botanic Gardens, Kew.
    • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
    • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
    • Miller, A.G. & Morris, M. (2004). Ethnoflora of Soqotra Archipelago: 1-759. The Royal Botanic Garden, Edinburgh.
    • Germishuizen, G. & Meyer, N.L. (eds.) (2003). Plants of Southern Africa an annotated checklist Strelitzia 14: 1-1231. National Botanical Institute, Pretoria.
    • Edwards, S., Tadesse, M., Demissew, S. & Hedberg, I. (eds.) (2000). Flora of Ethiopia and Eritrea 2(1): 1-532. The National Herbarium, Addis Ababa University, Ethiopia & The Department of Systematic Botany, Upps.
    • Audru, J., Cesar, J. & Lebrun, J.-P. (1994). Les Plantes Vasculaires de la République de Djibouti. Flore Illustrée 1: 1-336. CIRAD, Départerment d'Elevage et de Médecine vétérinaire, Djibouti.
    • Barry, J. P. & Celles, J.S. (1991). Flore de Mauritanie 1: 1-359. Centre Regional de Documentation Pedagogique, Nice.
    • Lebrun, J.p., Toutain, B., Gaston, A. & Boudet, G. (1991). Catalogue des Plantes Vasculaires du Burkina Faso: 1-341. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
    • Boudet, G., Lebrun, J.P. & Demange, R. (1986). Catalogue des plantes vasculaires du Mali: 1-465. Etudes d'Elevage et de Médecine Vétérinaire des Pays Tropicaux.
    • Boulvert, Y. (1977). Catalogue de la Flore de Centrafrique 3: 1-89. ORSTROM, Bangui.
    • Peyre de Fabregues, B. & Lebrun, J.-P. (1976). Catalogue des Plantes Vascularies du Niger: 1-433. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
    • Lebrun, J.-P., Audru, J., Gaston, A. & Mosnier, M. (1972). Catalogue des Plantes Vasculaires du Tchad Méridional: 1-289. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
    • Lewalle, J. (1970). Liste floristique et répartition altitudinale de la flore du Burundi occidental: 1-84. Université officielle de Bujumbura.
    • Elffers, J., Graham, R.A. & Dewolf, G.P. (1964). Flora of Tropical East Africa, Capparidaceae: 1-88.
    • Exell, A.W. & Wild, H. (eds.) (1960). Flora Zambesiaca 1(1): 1-336. Royal Botanic Gardens, Kew.
    • Hutchinson, J., Dalziel, J.M. & Keay, R.W.J. (1954-1958). Flora of West Tropical Africa, ed. 2, 1: 1-828.
    • Robyns, W. & al. (eds.) (1948-1963). Flore du Congo Belge et du Ruanda-Urundi 1-10.

    Literature

    Flora of West Tropical Africa
    • Aubrév. Fl. For. Soud.-Guln. 55.
    • Chev. Fl. Viv. 1: 165
    • Gilg & Benedict in Engl. Bot. Jahrb. 53: 257
    • —F.T.A. 1: 86
    Kew Backbone Distributions
    • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
    • Kalema, J. & Beentje, H. (2012). Conservation checklist of the trees of Uganda: 1-235. Kew Publishing, Royal Botanic Gardens, Kew.
    • Onana, J.M. (2011). The vascular plants of Cameroon a taxonomic checklist with IUCN assessments: 1-195. National herbarium of Cameroon, Yaoundé.
    • Lisowski, S. (2009). Flore (Angiospermes) de la République de Guinée Scripta Botanica Belgica 41: 1-517.
    • Timberlake, J.R., Bayliss, J., Alves, T., Francisco, J., Harris, T., Nangoma, D. & de Sousa, C. (2009). Biodiversity and Conservation of Mchese Mountain, Malawi. Report produced under the Darwin Initiative Award 15/036: 1-71. Royal Botanic Gardens, Kew.
    • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
    • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
    • Germishuizen, G. & Meyer, N.L. (eds.) (2003). Plants of Southern Africa an annotated checklist Strelitzia 14: 1-1231. National Botanical Institute, Pretoria.
    • Edwards, S., Tadesse, M., Demissew, S. & Hedberg, I. (eds.) (2000). Flora of Ethiopia and Eritrea 2(1): 1-532. The National Herbarium, Addis Ababa University, Ethiopia & The Department of Systematic Botany, Upps.
    • Audru, J., Cesar, J. & Lebrun, J.-P. (1994). Les Plantes Vasculaires de la République de Djibouti. Flore Illustrée 1: 1-336. CIRAD, Départerment d'Elevage et de Médecine vétérinaire, Djibouti.
    • Barry, J. P. & Celles, J.S. (1991). Flore de Mauritanie 1: 1-359. Centre Regional de Documentation Pedagogique, Nice.
    • Lebrun, J.p., Toutain, B., Gaston, A. & Boudet, G. (1991). Catalogue des Plantes Vasculaires du Burkina Faso: 1-341. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
    • Boudet, G., Lebrun, J.P. & Demange, R. (1986). Catalogue des plantes vasculaires du Mali: 1-465. Etudes d'Elevage et de Médecine Vétérinaire des Pays Tropicaux.
    • Troupin, G. (ed.) (1978). Flora du Rwanda 1: 1-413. Musee Royal de l'Afrique Centrale.
    • Boulvert, Y. (1977). Catalogue de la Flore de Centrafrique 3: 1-89. ORSTROM, Bangui.
    • Peyre de Fabregues, B. & Lebrun, J.-P. (1976). Catalogue des Plantes Vascularies du Niger: 1-433. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
    • Lebrun, J.-P., Audru, J., Gaston, A. & Mosnier, M. (1972). Catalogue des Plantes Vasculaires du Tchad Méridional: 1-289. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
    • Lewalle, J. (1970). Liste floristique et répartition altitudinale de la flore du Burundi occidental: 1-84. Université officielle de Bujumbura.
    • Elffers, J., Graham, R.A. & Dewolf, G.P. (1964). Flora of Tropical East Africa, Capparidaceae: 1-88.
    • Exell, A.W. & Wild, H. (eds.) (1960). Flora Zambesiaca 1(1): 1-336. Royal Botanic Gardens, Kew.
    • Hutchinson, J., Dalziel, J.M. & Keay, R.W.J. (1954-1958). Flora of West Tropical Africa, ed. 2, 1: 1-828.
    • Robyns, W. & al. (eds.) (1948-1963). Flore du Congo Belge et du Ruanda-Urundi 1-10.
    Flora of Tropical East Africa
    • F. White, Forest Flora of Northern Rhodesia p. 64 (1962).
    • Dale & Greenway, Kenya Trees and Shrubs p. 125, fig. 24 (1961).
    • Wild in Flora Zambesiaca 1: 228, t. 35/F (1960).
    • Keay, Flora of West Tropical Africa, ed. 2, 1: 88 (1954).
    • Milne-Redh, in Mem. N.Y. Bot. Gard. 8: 217 (1953).
    • W.J. Eggeling, Indigenous Trees of the Uganda Protectorate, ed. 2: 75, fig. 19 (1952).
    • Hauman & Wilczek in Flore du Congo Belge et du Ruanda-Urundi, 2: 487 (1951).
    • F. W. Andr., Fl. Pl. A.-E. Sudan 1: 52, fig. 39 (1950).
    • J.P.M. Brenan, Check-lists of the Forest Trees and Shrubs of the British Empire no. 5, part II, Tanganyika Territory p. 118 (1949).
    • Robyns, Flore des Spermatophytes du Parc National Albert 1: 208 (1948).
    • Exell & Mendonça in Consp. Fl. Angol. 1: 59 (1937).
    • Battiscombe & Dale, Trees and Shrubs of Kenya Colony p. 10 (1936).
    • Chiov., Fl. Somala 2: 7 (1932).
    • Gilg in A. Engler & O. Drude, Die Vegetation Der Erde, IX, Pflanzenwelt Afrikas 3(1): 253, fig. 167/A–D (1915).
    • Gilg & Bene. in A. Engler, Botanische Jahrbücher für Systematik, Pflanzengeschichte und Pflanzengeographie 53: 257 (1915).
    • Gilg in Die Pflanzenwelt Ost-Afrikas und der Nachbargebiete, Theile C: 187 (1895).
    • Oliv. in Flora of Tropical Africa 1: 86 (1868).
    • DC., Prodr. 1: 254 (1824).

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of West Tropical Africa
    Flora of West Tropical Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2018. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2018. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0