1. Family: Phyllanthaceae Martinov
    1. Cleistanthus Hook.f. ex Planch.

      1. This genus is accepted, and its native range is Tropical & S. Africa, Madagascar, Tropical & Subtropical Asia to W. Pacific.

    [FZ]

    Euphorbiaceae, A. Radcliffe-Smith. Flora Zambesiaca 9:4. 1996

    Habit
    Monoecious or more rarely dioecious shrubs or trees.
    Indumentum
    Indumentum simple.
    Leaves
    Leaves alternate, petiolate, stipulate, simple, entire, penninerved.
    Inflorescences
    Inflorescences axillary, rarely terminal, fasciculate or pesudo-racemose, uni- or bisexual, bracteate; bracts soon falling.
    Flowers
    Male flowers: sepals (4)5(6), valvate; petals (4)5(6), minute; disk ± annular or cupular, ± entire; stamens (4)5(6), filaments united below into a staminal column, free and spreading above, anthers basifixed, introrse, longitudinally dehiscent; pistillode (non-functional ovary) 3-lobed to tripartite. Female flowers: sepals and petals ± as in the male; disk annular, hypogynous or cupuliform, enveloping the ovary; ovary (2)3(4)-locular, ovules 2 per locule; styles (2)3(4), free or united at the base, 1–4 times bifid. Flowers proterogynous, pedicellate, fragrant.
    Fruits
    Fruit (2)3(4)-lobed, septicidally dehiscent into as many bivalved cocci; endocarp woody, separating from the pericarp; columella persistent, apically dilated.
    Seeds
    Seeds (1)2 per coccus, ecarunculate; endosperm copious or scanty; cotyledons broad, thin or thick and fleshy, flat or folded.
    [FTEA]

    Euphorbiaceae, A. R.-Smith. Flora of Tropical East Africa. 1987

    Habit
    Monoecious or more rarely dioecious shrubs or trees with a simple indumentum
    Leaves
    Leaves alternate, petiolate, stipulate, simple, entire, penninerved
    Inflorescences
    Inflorescences axillary, rarely terminal, glomerulate, fasciculate or pseudospicate to pseudoracemose, bracteate, the bracts soon caducous, unisexual or bisexual
    Flowers
    Flowers pedicellate Male flowers: sepals (4–)5(–6), valvate, or slightly imbricate at the apex; petals (4–)5(–6), minute; disc pulvinate, annular or cupuliform, entire or sinuately lobed; stamens (4–)5(–6), united at the base into a staminal column, with the anthers longitudinally dehiscent; pistillode trilobate or tripartite, the lobes adherent Female flowers: sepals and petals ± as in the ♂ flowers; outer disc annular, inner disc annular or cupuliform and partially or completely enveloping the ovary; ovary (2–)3(–4)-locular, with 2 ovules per locule; styles (2–)3(–4), free or united at the base, 1–4 times bilobed or bifid
    Male
    Male flowers: sepals (4–)5(–6), valvate, or slightly imbricate at the apex; petals (4–)5(–6), minute; disc pulvinate, annular or cupuliform, entire or sinuately lobed; stamens (4–)5(–6), united at the base into a staminal column, with the anthers longitudinally dehiscent; pistillode trilobate or tripartite, the lobes adherent
    Female
    Female flowers: sepals and petals ± as in the ♂ flowers; outer disc annular, inner disc annular or cupuliform and partially or completely enveloping the ovary; ovary (2–)3(–4)-locular, with 2 ovules per locule; styles (2–)3(–4), free or united at the base, 1–4 times bilobed or bifid
    Fruits
    Fruit (2–)3(–4)-lobed, dehiscing into (2–)3(–4) bivalved cocci; endocarp woody, separating from the pericarp at maturity; columella persistent
    Seeds
    Seeds (1–)2 per coccus; albumen scanty or copious; cotyledons thick and fleshy, thin and flat or thin and plicate.

    Images

    Distribution

    Native to:

    Andaman Is., Angola, Assam, Bangladesh, Bismarck Archipelago, Borneo, Burundi, Cabinda, Cambodia, Cameroon, Caroline Is., Central African Repu, China South-Central, China Southeast, Comoros, Congo, Equatorial Guinea, Fiji, Gabon, Ghana, Gulf of Guinea Is., Hainan, India, Ivory Coast, Jawa, Kenya, KwaZulu-Natal, Lesser Sunda Is., Liberia, Madagascar, Malawi, Malaya, Maluku, Mozambique, Myanmar, New Caledonia, New Guinea, New South Wales, Nicobar Is., Nigeria, Northern Provinces, Northern Territory, Pakistan, Philippines, Queensland, Rwanda, Sierra Leone, Solomon Is., Sri Lanka, Sulawesi, Sumatera, Tanzania, Thailand, Uganda, Vietnam, Zambia, Zaïre, Zimbabwe

    Cleistanthus Hook.f. ex Planch. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Jan 1, 1994 Tchouto (Mbatchou), P. [853], Cameroon K000540097
    Jan 1, 1994 Tchouto (Mbatchou), P. [818], Cameroon K000540098

    First published in Hooker's Icon. Pl. 8: t. 779 (1848)

    Accepted by

    • Govaerts, R., Frodin, D.G. & Radcliffe-Smith, A. (2000). World Checklist and Bibliography of Euphorbiaceae (and Pandaceae) 1-4: 1-1622. The Board of Trustees of the Royal Botanic Gardens, Kew.
    • Govaerts, R. (1999). World Checklist of Seed Plants 3(1, 2a & 2b): 1-1532. MIM, Deurne.

    Literature

    Flora of West Tropical Africa
    • Jablonszky in Engl. Pflanzenr. Euph. 8: 8 (1915).
    • —F.T.A. 6, 1: 621
    Flora Zambesiaca
    • Jablonszky in Engler, Pflanzenr. [IV, fam. 147, viii] 65: 8 (1915).
    • Müller Argoviensis in De Candolle, Prodr. 15, 2: 503 (1866).
    • in Hooker’s Icon. Pl. 8: t. 779 (1848).
    Flora of Tropical East Africa
    • in F.C.B. 8(1): 5 (1962)
    • J. Léon in B.J.B.B. 30: 421 (1960)
    • Jabl. in E.P. IV. 147(8): 8 (1915)
    • Hutch. in F.T.A. 6(1): 621 (1912)
    • G.P. 3(1): 268 (1880)
    • Muell. Arg. in DC., Prodr. 15(2): 503 (1866)
    • in Hook., Ic. Pl. 8, t. 779 (1848)

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2018. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2018. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0