1. Family: Phyllanthaceae Martinov
    1. Heywoodia Sim

      1. This genus is accepted, and its native range is Kenya to S. Africa.

    [FTEA]

    Euphorbiaceae, A. R.-Smith. Flora of Tropical East Africa. 1987

    Habit
    Dioecious evergreen tree lacking indumentum
    Leaves
    Leaves alternate, petiolate, peltate in the seedling stage and on sucker-shoots, otherwise basally attached, stipulate, simple, entire, symmetrical at the base, coriaceous, penninerved
    Flowers
    Flowers fasciculate, the males densely so, and sessile, the females in 1–3-flowered cymes and shortly pedicellate, axillary Male flowers: bracts 4, free, imbricate, broad, concave, unequal; sepals 3, free, imbricate, broad, concave, unequal, in size and texture intermediate between the bracts and the petals; petals 5, free, imbricate, concave, subequal; disc peripheral, crisped-lobulate, invaginated amongst the filaments of the outer whorl of stamens, fleshy; stamens (8–)10–11(–12), in 2 whorls, filaments free, anthers erect, slightly recurved, dorsifixed, introrse, thecae parallel, longitudinally dehiscent; pistillode minute, trifid Female flowers: bracts, sepals and petals ± as in the ♂; 1–2 bracteoles also present; disc ± annular, slightly lobulate, shallowly cupular; staminodes 6–8, filiform; ovary 4–5-locular, with 2 ovules per locule; stigmas 4–5, sessile, bilobed
    Male
    Male flowers: bracts 4, free, imbricate, broad, concave, unequal; sepals 3, free, imbricate, broad, concave, unequal, in size and texture intermediate between the bracts and the petals; petals 5, free, imbricate, concave, subequal; disc peripheral, crisped-lobulate, invaginated amongst the filaments of the outer whorl of stamens, fleshy; stamens (8–)10–11(–12), in 2 whorls, filaments free, anthers erect, slightly recurved, dorsifixed, introrse, thecae parallel, longitudinally dehiscent; pistillode minute, trifid
    Female
    Female flowers: bracts, sepals and petals ± as in the ♂; 1–2 bracteoles also present; disc ± annular, slightly lobulate, shallowly cupular; staminodes 6–8, filiform; ovary 4–5-locular, with 2 ovules per locule; stigmas 4–5, sessile, bilobed
    Fruits
    Fruit slightly depressed-subglobose, shallowly 8–10-lobed and 4–5-ridged, tardily dehiscent, leaving a columella; pericarp thinly fleshy, smooth, becoming rugulose when dried; endocarp bony
    Seeds
    Seeds solitary per locule by abortion, ecarunculate, obliquely ovoid, rounded-triangular in section, ± smooth; endosperm papery; embryo minute, cotyledons broad, flat.
    [FZ]

    Euphorbiaceae, A. Radcliffe-Smith. Flora Zambesiaca 9:4. 1996

    Habit
    Dioecious, completely glabrous evergreen tree.
    Leaves
    Leaves alternate, petiolate, sometimes peltate in juvenile foliage, stipulate; blades simple, entire, basally symmetrical, coriaceous, penninerved.
    Flowers
    Male flowers: sepals 3, free, imbricate, concave, unequal; petals 5, free, imbricate, concave, subequal; disk extrastraminal, lobulate, invaginated amongst the outer stamens, fleshy; stamens (8)10–11(12), in 2 whorls, filaments free, anthers dorsifixed, introrse, thecae parallel, longitudinally dehiscent; pistillode (non-functional ovary) minute, trifid. Female flowers: bracteoles 1–2, sepals and petals similar to those of the male; disk annular, slightly lobulate, shallowly cupular; staminodes 6–8, filiform; ovary 4–5-locular, with 2 ovules per loculus; stigmas 4–5, sessile, 2-lobed. Flowers axillary, fasciculate, the males densely so, sessile, bracteate, the females in 1–3-flowered cymes, shortly pedicellate; bracts 4, free, imbricate, concave, unequal.
    Fruits
    Fruit depressed-subglobose, shallowly 8–10-lobed, tardily septicidally dehiscent, leaving a columella; pericarp thinly fleshy, smooth, becoming rugulose on drying; endocarp bony.
    Seeds
    Seeds solitary per loculus by abortion, ecarunculate; endosperm papery; embryo minute; cotyledons broad, flat.

    Distribution

    Native to:

    Cape Provinces, Kenya, KwaZulu-Natal, Mozambique, Swaziland, Tanzania, Uganda

    Heywoodia Sim appears in other Kew resources:

    First published in Forest Fl. Cape: 326 (1907)

    Accepted by

    • Govaerts, R., Frodin, D.G. & Radcliffe-Smith, A. (2000). World Checklist and Bibliography of Euphorbiaceae (and Pandaceae) 1-4: 1-1622. The Board of Trustees of the Royal Botanic Gardens, Kew.

    Literature

    Flora Zambesiaca
    • Milne-Redhead in Bull. Jard. Bot. État 27: 329, f. 32 & t. 10 (1957).
    • For. Fl. Col. Cape Good Hope: 326, t. 140, fig.1 (1907).
    Flora of Tropical East Africa
    • Milne-Redh. in B.J.B.B. 27: 329, fig. 32 (map) & t. 10 (1957)
    • in K.B. 1922: 115 (1922), descr. emend. et ampl.
    • Pax in E.P. IV. 147(15): 280 (1922)
    • Hutch. in Fl. Cap. 5(2): 384 (1915)
    • For. Fl. Cape Col.: 326, t. 140/1 (1907)

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0