1. Family: Phyllanthaceae Martinov
    1. Uapaca Baill.

      1. This genus is accepted, and its native range is Tropical Africa, Madagascar.

    [FZ]

    Euphorbiaceae, A. Radcliffe-Smith. Flora Zambesiaca 9:4. 1996

    Habit
    Dioecious trees or shrubs.
    Indumentum
    Indumentum simple and/or minutely pseudolepidote.
    Stem
    Trunks occasionally stilt-rooted. Twigs stout, often with pronounced leaf scars, producing an exudate when cut, which hardens and darkens on drying.
    Leaves
    Leaves alternate, usually crowded towards the end of the twigs, petiolate or subsessile, stipulate or not; blades simple, often obovate, entire, penninerved.
    Inflorescences
    Inflorescences axillary, or borne on older wood, solitary or fasciculate, pedunculate, with a whorl of 5–12 involucrate imbricate tepaloid bracts surrounding the flowers; male inflorescences many-flowered, the flowers in dense globose capitula; female inflorescences 1-flowered.
    Flowers
    Male flowers sessile; calyx campanulate or turbinate, truncate, dentate or irregularly or regularly lobed, the lobes imbricate; petals absent; disk absent; stamens (4)5(6), free, episepalous, anthers erect, subbasifixed, introrse, thecae parallel, longitudinally dehiscent; pistillode cylindric-obconic, infundibuliform, hypocrateriform, pileiform or sometimes lobed. Female flowers sessile; calyx minute, truncate, sinuate or lobed, disciform; petals absent; disk absent; ovary (2)3(5)-locular, with 2 ovules per loculus; styles (2)3(5), free, thick, recurved, covering the ovary, multipartite to laciniate.
    Fruits
    Fruit drupaceous, indehiscent; mesocarp spongy; pyrenes (2)3(4), dorsally carinate and bisulcate, indurate, tardily bivalved.
    Seeds
    Seeds mostly 1 per pyrene, compressed, ecarunculate; endosperm fleshy; embryo straight; cotyledons broad, flat, green.
    [FTEA]

    Euphorbiaceae, A. R.-Smith. Flora of Tropical East Africa. 1987

    Habit
    Dioecious pachycaul vernicifluous trees or shrubs with a simple and/or lepidote indumentum; trunks often with buttress-roots; twigs often strongly marked with leaf-scars
    Leaves
    Leaves alternate, usually crowded towards the ends of the branches, petiolate or subsessile, stipulate or not, simple, entire, penninerved
    Inflorescences
    Inflorescences axillary, solitary or fasciculate, pedunculate, with a whorl of 5–10 imbricate involucrate tepaloid bracts surrounding the flowers; ♂ inflorescences many-flowered, the flowers in dense globose capitula; ♀ inflorescences 1-flowered
    Flowers
    Male flowers sessile; calyx campanulate or turbinate, truncate, dentate, irregularly split or regularly 5–more-lobed, the lobes imbricate; petals 0; disc 0; stamens (4–)5(–6), free, episepalous, anthers erect, basifixed, introrse, thecae parallel, longitudinally dehiscent; pistillode cylindric-obconic, and often infundibuliform, hypocrateriform, or pileiform and sometimes lobate Female flowers sessile; calyx minute, truncate or sinuate, disc-like; petals 0; disc 0; ovary (2–)3(–5)-locular, with 2 ovules per locule; styles (2–)3(–5), free, thick, recurved, enveloping the ovary, multipartite or laciniate
    Male
    Male flowers sessile; calyx campanulate or turbinate, truncate, dentate, irregularly split or regularly 5–more-lobed, the lobes imbricate; petals 0; disc 0; stamens (4–)5(–6), free, episepalous, anthers erect, basifixed, introrse, thecae parallel, longitudinally dehiscent; pistillode cylindric-obconic, and often infundibuliform, hypocrateriform, or pileiform and sometimes lobate
    Female
    Female flowers sessile; calyx minute, truncate or sinuate, disc-like; petals 0; disc 0; ovary (2–)3(–5)-locular, with 2 ovules per locule; styles (2–)3(–5), free, thick, recurved, enveloping the ovary, multipartite or laciniate
    Fruits
    Fruit drupaceous, indehiscent; mesocarp spongy; pyrenes (2–) 3(–4), dorsally bisulcate, indurate
    Seeds
    Seeds mostly 1 per pyrene, compressed, ecarunculate; endosperm fleshy; cotyledons broad, flat.

    Images

    Distribution

    Native to:

    Angola, Benin, Burkina, Burundi, Cabinda, Cameroon, Central African Repu, Chad, Congo, Equatorial Guinea, Gabon, Ghana, Guinea, Guinea-Bissau, Gulf of Guinea Is., Ivory Coast, Kenya, Liberia, Madagascar, Malawi, Mali, Mozambique, Nigeria, Senegal, Sierra Leone, Sudan, Tanzania, Togo, Uganda, Zambia, Zaïre, Zimbabwe

    Uapaca Baill. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Jun 1, 2010 Cheek, M. [11998], Cameroon K000678386
    Troupin, G. [3432], Congo 15847.000
    Etuge, M. [73], Cameroon K000180781

    First published in Étude Euphorb.: 595 (1858)

    Accepted by

    • Breteler, F.J. (2013). Uapaca (Phyllanthaceae) in the Guineo-Congolian forest region: a synoptic revision Plant ecology and evolution 146: 75-94.
    • Breteler, F.J. (2012). Flore du Gabon 43: 1-107. Muséum National D'Histoire Naturelle, Paris.
    • McPherson, G. (2011). A review of Madagascan Uapaca (Euphorbiaceae s.l.) Adansonia, III, 33: 221-231.
    • Govaerts, R., Frodin, D.G. & Radcliffe-Smith, A. (2000). World Checklist and Bibliography of Euphorbiaceae (and Pandaceae) 1-4: 1-1622. The Board of Trustees of the Royal Botanic Gardens, Kew.

    Literature

    Flora of West Tropical Africa
    • Pax & K. Hoffm. in Engl. Pflanzenr. Euph. 15: 298–311 (1922).
    • —F.T.A. 6, 1: 634
    Flora Zambesiaca
    • De Wildeman, Contrib. Ét. Esp. Gen. Uapaca (1936).
    • Müller Argoviensis in De Candolle, Prodr. 15, 2: 489 (1866).
    • Étud. Gén. Euphorb.: 595 (1858).
    Flora of Tropical East Africa
    • De Wild, Contrib. Étud. Esp. Uapaca (1936)
    • Pax & K. Hoffm. in E.P. IV. 147 (15): 298 (1922)
    • Hutch. in F.T.A. 6(1): 634 (1912)
    • G.P. 3 (1): 282 (1880)
    • Muell. Arg. in DC., Prodr. 15(2): 489 (1866)
    • Étud. Gén. Euph.: 595 (1858)

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2018. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2018. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0