1. Family: Combretaceae R.Br.
    1. Genus: Combretum Loefl.
      1. Combretum hereroense Schinz

        This species is accepted, and its native range is Kenya to S. Africa.

    [FZ]

    Combretaceae, A. W. Exell. Flora Zambesiaca 4. 1978

    Habit
    Small tree up to 9(12) m. high or more rarely a shrub, often with arching stems, frequently flowering before the leaves; crown dense, irregular; bark creamy-grey to grey-black, fissured; branchlets usually tomentose to densely pubescent when young but often quickly glabrescent.
    Leaves
    Leaves opposite or subopposite, often borne on short shoots; lamina usually c. 2–7 x 1–4·5 cm., from very narrowly elliptic to very broadly obovate or subcircular (especially on sucker shoots when they may be as much as 5·5 cm. broad and broader than long), from densely tomentose to nearly glabrous (except for scales), densely golden-(rarely silvery-)lepidote beneath, scales usually contiguous but sometimes hidden by the indumentum, apex sometimes acute but often rounded or even retuse, base cuneate to rounded; lateral nerves usually only 3–4(5) pairs, prominent beneath; petiole up to 5 mm. long.
    Inflorescences
    Inflorescences of short rather congested spikes up to c. 3 cm. long, occasionally branched, axillary, often appearing on leafless branchlets in the axils of the scars of the fallen leaves; rhachis tomentose to glabrous (except for scales); bracts 3 mm. long, filiform, often somewhat persistent.
    Flowers
    Flowers sessile, yellow.
    Receptacle
    Lower receptacle 2·3–3 mm. long; upper receptacle 2·5–3 x 2–3 mm., campanulate or shortly infundibuliform, lepidote and otherwise pubescent to nearly glabrous.
    Calyx
    Sepals 1–1·5 x l–1·5 mm., deltate.
    Corolla
    Petals 1·5–2·5 x 1–2·5 mm., broadly to very broadly obovate or subcircular or very broadly transversely elliptic, emarginate, shortly unguiculate, glabrous.
    Stamens
    Stamen-filaments 3–1·5 mm. long; anthers 0·4–0·6 mm. long.
    Disc
    Disk c. 2 mm. in diam. with pilose margin free for 0·7–0·8 mm.
    Style
    Style 3·5–4·5 mm. long, not expanded at the apex.
    Fruits
    Fruit usually c. 2 x 2 cm. but sometimes smaller and occasionally up to 3·5 x 3·5 cm., subcircular in outline, densely reddish- or golden-lepidote, apical peg absent or very short, wings c. 5(10) mm. wide, subcoriaceous, stipe up to 11 mm. long.
    Cotyledons
    Cotyledons 2 1·5 x 2·5 cm., transversely elliptic, arising above soil level; petioles 10–11 mm. long.
    Scales
    Scales irregularly undulate in outline, marginal cells 8–12 mostly retusely scalloped, delimited by usually 8 radial walls, 8 tangential walls and sometimes a few extra radial walls.
    [FTEA]

    Combretaceae, G. E. Wickens. Flora of Tropical East Africa. 1973

    Habit
    Small tree up to 8 (–12) m. high or more frequently (in E. Africa) a much branched coppicing shrub up to 5 m. high.
    Bark
    Bark slightly fibrous, grey-brown to blackish; branchlets reddish brown to light grey, fibrous; young shoots tomentose to densely pubescent, later becoming glabrescent, or glabrous except for the ± contiguous rufous scales.
    Shoots
    Short horizontal spur shoots usually present.
    Leaves
    Leaves opposite or subopposite, usually borne along the length of the spur shoots but tending to cluster towards the apex; lamina coriaceous, narrowly elliptic to broadly obovate or subcircular (especially on sucker shoots), 3–6.5(–8) cm. long, l.5–4(–4.5) cm. wide, apex slightly refuse to rounded or acute, mucronate, base cuneate to rounded, from densely tomentose to glabrous (except for the scales); scales silvery or golden and due to their wavy margins may present a rather scurfy appearance, contiguous to scattered, sometimes appearing impressed in subsp. volkensii; lateral nerves 3–9 pairs, subprominent to prominent beneath, visible above; petiole up to 5 mm. long.
    Inflorescences
    Inflorescence of short, sometimes rather congested spikes up to 3 cm. long, axillary, occasionally branched, often appearing on leafless branches (especially on spur shoots) in the axils of scars of fallen leaves; peduncles ± 0.7 cm. long.
    Flowers
    Flowers (fig. 3/20, p. 14) pale yellow to yellow, fragrant.
    Receptacle
    Lower receptacle 1.5 mm. long; upper receptacle campanulate to shortly infundibuliform, 2–3 mm. long and wide, lepidote and otherwise densely pubescent to nearly glabrous.
    Calyx
    Sepals deltate.
    Corolla
    Petals spathulate to very broadly obovate or subcircular, 1.5–2.5 mm. long, 1–2.5 mm. wide, emarginate, clawed, glabrous.
    Stamens
    Stamen-filaments 2–4.5 mm. long; anthers 0.4–0.6 mm. long.
    Disc
    Disk with a pilose margin free for 0.7–0.8 mm.
    Style
    Style 3–4 mm. long.
    Fruits
    Fruit (fig. 5/20, p. 16) oval to subcircular in outline, up to 3.5 cm. long and wide, often much smaller, glabrous except for the scales; apical peg absent or very short; stipe up to 11 mm. long.
    Germination
    Cotyledons with petioles 10–11 mm. long; germination epigeal.
    Scales
    Scales (fig. 2/20, p. 13) irregularly undulate in outline, (50–)75–100 μ in diameter, marginal cells 8–12, mostly retusely scalloped.
    Habitat
    (of species as a whole). Wooded grassland and Acacia, Commiphora bushland, sometimes dominant; 50–2700 m.
    Distribution
    K1 K2 K3 K4 K7 T1 T2 T3 T5 T6 T7 T8 U1 U3 Sudan and Somali Republic (S.), southwards to Lesotho and Natal and west to Angola

    Images

    Distribution

    Native to:

    Angola, Botswana, Caprivi Strip, Kenya, KwaZulu-Natal, Malawi, Mozambique, Namibia, Northern Provinces, Swaziland, Tanzania, Zambia, Zimbabwe

    Common Names

    English
    Mouse-eared combretum, Russet bush willow

    Combretum hereroense Schinz appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Sep 23, 2008 Crawford, F. [57], Namibia K000450441
    Aug 28, 1969 Bolus, H. [7769], South Africa K000226631 syntype
    Aug 26, 1969 Bolus, H. [7763], South Africa K000226630 holotype
    Aug 26, 1969 Bolus, H. [7768], South Africa K000226632 syntype
    Jan 1, 1968 Harbor, C.C. [6609], Botswana K000489077
    Jan 1, 1967 Lugard, E.J. [9], Botswana K000489078
    Jan 1, 1967 Menyhart [s.n.] K000226511
    Jan 1, 1967 Stony, R. [5034], Botswana K000489080
    Jan 1, 1961 Engler, A. [6217], Namibia K000226635
    Brenan, J.P.M. [8274], Tanzania 25155.000
    Ash [1920], Ethiopia 35554.000
    Leach, L.C. [12503], Botswana K000489084
    Galpin, E.E. [561], South Africa K000226625
    Galpin, E.E. [560], South Africa K000226626 Unknown type material
    Galpin, E.E. [756], South Africa K000226627 Unknown type material
    Galpin, E.E. [756], South Africa K000226628
    Pope, G.V. [397], Botswana K000489083
    Smith, P. [82], Botswana K000489086
    Schlechter [11772], South Africa K000226629
    Bolus, H. [11017], South Africa K000226633 paratype
    Legat, C.E. [66], South Africa K000226634 paratype
    Biegel, H. [3887], Botswana K000489079
    Hansen, O.J. [3034], Botswana K000489082
    Brown, R.C. [6609], Botswana K000489085
    Allen, A. [266], Botswana K000489081

    First published in Verh. Bot. Vereins Prov. Brandenburg 30: 245 (1888)

    Accepted by

    • Mannheimer, C.A. & Curtis, B.A. (eds.) (2009). Le Roux and Müller's field guide to the trees and shrubs of Namibia, rev. ed.: 1-525. Macmillan Education Namibia, Windhoek.
    • Germishuizen, G. & Meyer, N.L. (eds.) (2003). Plants of Southern Africa an annotated checklist Strelitzia 14: 1-1231. National Botanical Institute, Pretoria.
    • Govaerts, R. (1999). World Checklist of Seed Plants 3(1, 2a & 2b): 1-1532. MIM, Deurne.
    • Edwards, S., Tadesse, M. & Hedberg, I. (eds.) (1995). Flora of Ethiopia and Eritrea 2(2): 1-456. The National Herbarium, Addis Ababa University, Ethiopia & The Department of Systematic Botany, Upps.
    • Launert, E. (ed.) (1978). Flora Zambesiaca 4: 1-658. Royal Botanic Gardens, Kew.
    • Wickens, G.E. (1973). Flora of Tropical East Africa, Combretaceae: 1-99.

    Literature

    Kew Backbone Distributions
    • Wickens, G.E. (1973). Flora of Tropical East Africa, Combretaceae: 1-99.
    Flora of Tropical East Africa
    • Wickens in Kew Bulletin 25: 413 (1971).
    • Exell in Kirkia 7: 199 (1970).
    • F. White, Forest Flora of Northern Rhodesia p. 285 (1962).
    • Engl. & Diels in Monographieen Afrikanischer Pflanzen-Familien und Gattungen 3: 63 (1899).
    • Schinz in Verh. Bot. Ver. Brand. 30: 245 (1888).

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2018. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2018. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Plants and People Africa
    Common Names from Plants and People Africa http://www.plantsandpeopleafrica.com/
    © Plants and People Africa http://www.plantsandpeopleafrica.com http://creativecommons.org/licenses/by/3.0/