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  1. Family: Aspleniaceae Newman
    1. Genus: Asplenium L.
      1. Asplenium warneckii Hieron.

        This species is accepted, and its native range is Ghana, Cameroon, Tanzania, Madagascar.


    Aspleniaceae, Henk Beentje. Flora of Tropical East Africa. 2008

    Type: Tanzania, Usambara, Amani, Warnecke 338 (B!, holo.; B!, K!, P!, iso.)
    Terrestrial or epiphyte.
    Rhizome short-creeping or short-ascending, to 5 mm thick, with black narrowly triangular rhizome scales to 3.5 × 0.4–0.5 mm, with very thin brown margin, attenuate into hair-tip.
    Fronds almost tufted to shortly spaced, 30–80 cm long, not proliferous (rarely proliferous near apex).
    Stipe grey or black, green on upper surface near apex, 10–35 cm long, at base with scales like rhizome scales, otherwise with brown sparse scales to 2.5 mm.
    Leaf lamina
    Lamina oblong, 20–45 × 8–18 cm, 1-pinnate, the lower pinnae rarely sub-3-lobed or lobed at base, the terminal pinna often 3-lobed and acuminate.
    Pinnae in 5–9 pairs, patent or spreading, (sub)opposite, chartaceous, glaucous green, shortly stalked, inaequilateral, trapezo-rhomboid, to 10 × 4.8 cm, acroscopic base subtruncate-cuneate, basiscopic base cuneate, margin irregularly serrate-crenate, glabrous or nearly so.
    Rachis black with thin green lines connecting pinna bases, with dense scales when young, later with scattered subulate scales.
    Sori 9–18 per pinna, on the veins at a very slight angle to costa, 0.5–3 cm long, not reaching margin.
    Indusium membranous, entire, 0.7–0.8 mm wide.
    Fig. 4: 3, p. 25.
    Moist forest near water; 900–1350 m
    Widespread; least concern (LC)
    I have made var. prolifera into a synonym, though Hieronymus felt it differed in stipe colour, dissection of pinnae margins, and being proliferous near the apex; as the type is sterile and lacks basal parts, it is not possible to be entirely certain, but I believe this is only a form. Two other specimens are proliferous: Uganda, Mengo District: Kipayo, Jan. 1914, Dummer 585! & 15 km from Kampala on Mascha road, May 1937, Chandler 1625! A specimen from T 3, Zaraninge forest, collected in 1990 by Frontier Tanzania 1099! (MO) looks superficially like this taxon, but is from 300 m altitude, has longer rhizome scales and narrower pinnae. It might represent a new taxon. Very close to A. hemitomum, from West and central Africa, but distinct in the size of the rhizome scales (longer in hemitomum), the texture of the leaf (more leathery in hemitomum), and spores: irregularly reticulate in hemitomum, crested in warneckei. This taxon ( warneckei) also looks like A. macrophyllum from Asia, and Peter cited his number 10000 (cited above) as such, but warneckei has consistently wider pinnae and a different feel to it. I prefer to keep it separate.
    Range: Ghana, Cameroon Flora districts: U2 U4 T1 T3 T6 T7



    Native to:

    Cameroon, Ghana, Madagascar, Tanzania

    Other Data

    Asplenium warneckii Hieron. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Aug 1, 2001 Cable, S. [3621], Cameroon K000108934
    Warnecke, O. [338], Tanzania K000804576 isotype


    First published in Bot. Jahrb. Syst. 46: 367 (1911)

    Accepted by

    • Roskov Y. & al. (eds.) (2018). World Ferns: Checklist of Ferns and Lycophytes of the World Species 2000 & ITIS Catalogue of Life Naturalis, Leiden, the Netherlands.


    Flora of West Tropical Africa

    • in Engl. Bot. Jahrb. 46 : 367 (1911).

    Flora of Tropical East Africa

    • Johns, Pterid. trop. East Africa checklist: 68 (1991).
    • Tardieu, Fl. Cameroun Pteridophytes: 204, t. 32/6–7 (1964)
    • Alston, Ferns W.T.A.: 59 (1959)
    • E.J. 46: 367 (1911)


    Flora of Tropical East Africa
    Flora of Tropical East Africa

    Herbarium Catalogue Specimens
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    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2020. Published on the Internet at and
    © Copyright 2017 World Checklist of Selected Plant Families.

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2020. Published on the Internet at and
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families.