1. Family: Aspleniaceae Newman
    1. Genus: Thelypteris Schmidel
      1. Thelypteris heterocarpos (Blume) Morton

        This species is accepted, and its native range is SE. China to Indo-China and Pacific.

    Distribution

    Native to:

    Borneo, Cambodia, Caroline Is., China Southeast, Fiji, Hainan, Jawa, Laos, Lesser Sunda Is., Malaya, Maluku, Marianas, Marquesas, New Guinea, Philippines, Queensland, Samoa, Solomon Is., Sulawesi, Sumatera, Thailand, Vanuatu, Vietnam

    Thelypteris heterocarpos (Blume) Morton appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Mar 1, 2019 Schomburgk, R.H. [30], Singapore Sphaerostephanos heterocarpos K000117141
    Mar 1, 2019 Lee, D.W. [UL21], Malaysia Sphaerostephanos heterocarpos K000117120
    Mar 1, 2019 Pestama, J.L. [21125], Singapore Sphaerostephanos heterocarpos K000117140
    Feb 1, 2006 Beaman, J.H. [18950], Sabah Sphaerostephanos heterocarpos K001053157
    Dec 1, 2004 Miguel, G. [s.n.], Indonesia Sphaerostephanos heterocarpos K000117458
    Sep 10, 1992 Cook, M.A. [4], Brunei Sphaerostephanos heterocarpos K000408126
    Sep 10, 1992 Cook, M.A. [4], Brunei Sphaerostephanos heterocarpos K000408127
    Sep 10, 1992 Cook, M.A. [4], Brunei Sphaerostephanos heterocarpos K000408128
    Jan 1, 1986 Parris, B.S. [10716], Malaysia Sphaerostephanos heterocarpos K000397505
    Jan 1, 1986 Beaman, J.H. [10725], Malaysia Sphaerostephanos heterocarpos K000397502
    Jan 1, 1986 Beaman, J.H. [10725], Malaysia Sphaerostephanos heterocarpos K000397503
    Jan 1, 1986 Ridsdale, C.E. [5330], Philippines Sphaerostephanos heterocarpos K000399069
    Jan 1, 1986 Ridsdale, C.E. [5330], Philippines Sphaerostephanos heterocarpos K000399070
    Jan 1, 1986 Ridsdale, C.E. [5330], Philippines Sphaerostephanos heterocarpos K000399071
    Jan 1, 1982 Kerenga, K. [LAE73924], Papua New Guinea Sphaerostephanos heterocarpos K000422448
    May 11, 1979 Parris, B.S. [6652], Malaysia Sphaerostephanos heterocarpos K000117488
    May 11, 1979 Parris, B.S. [6652], Malaysia Sphaerostephanos heterocarpos K000117489
    Feb 23, 1979 Lee, D.W. [UL21], Malaysia Sphaerostephanos heterocarpos K000117122
    Jan 1, 1978 Jermy, A.C. [13041], Malaysia Sphaerostephanos heterocarpos K000117483
    Jun 30, 1976 Piggott, A.G. [1477], Malaysia Sphaerostephanos heterocarpos K000116950
    Jun 30, 1976 Piggott, A.G. [1457], Malaysia Sphaerostephanos heterocarpos K000116957
    Jan 1, 1976 Anonymous [s.n.], Indonesia Sphaerostephanos heterocarpos K000117472
    Jan 1, 1976 Korthals, P.W. [s.n.], Indonesia Sphaerostephanos heterocarpos K000117452
    Mar 1, 1975 Anonymous [156], Indonesia Sphaerostephanos heterocarpos K000117460
    Mar 1, 1975 Anonymous [94], Indonesia Sphaerostephanos heterocarpos K000117461
    Oct 1, 1974 Price, M.G. [724], Philippines Sphaerostephanos heterocarpos K000399064
    Oct 1, 1974 Price, M.G. [676], Philippines Sphaerostephanos heterocarpos K000399065
    Aug 1, 1974 Jermy, A.C. [7054], Indonesia Sphaerostephanos heterocarpos K000117466
    Jan 1, 1974 Woodhams, J.R. [212B], Papua New Guinea Sphaerostephanos heterocarpos K000422443
    Jul 11, 1973 Jermy, A.C. [7061], Indonesia Sphaerostephanos heterocarpos K000117464
    Jul 11, 1973 Jermy, A.C. [7061], Indonesia Sphaerostephanos heterocarpos K000117465
    May 5, 1971 Jermy, A.C. [7054], Indonesia Sphaerostephanos heterocarpos K000117462
    Feb 23, 1968 Elmer, A.D.E. [6823], Philippines Sphaerostephanos heterocarpos K000399063
    Carr, C.E. [12497], Papua New Guinea Sphaerostephanos heterocarpos K000422447
    Brass, L.J. [32034], Papua New Guinea Sphaerostephanos heterocarpos K000422239
    Clemens, J. [21018], Malaysia Sphaerostephanos heterocarpos K000117480
    Corner, E.J.H. [s.n.], Malaysia Sphaerostephanos heterocarpos K000116791
    Corner, E.J.H. [30106], Malaysia Sphaerostephanos heterocarpos K000451763
    Corner, E.J.H. [30106], Malaysia Sphaerostephanos heterocarpos K000451764
    Ridley, H.N. [6963], Malaysia Sphaerostephanos heterocarpos K000116786
    Ridley, H.N. [s.n.], Malaysia Sphaerostephanos heterocarpos K000116958
    Jermy, A.C. [7054], Indonesia Sphaerostephanos heterocarpos K000117463
    Jermy, A.C. [13063], Malaysia Sphaerostephanos heterocarpos K000117481
    Jermy, A.C. [13041], Malaysia Sphaerostephanos heterocarpos K000117482
    Jermy, A.C. [13041], Malaysia Sphaerostephanos heterocarpos K000117484
    Jermy, A.C. [14244], Malaysia Sphaerostephanos heterocarpos K000117492
    Jermy, A.C. [14244], Malaysia Sphaerostephanos heterocarpos K000117493
    Holttum, R.E. [s.n.], Malaysia Sphaerostephanos heterocarpos K000116796
    Holttum, R.E. [31382], Malaysia Sphaerostephanos heterocarpos K000116797
    Holttum, R.E. [31378], Malaysia Sphaerostephanos heterocarpos K000116941
    Holttum, R.E. [24564], Malaysia Sphaerostephanos heterocarpos K000116942
    Holttum, R.E. [s.n.], Malaysia Sphaerostephanos heterocarpos K000116943
    Holttum, R.E. [s.n.], Malaysia Sphaerostephanos heterocarpos K000116944
    Holttum, R.E. [31308], Malaysia Sphaerostephanos heterocarpos K000116945
    Holttum, R.E. [31308], Malaysia Sphaerostephanos heterocarpos K000116946
    Holttum, R.E. [9474], Malaysia Sphaerostephanos heterocarpos K000117139
    Holttum, R.E. [34], Malaysia Sphaerostephanos heterocarpos K000117498
    Holttum, R.E. [34], Malaysia Sphaerostephanos heterocarpos K000117499
    Holttum, R.E. [34], Malaysia Sphaerostephanos heterocarpos K000117500
    Meijer, W. [10540], Indonesia Sphaerostephanos heterocarpos K000117473
    Hoogland, R.D. [8968], Papua New Guinea Sphaerostephanos heterocarpos K000422230
    Anderson, J.A.R. [S29884], Malaysia Sphaerostephanos heterocarpos K000117485
    Anderson, J.A.R. [S29884], Malaysia Sphaerostephanos heterocarpos K000117486
    Price, M.G. [3298], Philippines Sphaerostephanos heterocarpos K000399066
    Edwards, P.J. [1946], Malaysia Sphaerostephanos heterocarpos K000117135
    Edwards, P.J. [1946], Malaysia Sphaerostephanos heterocarpos K000117136
    Piggott, A.G. [1928], Malaysia Sphaerostephanos heterocarpos K000116792
    Piggott, A.G. [1928], Malaysia Sphaerostephanos heterocarpos K000116793
    Piggott, A.G. [1928], Malaysia Sphaerostephanos heterocarpos K000116794
    Piggott, A.G. [1930], Malaysia Sphaerostephanos heterocarpos K000116795
    Piggott, A.G. [1501], Malaysia Sphaerostephanos heterocarpos K000116948
    Piggott, A.G. [1501], Malaysia Sphaerostephanos heterocarpos K000116949
    Piggott, A.G. [3133], Malaysia Sphaerostephanos heterocarpos K000117119
    Piggott, A.G. [3133], Malaysia Sphaerostephanos heterocarpos K000117121
    Piggott, A.G. [1956], Malaysia Sphaerostephanos heterocarpos K000117133
    Piggott, A.G. [3133], Malaysia Sphaerostephanos heterocarpos K000117118
    Shim, P.S. [134746], Malaysia Sphaerostephanos heterocarpos K000117496
    Shim, P.S. [134746], Malaysia Sphaerostephanos heterocarpos K000117497
    Parris, B.S. [10411], Malaysia Sphaerostephanos heterocarpos K000116940
    Parris, B.S. [6741], Malaysia Sphaerostephanos heterocarpos K000117474
    Parris, B.S. [6741], Malaysia Sphaerostephanos heterocarpos K000117475
    Parris, B.S. [6741], Malaysia Sphaerostephanos heterocarpos K000117476
    Parris, B.S. [6741], Malaysia Sphaerostephanos heterocarpos K000117477
    Parris, B.S. [7004], Malaysia Sphaerostephanos heterocarpos K000117478
    Parris, B.S. [5969], Papua New Guinea Sphaerostephanos heterocarpos K000422228
    Parris, B.S. [9515], Papua New Guinea Sphaerostephanos heterocarpos K000422232
    Parris, B.S. [9515], Papua New Guinea Sphaerostephanos heterocarpos K000422233
    Parris, B.S. [9515], Papua New Guinea Sphaerostephanos heterocarpos K000422234
    Parris, B.S. [5976], Papua New Guinea Sphaerostephanos heterocarpos K000422226
    Parris, B.S. [5969], Papua New Guinea Sphaerostephanos heterocarpos K000422227
    Parris, B.S. [4377], Papua New Guinea Sphaerostephanos heterocarpos K000422229
    Parris, B.S. [9515], Papua New Guinea Sphaerostephanos heterocarpos K000422231
    Parris, B.S. [11644], Malaysia Sphaerostephanos heterocarpos K000397501
    Parris, B.S. [8934], Malaysia Sphaerostephanos heterocarpos K000397504
    Kiew, R. [2057], Malaysia Sphaerostephanos heterocarpos K000117137
    Dr. King's Collector [1080], Malaysia Sphaerostephanos heterocarpos K000116787
    Dr. King's Collector [491], Malaysia Sphaerostephanos heterocarpos K000116790
    Burkill, I.H. [8814], Malaysia Sphaerostephanos heterocarpos K000116947
    Ramos, M. [34476], Philippines Sphaerostephanos heterocarpos K000399068
    Elmer, A.D.E. [18386], Philippines Sphaerostephanos heterocarpos K000399062
    Kloss, C.B. [18698], Malaysia Sphaerostephanos heterocarpos K000397536
    Beccari, O. [s.n.], Indonesia Sphaerostephanos heterocarpos K000117451
    Matthew, C.G. [s.n], Malaysia Sphaerostephanos heterocarpos K000117490
    Matthew, C.G. [s.n], Malaysia Sphaerostephanos heterocarpos K000117491
    Merrill, E.D. [720], Philippines Sphaerostephanos heterocarpos K000399072
    Merrill, E.D. [720], Philippines Sphaerostephanos heterocarpos K000399073
    Croft, J.R. [323], Papua New Guinea Sphaerostephanos heterocarpos K000422449
    Croft, J.R. [LAE65360], Papua New Guinea Sphaerostephanos heterocarpos K000547820
    Croft, J.R. [LAE65360], Papua New Guinea Sphaerostephanos heterocarpos K000547821
    Croft, J.R. [323], Papua New Guinea Sphaerostephanos heterocarpos K000422450
    Croft, J.R. [LAE68574], Papua New Guinea Sphaerostephanos heterocarpos K000422237
    Croft, J.R. [419], Papua New Guinea Sphaerostephanos heterocarpos K000422238
    Anonymous [147], Malaysia Sphaerostephanos heterocarpos K000117479
    Robinson, H.C. [s.n.], Malaysia Sphaerostephanos heterocarpos K000116789
    Hervey, D.F.A. [3212], Malaysia Sphaerostephanos heterocarpos K000117134
    Copeland, E.B. [1643], Philippines Sphaerostephanos heterocarpos K000399074
    Nur, M. [32934], Malaysia Sphaerostephanos heterocarpos K000116798
    Nur, M. [32934], Malaysia Sphaerostephanos heterocarpos K000116799
    Reillo, J. [16210], Philippines Sphaerostephanos heterocarpos K000399067
    Iwatsuki, k. [13674], Malaysia Sphaerostephanos heterocarpos K000116801
    Iwatsuki, k. [S78], Indonesia Sphaerostephanos heterocarpos K000117454
    Iwatsuki, k. [S78], Indonesia Sphaerostephanos heterocarpos K000117455
    Iwatsuki, k. [B2950], Indonesia Sphaerostephanos heterocarpos K000397529
    Iwatsuki, k. [B1023], Indonesia Sphaerostephanos heterocarpos K000397530
    Iwatsuki, k. [B1023], Indonesia Sphaerostephanos heterocarpos K000397531
    Iwatsuki, k. [B2166], Indonesia Sphaerostephanos heterocarpos K000397532
    Iwatsuki, k. [B2166], Indonesia Sphaerostephanos heterocarpos K000397533
    Iwatsuki, k. [M14161], Malaysia Sphaerostephanos heterocarpos K000117138
    Kato, M. [C12916], Indonesia Sphaerostephanos heterocarpos K000397512
    Kato, M. [C12916], Indonesia Sphaerostephanos heterocarpos K000397513
    Kato, M. [C5025], Indonesia Sphaerostephanos heterocarpos K000397514
    Ratcliffe, J.B. [132], Indonesia Sphaerostephanos heterocarpos K000422224
    Clunie, N.M.U. [LAE63341], Papua New Guinea Sphaerostephanos heterocarpos K000422240
    Clunie, N.M.U. [LAE63341], Papua New Guinea Sphaerostephanos heterocarpos K000422241
    Hose, E.S. [161], Malaysia Sphaerostephanos heterocarpos K000117123
    Hose, E.S. [161], Malaysia Sphaerostephanos heterocarpos K000117124
    Hose, E.S. [4821], Malaysia Sphaerostephanos heterocarpos K000117125
    Hose, E.S. [4821], Malaysia Sphaerostephanos heterocarpos K000117126
    Hose, E.S. [4821], Malaysia Sphaerostephanos heterocarpos K000117127
    Hose, E.S. [4821], Malaysia Sphaerostephanos heterocarpos K000117128
    Hose, E.S. [178], Malaysia Sphaerostephanos heterocarpos K000117129
    Hose, E.S. [178], Malaysia Sphaerostephanos heterocarpos K000117130
    Hose, E.S. [178], Malaysia Sphaerostephanos heterocarpos K000117131
    Hose, E.S. [178], Malaysia Sphaerostephanos heterocarpos K000117132
    Hose, G.F. [294], Malaysia Sphaerostephanos heterocarpos K000397517
    Ernst, A. [936], Indonesia Sphaerostephanos heterocarpos K000117467
    Lee, D.W. [UL47], Malaysia Sphaerostephanos heterocarpos K000116953
    Lee, D.W. [UL47], Malaysia Sphaerostephanos heterocarpos K000116954
    Lee, D.W. [UL11], Malaysia Sphaerostephanos heterocarpos K000116955
    Lee, D.W. [UL11], Malaysia Sphaerostephanos heterocarpos K000116956
    Joncheere, G.J. de [1595], Indonesia Sphaerostephanos heterocarpos K000397506
    Joncheere, G.J. de [1743], Indonesia Sphaerostephanos heterocarpos K000397507
    Joncheere, G.J. de [1743], Indonesia Sphaerostephanos heterocarpos K000397508
    Joncheere, G.J. de [1567], Indonesia Sphaerostephanos heterocarpos K000397509
    Joncheere, G.J. de [1567], Indonesia Sphaerostephanos heterocarpos K000397510
    Joncheere, G.J. de [1552], Indonesia Sphaerostephanos heterocarpos K000397511
    Bünnemeijer, H.A.B. [5935], Indonesia Sphaerostephanos heterocarpos K000117449
    Hallier, J.G. [172], Indonesia Sphaerostephanos heterocarpos K000117459
    Manseima, J. [17], Papua New Guinea Sphaerostephanos heterocarpos K000422236
    Manseima, J. [17], Papua New Guinea Sphaerostephanos heterocarpos K000422235
    Bamler, G. [124], Papua New Guinea Sphaerostephanos heterocarpos K000547826
    Bamler, G. [124], Papua New Guinea Sphaerostephanos heterocarpos K000547825
    Bamler, G. [124], Papua New Guinea Sphaerostephanos heterocarpos K000422225
    Mousset, J.P. [103], Indonesia Sphaerostephanos heterocarpos K000547968
    Mousset, J.P. [103], Indonesia Sphaerostephanos heterocarpos K000117456
    Mousset, J.P. [301], Indonesia Sphaerostephanos heterocarpos K000117457
    Molesworth Allen, B.E.G. [4613], Malaysia Sphaerostephanos heterocarpos K000116788
    Norris, W. [s.n.], Malaysia Sphaerostephanos heterocarpos K000116785
    Burchard, W.F.G. [s.n.], Indonesia Sphaerostephanos heterocarpos K000117450
    Allen, B. [1977], Malaysia Sphaerostephanos heterocarpos K000116800
    Allen, B. [1697], Malaysia Sphaerostephanos heterocarpos K000116951
    Allen, B. [1697], Malaysia Sphaerostephanos heterocarpos K000116952
    Anderson, J.W. [31], Malaysia Sphaerostephanos heterocarpos K000117487
    Addison, G.H. [37151], Malaysia Sphaerostephanos heterocarpos K000116959
    Gebo, A. [4376], Papua New Guinea Sphaerostephanos heterocarpos K000422444
    Gebo, A. [4376], Papua New Guinea Sphaerostephanos heterocarpos K000422445
    Main, T.W. [1], Indonesia Sphaerostephanos heterocarpos K000117469
    Main, T.W. [1], Indonesia Sphaerostephanos heterocarpos K000117470
    Main, T.W. [1], Indonesia Sphaerostephanos heterocarpos K000117471
    Kadir, A. [A548], Malaysia Sphaerostephanos heterocarpos K000397539
    Iwatsuki. K. [S1633], Indonesia Sphaerostephanos heterocarpos K000117448
    Sapiën [2683], Indonesia Sphaerostephanos heterocarpos K000117468
    Gebo, A. [Parris B.S. [4376], Papua New Guinea Sphaerostephanos heterocarpos K000422446
    Hose., G.F. [s.n.], Malaysia Sphaerostephanos heterocarpos K000117494

    First published in Amer. Fern J. 49: 113 (1959)

    Accepted by

    • Fraser-Jenkins, C.R. & al. (2017). An Annotated Checklist of Indian Pteridophytes 1: 1-562. Dehra Dun : BSMPS.
    • Newman, M., Ketphanh, S., Svengsuksa, B., Thomas, P., Sengdala, K., Lamxay, V. & Armstrong, K. (2007). A checklist of the vascular plants of Lao PDR: 1-394. Royal Botanic Gardens, Edinburgh.

    Not accepted by

    • Roskov Y. & al. (eds.) (2018). World Ferns: Checklist of Ferns and Lycophytes of the World Species 2000 & ITIS Catalogue of Life Naturalis, Leiden, the Netherlands. [Cited as Sphaerostephanos heterocarpus.]

    Literature

    Kew Backbone Distributions
    • Newman, M., Ketphanh, S., Svengsuksa, B., Thomas, P., Sengdala, K., Lamxay, V. & Armstrong, K. (2007). A checklist of the vascular plants of Lao PDR: 1-394. Royal Botanic Gardens, Edinburgh.

    Sources

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0