1. Family: Aspleniaceae Newman
    1. Genus: Thelypteris Schmidel
      1. Thelypteris hispidula (Decne.) C.F.Reed

        This species is accepted, and its native range is Tropics & Subtropics.

    [FWTA]

    Thelypteridaceae, A. H. G. Alston. Flora of West Tropical Africa. Ferns and Fern-Allies Supplement. 1959

    Ecology
    In shade usually, in secondary growth or cultivated areas; at low elevations.
    [FTEA]

    Thelypteridaceae, Bernard Verdcourt. Flora of Tropical East Africa. 2006

    Type
    Type: Timor, Guichenot s.n. (P, holo.)
    Rhizomes
    Rhizome erect with brown to dark brown lanceolate entire or ciliate often thinly pilose scales up to 6 mm long
    Leaves
    Fronds tufted, 0.3–1 m tall
    Stipes
    Stipe 12–40(–60) cm long, thinly pubescent with minute whitish hairs and scaly at base
    Leaf lamina
    Lamina lanceolate, deeply 2-pinnatifid, 25–60 cm long, ± 30 cm wide, apex acuminate with deeply pinnatifid terminal segment; lower 2–4 pair of pinnae deflexed and somewhat reduced; middle pinnae 8–16 cm long, 1.2–2.2 cm wide, narrowly oblong, acuminate, deeply pinnatifid; lobes narrowly oblong, 4–10 mm long, 2–5 mm wide, somewhat falcate, entire, obtuse, hairy on both surfaces and sometimes with small capitate hairs; veins 7–8 pairs, the basal pair anastomosing at a very obtuse angle below the sinus with next pair to edge of sinus
    Indusium
    Indusium hairy, the hairs sometimes very long.
    Ecology
    Evergreen rain forest, often riverine, and on rocks near falls; 150–2400 m
    Note
    R.B. & A.J. Faden 74/369 (Lushoto District: E Usambaras, Derema, Hunga R., near small falls, 30 Mar. 1974) and other specimens show considerable variation in the anastomosing of the basal veins, some joining and having an excurrent vein to the sinus others quite free but close at base of sinus (all on one pinna). This is, I feel, just normal variation rather than hybridisation which is very often a facile explanation based on no evidence. Some specimens I have merely annotated as C. hispidula/gueinziana intermediates.
    Distribution
    Flora districts: U2 U3 U 4 ; K4 K5 K7 T2 T3 T 4 , T6 T7 T ?8 Range: West Africa, Bioko, Cameroon, Sudan, Malawi, Mozambique, Zimbabwe; Seychelles, Mascarene Is., Madagascar; widespread in tropical Asia and America

    Images

    Distribution

    Native to:

    Angola, Argentina Northeast, Argentina Northwest, Belize, Bolivia, Borneo, Brazil North, Brazil Northeast, Brazil South, Brazil Southeast, Brazil West-Central, Cameroon, Central African Repu, Colombia, Comoros, Costa Rica, Ecuador, El Salvador, Ethiopia, French Guiana, Galápagos, Ghana, Guatemala, Gulf of Guinea Is., Guyana, Honduras, India, Ivory Coast, Jamaica, Jawa, Kenya, KwaZulu-Natal, Leeward Is., Lesser Sunda Is., Liberia, Madagascar, Malawi, Malaya, Maluku, Mauritius, Mexico Central, Mexico Gulf, Mexico Northeast, Mexico Northwest, Mexico Southeast, Mexico Southwest, Mozambique, Myanmar, New Guinea, New South Wales, Nicaragua, Nigeria, Northern Provinces, Panamá, Paraguay, Peru, Philippines, Puerto Rico, Queensland, Réunion, Seychelles, Sierra Leone, South Australia, Sri Lanka, Sudan, Sulawesi, Sumatera, Suriname, Tanzania, Thailand, Uganda, Uruguay, Venezuela, Vietnam, Windward Is., Zimbabwe

    Thelypteris hispidula (Decne.) C.F.Reed appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Piggott, A.G. [3096], Malaysia Christella siamensis K000506591
    Piggott, A.G. [3096], Malaysia Christella siamensis K000506592
    Jul 1, 2009 Rosu [17], Malaysia Christella hispidula K000610854
    Oct 1, 2005 Darbyshire, I. [4814], Cameroon Christella hispidula K000338355
    Aug 1, 1996 Etuge, M. [1483], Cameroon Christella hispidula K000381606
    Aug 1, 1996 Etuge, M. [1483], Cameroon Christella hispidula K000381607
    Jan 1, 1985 Heringer, E.P. [6004], Brazil Christella hispidula K000956535
    Feb 23, 1981 Parris, B.S. [8953], Malaysia Christella hispidula K000519247
    Feb 23, 1981 Parris, B.S. [8921], Malaysia Christella hispidula K000519249
    Jan 1, 1981 Carr, C.E. [11263], Papua New Guinea Christella hispidula K000547845
    Jan 1, 1981 Clemens, J. [29238], Malaysia Christella hispidula K000519256
    Jan 1, 1981 Clemens, J. [27475], Malaysia Christella hispidula K000519257
    Jan 1, 1981 Clemens, J. [27475], Malaysia Christella hispidula K000519258
    Jan 1, 1981 Clemens, J. [27739bis], Malaysia Christella hispidula K000519259
    Jan 1, 1981 Clemens, J. [27469], Malaysia Christella hispidula K000519254
    Jan 1, 1981 Ridley, H.N. [14196], Malaysia Christella hispidula K000506363
    Jan 1, 1981 Ridley, H.N. [14196], Malaysia Christella hispidula K000506364
    Jan 1, 1981 Holttum, R.E. [SFN25727], Malaysia Christella hispidula K000519253
    Jan 1, 1981 Price, M.G. [1979], Philippines Christella hispidula K000519976
    Jan 1, 1981 Price, M.G. [585], Philippines Christella hispidula K000519977
    Jan 1, 1981 Price, M.G. [585], Philippines Christella hispidula K000519978
    Jan 1, 1981 Price, M.G. [1979], Philippines Christella hispidula K000519975
    Jan 1, 1981 Williams, R.S. [2432], Philippines Christella hispidula K000519979
    Jan 1, 1981 Dr. King's Collector [8657], Malaysia Christella hispidula K000506361
    Jan 1, 1981 Dr. King's Collector [8657], Malaysia Christella hispidula K000506362
    Jan 1, 1981 Ramos, M. [32959], Philippines Christella hispidula K000519980
    Jan 1, 1981 Elmer, A.D.E. [20383], Malaysia Christella hispidula K000519251
    Jan 1, 1981 Lam, H.J. [1058], Indonesia Christella hispidula K000547841 Unknown type material
    Jan 1, 1981 Merrill, E.D. [824], Philippines Christella hispidula K000519981
    Jan 1, 1981 Croft, J.R. [NGF41343], Papua New Guinea Christella hispidula K000547850
    Jan 1, 1981 Cuadra, A. [A2345], Malaysia Christella hispidula K000519250
    Jan 1, 1981 Kornassi [11], Indonesia Christella hispidula K000519265
    Jan 1, 1981 Yapp., R.H. [201], Malaysia Christella hispidula K000506359
    Jan 1, 1981 Kadir, A. [A2596], Malaysia Christella hispidula K000519252
    Oct 18, 1980 Eiten, G. [4735], Brazil Christella hispidula K000956543
    Nov 1, 1977 Luerssen [4030], Brazil Christella hispidula K000956555
    Feb 2, 1977 Giles, C. [817], Malaysia Christella hispidula K000519255
    Jan 18, 1977 Cuming, H. [338B], Philippines Christella hispidula K000548255
    Aug 25, 1976 Albertis, L.M. d' [s.n.], Papua New Guinea Christella hispidula K000547846
    Aug 24, 1976 Ridley, H.N. [s.n.], Indonesia Christella hispidula K000507897
    Aug 24, 1976 Matthew, C.G. [s.n.], Indonesia Christella hispidula K000507898
    Aug 24, 1976 Merrill, E.D. [7671], Philippines Christella hispidula K000519982 Unknown type material
    Aug 24, 1976 Motley, J. [39] Christella hispidula K000519240
    Aug 24, 1976 Motley, J. [39] Christella hispidula K000519241
    Aug 24, 1976 Barber [39] Christella hispidula K000519242
    Aug 20, 1976 Matthew, C.G. [s.n.], Malaysia Christella hispidula K000506360
    Aug 20, 1976 Anonymous [s.n], Malaysia Christella hispidula K000506358
    Jun 1, 1975 Blume, C.L. [s.n.], Indonesia Christella hispidula K000506172
    Comber, H.F. [4060], Malaysia Christella hispidula K000451750
    Comber, H.F. [4060], Malaysia Christella hispidula K000451751
    Comber, H.F. [4060], Malaysia Christella hispidula K000451752
    Giles, C. [s.n.], Malaysia Christella hispidula K000519243
    Giles, C. [s.n.], Malaysia Christella hispidula K000519244
    Krapovickas, A. [40833], Brazil Christella hispidula K000956525
    Parris, B.S. [4427], Papua New Guinea Christella hispidula K000547847
    Parris, B.S. [4423], Papua New Guinea Christella hispidula K000547842
    Parris, B.S. [4832], Papua New Guinea Christella hispidula K000547849
    Parris, B.S. [10677], Malaysia Christella hispidula K000519245
    Parris, B.S. [10697], Malaysia Christella hispidula K000519246
    Parris, B.S. [10832], Malaysia Christella hispidula K000519248
    Parris, B.S. [4423], Papua New Guinea Christella hispidula K000547843
    Parris, B.S. [4427], Papua New Guinea Christella hispidula K000547848
    Pereira, E. [7890], Brazil Christella hispidula K000956539
    Ratter, J.A. [4890], Brazil Christella hispidula K000956533
    Spruce, R. [1107], Brazil Christella hispidula K000956551
    Blanchet [s.n.], Brazil Christella hispidula K000956548
    Blanchet [2635], Brazil Christella hispidula K000956563
    Salzmann [s.n.], Brazil Christella hispidula K000956557
    Salzmann [s.n.], Brazil Christella hispidula K000956558
    Moricand [2457], Brazil Christella hispidula K000956549
    Anderson, W.R. [9450], Brazil Christella hispidula K000956530
    Bolland, G. [s.n.], Brazil Christella hispidula K000956537
    Lyra-Lemos, R.P. [995], Brazil Christella hispidula K000956528
    Lyra-Lemos, R.P. [995], Brazil Christella hispidula K000956529
    Cavalcante, P.B. [2597], Brazil Christella hispidula K000956526
    Hohenacker, R.F. [602], India Christella hispidula K000435796
    Sieber [s.n.], Mauritius Christella hispidula K000435795 Unknown type material
    Sieber [s.n.] Christella hispidula K000435797
    Gilpin, H. [s.n.], Madagascar Christella hispidula K000435798
    Fonseca, C.G. [53], Brazil Christella hispidula K000956534
    Robert, A. [335b], Brazil Christella hispidula K000956540
    Graham [s.n.], Brazil Christella hispidula K000956545
    Lindman, C.A.M. [A.963], Brazil Christella hispidula K000956538
    Lindberg [638], Brazil Christella hispidula K000956562
    Regel [s.n.], Brazil Christella hispidula K000956560
    Sellow [s.n.], Brazil Christella hispidula K000956553
    Kato, M. [C5953], Indonesia Christella hispidula K000519260
    Kato, M. [C5953], Indonesia Christella hispidula K000519261
    Kato, M. [C4769], Indonesia Christella hispidula K000519262
    Kato, M. [C4778], Indonesia Christella hispidula K000519263
    Kato, M. [C4778], Indonesia Christella hispidula K000519264
    Prance, G.T. [1855], Brazil Christella hispidula K000956527
    Prance, G.T. [14446], Brazil Christella hispidula K000956531
    Prance, G.T. [14446], Brazil Christella hispidula K000956532
    Mitchell, F. [3833], Papua New Guinea Christella hispidula K000547844
    Lindberg, G. [636], Brazil Christella hispidula K000956542
    Lindberg, G. [637], Brazil Christella hispidula K000956556
    Lindberg, G. [638], Brazil Christella hispidula K000956561
    Mexia, Y.E.J. [4863], Brazil Christella hispidula K000956541
    Gardner [1902], Brazil Christella hispidula K000956536
    Gardner [s.n.], Brazil Christella hispidula K000956546
    Gardner [s.n.], Brazil Christella hispidula K000956552
    Gardner [1902], Brazil Christella hispidula K000956554
    s.coll. [s.n.], French Guiana Christella hispidula K000956559
    Burchell [2104], Brazil Christella hispidula K000956547
    Burchell [4473], Brazil Christella hispidula K000956550
    Glaziou [14401/A], Brazil Christella hispidula K000956544

    First published in Phytologia 17: 283 (1968)

    Accepted by

    • Fraser-Jenkins, C.R. & al. (2017). An Annotated Checklist of Indian Pteridophytes 1: 1-562. Dehra Dun : BSMPS.

    Not accepted by

    • Roskov Y. & al. (eds.) (2018). World Ferns: Checklist of Ferns and Lycophytes of the World Species 2000 & ITIS Catalogue of Life Naturalis, Leiden, the Netherlands. [Cited as Christella quadrangularis.]

    Literature

    Kew Backbone Distributions
    • Fraser-Jenkins, C.R. & al. (2017). An Annotated Checklist of Indian Pteridophytes 1: 1-562. Dehra Dun : BSMPS.

    Sources

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of West Tropical Africa
    Flora of West Tropical Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0