1. Family: Poaceae Barnhart
    1. Genus: Pariana Aubl.
      1. Pariana bicolor Tutin

        This species is accepted, and its native range is Peru to Bolivia and N. Brazil.

    [GB]
    Habit
    Perennial. Culms erect; 40-80 cm long. Leaves cauline. Leaf-sheath oral hairs scanty, or lacking. Ligule an eciliate membrane; 2-3 mm long. Leaf-blade base with a brief petiole-like connection to sheath; petiole 0.2-0.3 cm long; petiole pubescent. Leaf-blades oblong, or ovate; 12-22 cm long; 25-50 mm wide; mid-green and glaucous; discolorous with last colour beneath. Leaf-blade surface scabrous; rough abaxially; glabrous. Leaf-blade apex acuminate.
    Inflorescences
    Monoecious; with male and female spikelets in the same inflorescence. Synflorescence on a separate leafless culm. Inflorescence composed of racemes. Racemes 1; single; erect; multilateral; 7-9 cm long; 8-10 mm wide. Rhachis fragile at the nodes; terminating in a spikelet. Spikelet packing lax. Rhachis internodes linear. Sexes mixed (in a cluster at each node). Spikelets subtended by an involucre. Fertile spikelets sessile; 1 in the cluster. Male spikelets pedicelled; 5 in a cluster. Involucre composed of imperfect spikelets; with pedicels forming a cup; oblong. Pedicels fused to each other; united along margins; oblong; flattened; 2-3 mm long; puberulous and villous (at base); tip rectangular.
    Fertile
    Spikelets comprising 1 fertile florets; without rhachilla extension. Spikelets ovate; laterally compressed; compressed slightly; 6 mm long; falling entire; deciduous with accessory branch structures.
    Glume
    Glumes similar; reaching apex of florets; thinner than fertile lemma. Lower glume ovate; 6 mm long; 1 length of upper glume; chartaceous; 1-keeled. Lower glume margins ciliate. Lower glume apex obtuse. Upper glume ovate; 6 mm long; 1 length of adjacent fertile lemma; chartaceous; 1-keeled. Upper glume margins ciliate. Upper glume apex obtuse.
    Florets
    Fertile florets female. Fertile lemma ovate; 6 mm long; coriaceous; without keel; 3 -veined. Lemma apex acute. Palea coriaceous; without keels.
    Flowers
    Lodicules 3. Anthers 25. Filaments united in a tube. Stigmas 2.
    Spikelets
    Male spikelets distinct from female; 1 flowered; deciduous with the fertile; ovate; 3.5-4.5 mm long; glabrous, or hairy. Male spikelet glumes 2; triangular; 2-2.5 mm long. Male spikelet lemma 3.5-4.5 mm long; 3 -veined; with cross-veins. Spikelets comprising 1 fertile florets; without rhachilla extension. Spikelets ovate; laterally compressed; compressed slightly; 6 mm long; falling entire; deciduous with accessory branch structures.
    Male
    Male spikelets distinct from female; 1 flowered; deciduous with the fertile; ovate; 3.5-4.5 mm long; glabrous, or hairy. Male spikelet glumes 2; triangular; 2-2.5 mm long. Male spikelet lemma 3.5-4.5 mm long; 3 -veined; with cross-veins.
    Distribution
    South America: western South America and Brazil.
    Reference
    Parianeae. Tutin 1995.

    Images

    Distribution

    Native to:

    Bolivia, Brazil North, Peru

    Pariana bicolor Tutin appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Apr 11, 1996 Maas, P.J.M. [P13016], Brazil K000947297
    Apr 11, 1996 Prance, G.T. [14238], Brazil K000947296
    Apr 4, 1996 Krukoff, B.A. [6957], Brazil K000947298
    Krukoff, B.A. [6957], Brazil K000947299

    First published in J. Linn. Soc., Bot. 50: 355 (1936)

    Accepted by

    • Clayton, W.D., Harman, K.T. & Williamson, H. (2006). World Grass Species - Synonymy database The Board of Trustees of the Royal Botanic Gardens, Kew.
    • Renvoize, S.A. (1998). Gramineas de Bolivia: 1-644. The Royal Botanic Gardens, Kew.
    • Brako, L. & Zarucchi, J.L. (1993). Catalogue of the Flowering Plants and Gymnosperms of Peru Monographs in Systematic Botany from the Missouri Botanical Garden 45: i-xl, 1-1286.

    Literature

    Kew Backbone Distributions
    • Forzza, R.C. & al. (2013). Lista de Espécies da Flora do Brasil http://floradobrasil.jbrj.gov.br/2010/.
    • Renvoize, S.A. (1998). Gramineas de Bolivia: 1-644. The Royal Botanic Gardens, Kew.
    • Brako, L. & Zarucchi, J.L. (1993). Catalogue of the Flowering Plants and Gymnosperms of Peru Monographs in Systematic Botany from the Missouri Botanical Garden 45: i-xl, 1-1286.

    Sources

    GrassBase - The Online World Grass Flora
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2020. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2020. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0