1. Family: Asteraceae Bercht. & J.Presl
    1. Genus: Carduus L.
      1. Carduus nyassanus R.E.Fr.

        This species is accepted, and its native range is Nigeria to Ethiopia and S. Tropical Africa.

    [FZ]

    Compositae, G. V. Pope. Flora Zambesiaca 6:1. 1992

    Habit
    An erect spinose perennial herb, to c. 1.5 m. tall.
    Stem
    Stems simple, or sparingly branched above, stout, coarsely striate, ± pubescent, ± crowded leafy below, leaves fewer and smaller above, longitudinally winged throughout with wings interrupted or sinuate; wing-segments up to c. 1.5 cm. wide and strongly spinose on the margins.
    Leaves
    Basal leaves 10–33(50) x 2–6.5 cm., narrowly oblanceolate in outline, pinnately divided, pubescent especially on the midribs, or glabrescent; leaf-segments up to c. 3.5 cm. long near the leaf apex and decreasing in size towards the leaf base, margins coarsely dentate with the teeth and apices strongly spine-tipped.
    Capitulum
    Capitula many, densely aggregated in terminal clusters 3.5–7 cm. in diam.; involucres up to c. 2.5 cm. long, narrowly campanulate.
    Phyllaries
    Phyllaries numerous, many-seriate, narrowly lanceolate, tapering to apical spines, the spines patent and increasingly longer on the inner phyllaries, the lamina stamineous or purple-tinged, sparsely puberulent outside and ciliolate on the margins, the outer phyllaries from c. 10 mm. long, the middle to c. 20 mm. long, the innermost shorter.
    Corolla
    Corollas white, reddish-tinged or mauve, 10–13 mm. long, very narrowly infundibuliform, limb 7–8 mm. long, glabrous, lobes 2.5–4 mm. long.
    Fruits
    Achenes stramineous or pale-brown, 4–4.5 mm. long, narrowly turbinate and somewhat 3–4-ribbed, glabrous; pappus several-seriate, setae 11–12 mm. long, numerous, flattened, barbellate, united at the base in a deciduous ring.
    [FTEA]

    Compositae, H. Beentje, C. Jeffrey & D.J.N. Hind. Flora of Tropical East Africa. 2005

    Habit
    Biennial or perennial herb, 0.5–3 m high, erect, spiny, branching slightly near the inflorescence; stems ribbed, glandular-setose (rarely araneose) with spiny lobed and interrupted wings.
    Leaves
    Leaves sessile, decurrent, spiny, pinnately lobed, the basal leaves largest, oblanceolate or elliptic in outline, 3–60 cm long, 1–21 cm wide, in basal leaves the base attenuate into a spiny-lobed petioloid base, in median leaves the base semi-amplexicaul, 5–47-lobed, the lobes alternate or opposite, with strong spines terminating the lobes and their lobules, with shorter marginal spines in between, sparsely or sometimes densely glandular-setose on both surfaces, rarely thinly lanate above, often araneose to arachnoid-tomentose beneath.
    Capitulum
    Capitula in terminal dense globose clusters of 3–10 together; involucre urn-shaped, 13–27 mm long; phyllaries many, in 3–4 series, lanceolate, 8–24 mm long, pungent with recurved apical spines in the median ones, the outermost setose or arachnoid proximally but otherwise glabrescent, ciliate; receptacle with setae 2.5–8.5 mm long.
    Florets
    Florets 38–102 per capitulum, sweet-scented; corolla white or pale mauve, tube 9.5–15.5 mm long, occasionally with a few glands, lobes 3–5.5 mm long, anthers grey.
    Fruits
    Achenes ovoid, 2.5–5 mm long, slightly angular and compressed, slightly curved, glabrous; pappus of many bristles 10–19 mm long, the longest slightly expanded distally, falling as a single unit.
    [FWTA]

    Compositae, C. D. Adams. Flora of West Tropical Africa 2. 1963

    Habit
    A perennial herb of damp montane grassland, up to 4 ft. high
    Florets
    Florets white.

    Images

    Distribution

    Native to:

    Burundi, Cameroon, Ethiopia, Kenya, Malawi, Nigeria, Rwanda, Sudan, Tanzania, Uganda, Zambia, Zaïre, Zimbabwe

    Carduus nyassanus R.E.Fr. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Apr 1, 2000 Cheek, M. [9815], Cameroon K000746894
    Hepper, F.N. [2719], Cameroon K000027537
    Maitland, T.D. [1649], Cameroon K000027536
    Schimper [44], Ethiopia K000418460 isotype

    First published in Acta Horti Berg. 8: 25 (1925)

    Accepted by

    • Roskov Y. & al. (eds.) (2018). Species 2000 & ITIS Catalogue of Life Naturalis, Leiden, the Netherlands.

    Literature

    Flora of West Tropical Africa
    • in Acta Hort. Berg. 8: 25 (1925).
    Flora of Tropical East Africa
    • Agnew, A.D.Q. & Agnew, S., Upland Kenya Wild Flowers, ed. 2: 227, t. 96 (1994).
    • G.V. Pope in Flora Zambesiaca 6: 51, t. 10 (1992).
    • Lisowski, Aster. Fl. Afr. Centr.: 589 (1991).
    • Maquet in Fl. Rwanda 3: 696 (1985).
    • C. Jeffrey in Kew Bulletin 22: 126 (1968).
    • C.D. Adams in Flora of West Tropical Africa ed. 2, 2: 291 (1963).
    • Kazmi in Mitt. Bot. Staats. München 5: 158 (1963).
    • R.E. Fr. in Acta Horti Berg. 8: 25, t. 3 (1923).

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of West Tropical Africa
    Flora of West Tropical Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0