1. Family: Phyllanthaceae Martinov
    1. Spondianthus Engl.

      1. This genus is accepted, and its native range is Tropical Africa.

    [FTEA]

    Euphorbiaceae, A. R.-Smith. Flora of Tropical East Africa. 1987

    Habit
    Dioecious trees, quite glabrous or with a simple indumentum
    Leaves
    Leaves alternate or subverticillate, often crowded at the shoot-apices, petiolate (petiole very variable in length), stipulate, simple, entire, penninerved, varying greatly in size
    Inflorescences
    Inflorescences terminal, subterminal or axillary in the axils of the uppermost leaves, shortly pedunculate, paniculate, solitary; bracts resembling the stipules, the males several-flowered with the flowers glomerulate, the females 1-flowered
    Male
    Male flowers subsessile; sepals (4–)5, imbricate; petals (4–)5, somewhat smaller than the sepals; disc-glands 5, epipetalous; stamens 5, episepalous, alternating with disc-glands, filaments free, anthers introrse, medifixed, connective apically pigmented and glandular, thecae parallel, longitudinally dehiscent; pistillode obconic-cylindric, truncate and slightly trilobate at the apex
    Flowers
    Female flowers shortly pedicellate; sepals and petals ± as in the ♂ flowers or petals 0; disc shallowly cupular, 5-lobed, the lobes alternating with the sepals; ovary 3-locular, with 2 ovules per locule; styles 3, ± free, short, shortly bilobed, papillose Male flowers subsessile; sepals (4–)5, imbricate; petals (4–)5, somewhat smaller than the sepals; disc-glands 5, epipetalous; stamens 5, episepalous, alternating with disc-glands, filaments free, anthers introrse, medifixed, connective apically pigmented and glandular, thecae parallel, longitudinally dehiscent; pistillode obconic-cylindric, truncate and slightly trilobate at the apex
    Female
    Female flowers shortly pedicellate; sepals and petals ± as in the ♂ flowers or petals 0; disc shallowly cupular, 5-lobed, the lobes alternating with the sepals; ovary 3-locular, with 2 ovules per locule; styles 3, ± free, short, shortly bilobed, papillose
    Fruits
    Fruit ± ellipsoid, loculicidally dehiscent into 3 valves; exocarp crustaceous; endocarp thin, horny; columella persistent, tripartite
    Seeds
    Seeds 1 per locule by abortion, compressed ovoid-ellipsoid, often remaining attached to the columella after dehiscence, shiny; exotesta papyraceous, reddish brown; mesotesta spongiose or chartaceous, white; endotesta thin, crustaceous; albumen scanty forming thin strata; embryo straight; cotyledons broad, flat.

    Distribution

    Native to:

    Angola, Benin, Burkina, Cameroon, Central African Repu, Chad, Congo, Equatorial Guinea, Gabon, Ghana, Guinea, Gulf of Guinea Is., Ivory Coast, Liberia, Mozambique, Nigeria, Sudan, Tanzania, Togo, Uganda, Zaïre

    Spondianthus Engl. appears in other Kew resources:

    First published in Bot. Jahrb. Syst. 36: 215 (1905)

    Accepted by

    • Govaerts, R., Frodin, D.G. & Radcliffe-Smith, A. (2000). World Checklist and Bibliography of Euphorbiaceae (and Pandaceae) 1-4: 1-1622. The Board of Trustees of the Royal Botanic Gardens, Kew.

    Literature

    Flora of West Tropical Africa
    • Pax & K. Hoffm. in Engl. Pflanzenr. Euph. 15: 13–15 (1922).
    • F.T.A. 6, 1: 1044
    • Bot. Jahrb. 36: 215 (1905)
    Flora of Tropical East Africa
    • Pax in E.P. IV. 147 (15): 13 (1922)
    • Hutch. in F.T.A. 6(1): 1044 (1913)
    • in N.B.G.B. 5: 240 (1911) (as Euphorbiacea)
    • in E.J. 36: 215 (1905) (as Anacardiacea)

    Sources

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2018. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2018. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0