1. Family: Verbenaceae J.St.-Hil.
    1. Genus: Phyla Lour.
      1. Phyla nodiflora (L.) Greene

        This species is accepted, and its native range is Tropics & Subtropics.

    [FTEA]

    Verbenaceae, B. Verdcourt. Flora of Tropical East Africa. 1992

    Habit
    Prostrate creeping herb, with slender often purplish stems 20–45 cm. long covered with fine medifixed hairs; rootstock woody.
    Leaves
    Leaves often purplish, oblanceolate to obovate, 0.5–5 cm. long, 0.2–3 cm. wide, acute and sharply serrate at the apex, cuneate and entire at the base, glabrous or thinly pubescent with adpressed medifixed hairs.
    Inflorescences
    Spikes purplish, cylindrical, 0.3–2.5 cm. long; bracts broadly obovate to oblate-cuspidate, 2–3 × 3–5 mm.; peduncles 1.3–8 cm. long.
    Calyx
    Calyx dorsiventrally flattened, 1.4 mm. long, split almost to base abaxially and to about the middle adaxially.
    Corolla
    Corolla mauve, pink or white with yellow centre, often white and purple in one inflorescence, and sometimes purple in bud, white when open; tube ± 1.5 mm. long; lobes unequal, 0.6–0.8 mm. in diameter.
    Stamens
    Stamens 4, didynamous.
    Stigma
    Stigma obliquely capitate.
    Fruits
    Fruit rounded obovoid, 1.2 × 1–1.2 mm., divided into 2 pyrenes.
    Figures
    Fig. 4.
    Habitat
    Sandy lake shores, shore mud, lava boulders by lakes, lakeside grassland, also roadsides and Avicennia salt marshes; 0–1170 m.
    Distribution
    K3 K5 K7 T2 T3 T4 T5 T6 T8 U1 U2 Z now widespread in tropics, subtropics and warmer temperate regionsthroughout tropical Africa from Senegal, Central African Republic, Sudan, Ethiopia and Somalia to South Africa and also in North Africa.
    [FZ]

    Flora Zambesiaca. Vol 8, Pt 7. Avicenniaceae, R. Fernandes. Nesogenaceae, M.A. Diniz. Verbenaceae, R. Fernandes. Lamiaceae, R. Fernandes. 2005.

    Type
    Type from N America (Virginia).
    Ecology
    Lake and dam shores, river banks, floodplains, pan margins, coastal dunes and swamp margins, in damp or seasonally inundated soils, sands and heavy black clays and in mud; 0–1300 m.
    Note
    Moldenke, in Fifth Summ. Verbenaceae: 252 (1971), has recorded Phyla nodiflora var. reptans (H.B. & K.) Moldenke for Moçambique.  However, we have not been able to confirm this record.
    Distribution
    Widespread in the tropics, subtropics and warmer temperate regions; throughout tropical Africa from Senegal, Central African Republic, Sudan, Ethiopia and Somalia to South Africa. Caprivi Strip
    Habit
    Prostrate perennial herb with long trailing stems radiating from a woody taproot. Prostrate perennial herb with long trailing stems radiating from a woody taproot
    Stem
    Stems up to 2(3) m long, decumbent, terete and somewhat fleshy, drying ± angular and sulcate, rooting at the nodes; internodes 0.7–9 cm long; nodes somewhat thickened; branches 1–2 at each node, from leaf axils, up to 30 cm long, erect or the longer branches ± decumbent and sometimes rooting at the nodes. Stems up to 2(3) m long, decumbent, terete and somewhat fleshy, drying ± angular and sulcate, rooting at the nodes; internodes 0.7–9 cm long; nodes somewhat thickened; branches 1–2 at each node, from leaf axils, up to 30 cm long, erect or the longer branches ± decumbent and sometimes rooting at the nodes
    Indumentum
    Indumentum of short appressed medifixed sharply pointed white hairs, sparse on old parts, ± dense on younger parts, mainly on leaves and floral bracts. Indumentum of short appressed medifixed sharply pointed white hairs, sparse on old parts, ± dense on younger parts, mainly on leaves and floral bracts
    Leaves
    Leaves somewhat fleshy, opposite, sessile or shortly petiolate, 0.8–7.2 × 0.4–2.5 cm, spathulate, oblanceolate or obovate, obtuse to rounded or subacute at the apex and sharply coarsely serrate at the upper part, cuneate and entire at the lower part; petiole up to 8 mm long. Leaves somewhat fleshy, opposite, sessile or shortly petiolate, 0.8–7.2 × 0.4–2.5 cm, spathulate, oblanceolate or obovate, obtuse to rounded or subacute at the apex and sharply coarsely serrate at the upper part, cuneate and entire at the lower part; petiole up to 8 mm long
    Inflorescences
    Spikes short, dense, subglobose or shortly-ovoid at flowering time, and up to 1–2.5(3) cm × 5–9 mm at maturity, cylindric; peduncles (0.5)1–11.5 cm long; bracts closely imbricate, up to c. 2.5 × 2 mm, broadly obovate to oblate-cuspidate, persistent and ± accrescent. Spikes short, dense, subglobose or shortly-ovoid at flowering time, and up to 1–2.5(3) cm × 5–9 mm at maturity, cylindric; peduncles (0.5)1–11.5 cm long; bracts closely imbricate, up to c. 2.5 × 2 mm, broadly obovate to oblate-cuspidate, persistent and ± accrescent
    Calyx
    Calyx compressed, membranous, deeply split almost to the base abaxially and to about the middle adaxially, persistent but hardly accrescent; lobes 1.5–2 mm long, hyaline or dark purple, shortly toothed on the keels. Calyx compressed, membranous, deeply split almost to the base abaxially and to about the middle adaxially, persistent but hardly accrescent; lobes 1.5–2 mm long, hyaline or dark purple, shortly toothed on the keels
    Corolla
    Corolla mauve-pink or white with a yellow throat, often white and purple in the same inflorescence, sometimes purple in bud, white when open, c. 2.5 mm long; tube 1.5–2 mm long and 1–1.5 mm broad at the mouth; limb 2–2.5 mm in diameter with the upper lip 2-lobed and erect, the lower lip larger, 3-lobed with the median lobe oblong. Corolla mauve-pink or white with a yellow throat, often white and purple in the same inflorescence, sometimes purple in bud, white when open, c. 2.5 mm long; tube 1.5–2 mm long and 1–1.5 mm broad at the mouth; limb 2–2.5 mm in diameter with the upper lip 2-lobed and erect, the lower lip larger, 3-lobed with the median lobe oblong
    Fruits
    Mericarps 1.5–2 mm long, sub-hemispherical, flattened at the commissural face, obtuse or rounded at the apex, glabrous, enclosed within the persistent calyx. Mericarps 1.5–2 mm long, sub-hemispherical, flattened at the commissural face, obtuse or rounded at the apex, glabrous, enclosed within the persistent calyx.
    [FWTA]

    Verbenaceae, H. Huber, F. N. Hepper & R. D. Meikle. Flora of West Tropical Africa 2. 1963

    Habit
    A prostrate creeping herb
    Ecology
    Of moist places
    Leaves
    Leaves often tinged purple
    Bracts
    Bracts often tinged purple
    Flowers
    Flowers mauve or pink.

    Images

    Distribution

    Native to:

    Alabama, Albania, Aldabra, Algeria, Andaman Is., Angola, Argentina Northeast, Argentina Northwest, Argentina South, Arizona, Arkansas, Aruba, Assam, Bahamas, Baleares, Bangladesh, Belize, Benin, Bermuda, Bolivia, Botswana, Brazil North, Brazil Northeast, Brazil South, Brazil Southeast, Brazil West-Central, Burkina, Burundi, California, Cambodia, Cape Provinces, Caprivi Strip, Caroline Is., Cayman Is., Central African Repu, Chad, Chagos Archipelago, Chile Central, Chile North, China South-Central, China Southeast, Colombia, Corse, Costa Rica, Cuba, Cyprus, Djibouti, Dominican Republic, East Himalaya, Ecuador, Egypt, El Salvador, Eritrea, Ethiopia, Florida, Free State, Galápagos, Georgia, Ghana, Greece, Guatemala, Guinea-Bissau, Gulf States, Hainan, Haiti, Honduras, India, Iran, Italy, Jamaica, Japan, Kansas, Kentucky, Kenya, Kriti, KwaZulu-Natal, Laccadive Is., Leeward Is., Lesotho, Lesser Sunda Is., Liberia, Libya, Louisiana, Madagascar, Malawi, Malaya, Maldives, Mali, Marianas, Mauritania, Mexico Central, Mexico Gulf, Mexico Northeast, Mexico Northwest, Mexico Southeast, Mexico Southwest, Mississippi, Missouri, Morocco, Mozambique, Myanmar, Namibia, Nansei-shoto, Nepal, Netherlands Antilles, Nevada, New Guinea, New Mexico, New South Wales, Nicaragua, Nicobar Is., Niger, Nigeria, North Carolina, Northern Provinces, Northern Territory, Oklahoma, Oman, Oregon, Pakistan, Panamá, Paraguay, Pennsylvania, Peru, Philippines, Puerto Rico, Queensland, Samoa, Saudi Arabia, Senegal, Seychelles, Sicilia, Sinai, Socotra, Solomon Is., Somalia, South Carolina, South China Sea, Southwest Caribbean, Spain, Sri Lanka, Sudan, Sulawesi, Swaziland, Tanzania, Texas, Thailand, Tibet, Togo, Trinidad-Tobago, Tunisia, Turkey-in-Europe, Turks-Caicos Is., Uganda, Uruguay, Utah, Venezuela, Venezuelan Antilles, West Himalaya, Western Australia, Windward Is., Zambia, Zaïre, Zimbabwe

    Introduced into:

    Canary Is., Cocos (Keeling) Is., East Aegean Is., France, Great Britain, Hawaii, Mauritius, New Caledonia, Portugal, Sardegna, South Australia, St.Helena, Victoria

    Common Names

    English
    Daisy lawn, Fog grass

    Phyla nodiflora (L.) Greene appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Meikle, R.D. [1361], Nigeria 11719.000
    Gossweiler, J. [s.n.], Angola K001009801
    Gossweiler, J. [s.n.], Angola K001009802
    Ward, C.J. [44], Angola K001009799
    Welwitsch [5694], Angola K001009805
    Welwitsch [5638], Angola K001009806
    Bourgeau, E. [1641], Spain K000215598
    Pearson, H.H.W. [2177], Angola K001009807
    Pearson, H.H.W. [2177], Angola K001009808
    Baum, H. [582], Angola K001009800
    Reverchon, E. [440], Spain K000215599
    Curror [s.n.], Angola K001009803
    Menezes, A. [3794], Angola K001009804
    Tackholm, V. [8977], Jordan K000318313
    Wallich, N. [Cat. no. 1824], India Zappania nodiflora K001114304
    s.coll. [Cat. no. 1824] Zappania nodiflora K001114305
    s.coll. [Cat. no. 1824] Zappania nodiflora K001114306
    s.coll. [Cat. no. 1824] Zappania nodiflora K001114307
    s.coll. [Cat. no. 1824] Zappania nodiflora K001114308
    s.coll. [Cat. no. 1824] Zappania nodiflora K001114309
    s.coll. [Cat. no. 1824], India Zappania nodiflora K001114310
    Mar 1, 2013 Glaziou, A.F.M. [13065], Brazil Lippia nodiflora K000925067
    Ribas, O.S. [1846], Brazil Lippia nodiflora K000925069
    Sellow, F. [1451], Brazil Lippia nodiflora K000925068
    Sello [2420], Brazil Lippia nodiflora K000925070
    Sellow [s.n.], Brazil Lippia nodiflora K000925066
    Glaziou, A.F.M. [13066], Brazil Lippia nodiflora K000925064
    Hatschbach, G.G. [50069], Brazil Lippia nodiflora K000925065

    First published in Pittonia 4: 46 (1899)

    Accepted by

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    • Vladimirov, V., Dane, F. & Kit Tan (2015). New floristic records in the Balkans: 26 Phytologia Balcanica 21: 53-91.
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    • Shaheen, H., Qureshi, R., Akram, A., Gulfraz, M. & Potter, D. (2014). A preliminary floristic checklist of Thal desert Punjab, Pakistan Pakistan Journal of Botany 46: 13-18.
    • Cossu, T.A, Camarda, I. & Brundu, G. (2014). A catalogue of non-native weeds in irrigated crops in Sardinia (Italy) Webbia; Raccolta de Scritti Botanici 69: 145-156.
    • Dimopoulos, P., Raus, T., Bergmeier, E., Constantinidis, T., Iatrou, G., Kokkini, S., Strid, A., & Tzanoudakis, D. (2013). Vascular plants of Greece. An annotated checklist: 1-372. Botanic gardens and botanical museum Berlin-Dahlem, Berlin and Hellenic botanical society, Athens.
    • Leti, M., Hul, S., Fouché, J.-G., Cheng, S.K. & David, B. (2013). Flore photographique du Cambodge: 1-589. Éditions Privat, Toulouse.
    • Davidse, G. & al. (eds.) (2012). Flora Mesoamericana 4(2): 1-533. Universidad Nacional Autónoma de México, México, D.F.
    • Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies Smithsonian Contributions to Botany 98: 1-1192.
    • O'Leary, N. & Múlgura, M.E. (2011). A taxonomic revision of the genus Phyla (Verbenaceae) Annals of the Missouri Botanical Garden 98: 578-596.
    • Idárraga-Piedrahita, A., Ortiz, R.D.C., Callejas Posada, R. & Merello, M. (eds.) (2011). Flora de Antioquia: Catálogo de las Plantas Vasculares 2: 1-939. Universidad de Antioquia, Medellín.
    • Pandey, R.P. (2009). Floristic diversity of Ferrargunj forest area in South Andaman Journal of Economic and Taxonomic Botany 33: 747-768.
    • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
    • Hokche, O., Berry, P.E. & Huber, O. (eds.) (2008). Nuevo Catálogo de la Flora Vascular de Venezuela: 1-859. Fundación Instituto Botánico de Venezuela.
    • Nelson Sutherland, C.H. (2008). Catálogo de las plantes vasculares de Honduras. Espermatofitas: 1-1576. SERNA/Guaymuras, Tegucigalpa, Honduras.
    • Hedberg, I., Kelbessa, E., Edwards, S., Demissew, S. & Persson, E. (eds.) (2006). Flora of Ethiopia and Eritrea 5: 1-690. The National Herbarium, Addis Ababa University, Ethiopia & The Department of Systematic Botany, Upps.
    • Fernandes, R.B. (2006). Contribução para o conhecimento das Avicenniaceae e Verbenaceae da Flora de Angola Garcia de Orta, Série de Botânica 17(2): 5-68.
    • Raffaelli, M., Mosti, S. & Tardelli, M. (2006). Boswellia sacra Flueck. (Burseraceae) in the Hasik area (Eastern Dhofar, Oman) and a list of surrounding flora Webbia; Raccolta de Scritti Botanici 61: 245-251.
    • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
    • Thulin, M. (ed.) (2006). Flora of Somalia 3: 1-626. The Royal Botanic Gardens, Kew.
    • Fernandes, R. & Diniz, M.A. (2005). Avicenniaceae, Nesogenaceae, Verbenaceae and Lamiaceae (subfams, Viticoideae and Ajugoideae) Flora Zambesiaca 8(7): 1-161. Royal Botanic Gardens, Kew.
    • Miller, A.G. & Morris, M. (2004). Ethnoflora of Soqotra Archipelago: 1-759. The Royal Botanic Garden, Edinburgh.
    • Govaerts, R. (2003). World Checklist of Selected Plant Families Database in ACCESS: 1-216203. The Board of Trustees of the Royal Botanic Gardens, Kew.
    • Turner, I.M. (1995 publ. 1997). A catalogue of the Vascular Plants of Malaya Gardens' Bulletin Singapore 47(2): 347-655.
    • Cirilo, N. & Proctor, G.R. (1994). Vascular plants of the Caribbean Swan islands of Honduras Brenesia 41-42: 73-80.
    • Friedmann, F. (1994). Flore des Seychelles Dicotylédones: 1-663. ORSTOM éditions.
    • MacKee, H.S. (1994). Catalogue des plantes introduites et cultivées en Nouvelle-Calédonie, ed. 2: 1-164. Museum national d'histoire naturelle, Paris.
    • Barry, J. P. & Celles, J.S. (1991). Flore de Mauritanie 1: 1-359. Centre Regional de Documentation Pedagogique, Nice.
    • Lebrun, J.P., Toutain, B., Gaston, A. & Boudet, G. (1991). Catalogue des Plantes Vasculaires du Burkina Faso: 1-341. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
    • Webb, C.J., Sykes, W.R & Garnock-Jones, P.J. (1988). Flora of New Zealand 4: 1-1365. R.E.Owen, Government Printer, Wellington.
    • Brunel, J.F., Hiepo, P. & Scholz, H. (eds.) (1984). Flore Analytique du Togo Phanérogames: 1-751. GTZ, Eschborn.

    Literature

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    Kew Backbone Distributions
    • Shaheen, H., Qureshi, R., Akram, A., Gulfraz, M. & Potter, D. (2014). A preliminary floristic checklist of Thal desert Punjab, Pakistan Pakistan Journal of Botany 46: 13-18.
    • Cossu, T.A, Camarda, I. & Brundu, G. (2014). A catalogue of non-native weeds in irrigated crops in Sardinia (Italy) Webbia; Raccolta de Scritti Botanici 69: 145-156.
    • Dobignard, A. & Chatelain, C. (2013). Index synonymique de la flore d'Afrique du nord 5: 1-451. Éditions des conservatoire et jardin botaniques, Genève.
    • Dimopoulos, P., Raus, T., Bergmeier, E., Constantinidis, T., Iatrou, G., Kokkini, S., Strid, A., & Tzanoudakis, D. (2013). Vascular plants of Greece. An annotated checklist: 1-372. Botanic gardens and botanical museum Berlin-Dahlem, Berlin and Hellenic botanical society, Athens.
    • Leti, M., Hul, S., Fouché, J.-G., Cheng, S.K. & David, B. (2013). Flore photographique du Cambodge: 1-589. Éditions Privat, Toulouse.
    • Hassler, M. (2012). Flora of Rhodes. Systematic list of flora of Rhodes http://www.rz.uni-karlsruhe.de/~db111/flora/rhodos/list.php.
    • Lambdon, P. (2012). Flowering plants & ferns of St Helena: 1-624. Pisces publications for St Helena nature conservation group.
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    • Pandey, R.P. (2009). Floristic diversity of Ferrargunj forest area in South Andaman Journal of Economic and Taxonomic Botany 33: 747-768.
    • Hokche, O., Berry, P.E. & Huber, O. (eds.) (2008). Nuevo Catálogo de la Flora Vascular de Venezuela: 1-859. Fundación Instituto Botánico de Venezuela.
    • Raffaelli, M., Mosti, S. & Tardelli, M. (2006). Boswellia sacra Flueck. (Burseraceae) in the Hasik area (Eastern Dhofar, Oman) and a list of surrounding flora Webbia; Raccolta de Scritti Botanici 61: 245-251.
    • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
    • Thulin, M. (ed.) (2006). Flora of Somalia 3: 1-626. The Royal Botanic Gardens, Kew.
    • Fernandes, R. & Diniz, M.A. (2005). Avicenniaceae, Nesogenaceae, Verbenaceae and Lamiaceae (subfams, Viticoideae and Ajugoideae) Flora Zambesiaca 8(7): 1-161. Royal Botanic Gardens, Kew.
    • Turner, I.M., Xing, F. & Corlett, R.T. (2000). An annotated check-list of the vascular plants of the South China Sea and its shores The Raffles bulletin of Zoology, Suppl. 8: 23-116.
    • Sheppard, C.R.C. & Seaward, M.R.D. (eds.) (1999). Ecology of the Chagos archipelago: 1-350. Westbury Academic & Scientific Publishing, Otley.
    • Ananda Rao, T. & Ellis, J.L. (1995). Flora of Lakshadweep islands off the Malabar coast, peninsular India, with emphasis on phytogeographical distribution of plants Journal of Economic and Taxonomic Botany 19: 235-250.
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    • MacKee, H.S. (1994). Catalogue des plantes introduites et cultivées en Nouvelle-Calédonie, ed. 2: 1-164. Museum national d'histoire naturelle, Paris.
    • Barker, R.M. & Telford, I.R.H. (1993). Fl. Australia Oceanic Islds. Convolvs Flora of Australia 50: 342-353. Australian Government Publishing Service, Canberra.
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    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of West Tropical Africa
    Flora of West Tropical Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2018. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2018. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Plants and People Africa
    Common Names from Plants and People Africa http://www.plantsandpeopleafrica.com/
    © Plants and People Africa http://www.plantsandpeopleafrica.com http://creativecommons.org/licenses/by/3.0/