- Family: Lamiaceae Martinov
Ajuga L.
[FTEA]
Lamiaceae (Labiatae), A.J. Paton, G. Bramley, O. Ryding, R.M. Polhill, Y.B. Harvey, M. Iwarsson, F. Willis, P.B. Phillipson, K. Balkwill, C.W. Lukhoba, D.F. Otieno, & R.M. Harley. Ajuga, C.W. Lukhoba. Flora of Tropical East Africa. 2009
- Morphology General Habit
- Perennial, annual and biennial herbs (rarely subshrubs)
- Morphology Reproductive morphology Inflorescences
- Inflorescence of dense, verticillate cymes, sometimes reduced to solitary flowers or merging to form a terminal, spiciform thyrse
- Morphology Reproductive morphology Flowers Calyx
- Calyx actinomorphic or weakly zygomorphic (posterior lobe reduced), not accrescent, 5-lobed
- Morphology Reproductive morphology Flowers Corolla
- Corolla highly variable in colour, 2-lipped, marcescent, persisting as a sheath around fruit; posterior lip short, much shorter than anterior, 2-lobed, from which the sex organs emerge; anterior lip 3-lobed, median lobe very large, broadening distally and usually emarginate or bipartite
- Morphology Reproductive morphology Flowers Androecium Stamens
- Stamens 4, didynamous, usually slightly exserted; filaments fused to corolla for more than half their length, and covered with short hyaline hairs all over; anther thecae divergent to divaricate, confluent at dehiscence
- Morphology Reproductive morphology Flowers Disc
- Disk poorly to well developed, usually asymmetrical
- Morphology Reproductive morphology Flowers Gynoecium Ovary
- Ovary shallowly 4-lobed; style subterminal, ± equalling filaments, bifid with equal or unequal lobes
- Morphology Reproductive morphology Fruits
- Fruit usually a schizocarp of 4 nutlets (rarely drupaceous, 4-lobed); nutlets obovoid, reticulately foveolate, glabrous, with prominent lateral attachment-scar; endosperm present.
- Distribution
- Range: A genus of about 30 species, mostly tropical and subtropical in the Old World.
[LKGF]
Harley, R.M. et al. (2004). Labiatae, in The Families and Genera of Vascular Plants (K. Kubitzki, ed. in chief) VI: 167-275. Springer-Verlag, Berlin, Heidelberg, New York.
- Morphology General Habit
- Perennial, annual, and biennial herbs (rarely subshrubs)
- Morphology Reproductive morphology Inflorescences
- Inflorescence of dense, verticillastrate cymes, sometimes reduced to solitary flowers or merging to form a terminal, spiciform or subcapitate thyrse
- Morphology Reproductive morphology Flowers Calyx
- Calyx actinomorphic or weakly zygomorphic (posterior lobe reduced), not accrescent, 5-lobed
- Morphology Reproductive morphology Flowers Corolla
- Corolla highly variable in colour, 2-lipped or 1-lipped and then with only anterior lip present, marcescent, persisting as a sheath around fruit, posterior lip short or obsolescent, usually 2-lobed or retuse, anterior lip 3-lobed, median lobe very large,
- Morphology Reproductive morphology Flowers Androecium Stamens
- Stamens 4, didynamous, usually slightly exserted, thecae divergent to divaricate, confluent at dehiscence
- Morphology Reproductive morphology Flowers Pollen
- Pollen suprareticulate, columellae simple or branched
- Morphology Reproductive morphology Flowers Gynoecium Ovary
- Ovary shallowly 4-lobed
- Morphology Reproductive morphology Flowers Gynoecium Stigma
- Stigma lobes equal to unequal
- Morphology Reproductive morphology Flowers Disc
- Disc poorly to well developed, usually asymmetrical
- Morphology Reproductive morphology Fruits
- Fruit usually a schizocarp of 4 nutlets (rarely drupaceous, 4-lobed), nutlets oblate to obovoid, reticulately ridged to foveolate, usually glabrous, attachment-scar lateral, 0.55-0.85x as long as nutlet
- Morphology Reproductive morphology Seeds Endosperm
- Endosperm present.
- Note
- 2n = 32 in most species (16, 24, 28, 30, 62, 64, ca. 86). Forty to 50 species, paleotemperate, mostly Eurasian.
- Distribution
- Native to: Europe, Northern Africa, Macaronesia, West-Central Tropical Africa, Northeast Tropical Africa, East Tropical Africa, Southern Africa (doubtfully present in South Tropical Africa), Western Indian Ocean, Siberia, Russian Far East, Middle Asia, Ca
>
Native to:
Afghanistan, Albania, Algeria, Amur, Assam, Austria, Baleares, Baltic States, Bangladesh, Belarus, Belgium, Bulgaria, Canary Is., Cape Provinces, Cape Verde, Central European Rus, China North-Central, China South-Central, China Southeast, Chita, Corse, Cyprus, Czechoslovakia, Denmark, East Aegean Is., East European Russia, East Himalaya, Egypt, Eritrea, Ethiopia, Finland, France, Free State, Germany, Great Britain, Greece, Hainan, Hungary, Iceland, India, Inner Mongolia, Iran, Iraq, Ireland, Italy, Japan, Kazakhstan, Kenya, Khabarovsk, Kirgizstan, Korea, Kriti, Krym, Kuril Is., KwaZulu-Natal, Laos, Lebanon-Syria, Lesotho, Libya, Madagascar, Madeira, Maluku, Manchuria, Morocco, Myanmar, Nansei-shoto, Nepal, Netherlands, New Guinea, New South Wales, North Caucasus, North European Russi, Northern Provinces, Northwest European R, Norway, Ogasawara-shoto, Pakistan, Palestine, Philippines, Poland, Portugal, Primorye, Qinghai, Queensland, Romania, Sakhalin, Sardegna, Saudi Arabia, Sicilia, Sinai, South Australia, South European Russi, Spain, Sudan, Sulawesi, Swaziland, Sweden, Switzerland, Tadzhikistan, Taiwan, Tanzania, Tasmania, Thailand, Tibet, Transcaucasus, Tunisia, Turkey, Turkey-in-Europe, Turkmenistan, Uganda, Ukraine, Victoria, Vietnam, West Himalaya, Western Sahara, Yemen, Yugoslavia, Zaïre
Introduced into:
Alabama, Arkansas, British Columbia, Connecticut, Delaware, District of Columbia, Florida, Georgia, Illinois, Indiana, Iowa, Irkutsk, Kansas, Kentucky, Louisiana, Maine, Maryland, Massachusetts, Michigan, Minnesota, Mississippi, New Hampshire, New Jersey, New York, New Zealand North, New Zealand South, Newfoundland, North Carolina, Nova Scotia, Ohio, Ontario, Oregon, Pennsylvania, Québec, Rhode I., South Carolina, Tennessee, Texas, Utah, Venezuela, Vermont, Virginia, Washington, West Virginia, Wisconsin
- Ajuga × adulterina Wallr.
- Ajuga arabica P.H.Davis
- Ajuga australis R.Br.
- Ajuga austroiranica Rech.f.
- Ajuga × bastarda Makino
- Ajuga bombycina Boiss.
- Ajuga boninsimae Maxim.
- Ajuga × borbasiana (Rouy) Prain
- Ajuga brachystemon Maxim.
- Ajuga campylantha Diels
- Ajuga campylanthoides C.Y.Wu & C.Chen
- Ajuga chamaecistus Ging. ex Benth.
- Ajuga chamaepitys (L.) Schreb.
- Ajuga chasmophila P.H.Davis
- Ajuga ciliata Bunge
- Ajuga davisiana Kit Tan & Yildiz
- Ajuga decaryana Danguy ex R.A.Clement
- Ajuga decumbens Thunb.
- Ajuga dictyocarpa Hayata
- Ajuga flaccida Baker
- Ajuga forrestii Diels
- Ajuga genevensis L.
- Ajuga grandiflora Stapf
- Ajuga × hybrida A.Kern.
- Ajuga incisa Maxim.
- Ajuga integrifolia Buch.-Ham. ex D.Don
- Ajuga iva (L.) Schreb.
- Ajuga japonica Miq.
- Ajuga laxmannii (Murray) Benth.
- Ajuga leucantha Lukhoba
- Ajuga linearifolia Pamp.
- Ajuga lobata D.Don
- Ajuga lupulina Maxim.
- Ajuga macrosperma Wall. ex Benth.
- Ajuga makinoi Nakai
- Ajuga × mixta Makino
- Ajuga mollis Gladkova
- Ajuga multiflora Bunge
- Ajuga nipponensis Makino
- Ajuga novoguineensis A.J.Paton & R.J.Johns
- Ajuga nubigena Diels
- Ajuga oblongata M.Bieb.
- Ajuga oocephala Baker
- Ajuga ophrydis Burch. ex Benth.
- Ajuga orientalis L.
- Ajuga ovalifolia Bureau & Franch.
- Ajuga pantantha Hand.-Mazz.
- Ajuga parviflora Benth.
- Ajuga piskoi Degen & Bald.
- Ajuga postii Briq.
- Ajuga × pseudopyramidalis Schur
- Ajuga pygmaea A.Gray
- Ajuga pyramidalis L.
- Ajuga relicta P.H.Davis
- Ajuga reptans L.
- Ajuga robusta Baker
- Ajuga salicifolia (L.) Schreb.
- Ajuga saxicola Assadi & Jamzad
- Ajuga sciaphila W.W.Sm.
- Ajuga shikotanensis Miyabe & Tatew.
- Ajuga sinuata R.Br.
- Ajuga spectabilis Nakai
- Ajuga taiwanensis Nakai ex Murata
- Ajuga tenorei C.Presl
- Ajuga turkestanica (Regel) Briq.
- Ajuga vesiculifera Herder
- Ajuga vestita Boiss.
- Ajuga xylorrhiza Kit Tan
- Ajuga yesoensis Maxim. ex Franch. & Sav.
- Ajuga zakhoensis Rech.f.
Ajuga L. appears in other Kew resources:
First published in Sp. Pl.: 561 (1753)
Accepted by
- Govaerts, R. (2003). World Checklist of Selected Plant Families Database in ACCESS: 1-216203. The Board of Trustees of the Royal Botanic Gardens, Kew.
- Govaerts, R. (1995). World Checklist of Seed Plants 1(1, 2): 1-483, 1-529. MIM, Deurne.
Literature
Flora of Tropical East Africa
- Kubitzki, Fam. Gen. Vasc. Pl. 7: 201 (2004)
- Gen. Pl., ed. 5: 246 (1754)
- Sp. Pl.: 561 (1753)
Lamiaceae Key Genus Fact Sheets
- Harley RM, Atkins S, Budantsev AL, Cantino PD, Conn BJ, Grayer R, Harley MM, de Kok RPJ, Krestovskaja T, Morales R, Paton AJ, Ryding O, and Upson T. 2004. Labiatae, in The Families and Genera of Vascular Plants (K. Kubitzki, ed. in chief) VI: 167-275. Sp
- Bulga Kuntze, Revis. Gen. Pl. 2: 512 (1891).
- Rosenbachia Regel, Trudy Imp. S.-Peterburgsk. Bot. Sada 9: 613 (1886).
- Abiga St.-Lag., Ann. Soc. Bot. Lyon 7: 85 (1880).
- Phleboanthe Tausch, Flora 11: 322 (1828).
- Moscharia Forssk., Fl. Aegypt.-Arab.: 158 (1775).
- Chamaepitys Hill, Brit. Herb.: 371 (1756).
- Gen. Pl. ed. 5: 246 (1754).
- Bugula Mill., Gard. Dict. Abr. ed. 4 (1754).
- Ajuga L., Sp. Pl. 2: 561 (1753)
Flora of Tropical East Africa
Flora of Tropical East Africa
http://creativecommons.org/licenses/by-nc-sa/3.0
Kew Backbone Distributions
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0
Kew Names and Taxonomic Backbone
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0
Lamiaceae Key Genus Fact Sheets
Nina Davies, Gemma Bramley and Don Kirkup, Royal Botanic Gardens, Kew
http://creativecommons.org/licenses/by/3.0