1. Family: Petiveriaceae C.Agardh
    1. Genus: Rivina Plum. ex L.
      1. Rivina humilis L.

        This species is accepted, and its native range is Tropical & Subtropical America.

    [FZ]

    Phytolaccaceae, B. L. Stannard. Flora Zambesiaca 9:1. 1988

    Habit
    Herbs or (?)sub-shrubs 0.30–1.2 m. tall.
    Stem
    Stems glabrous or slightly pubescent at nodes.
    Leaves
    Leaves: lamina 2.0–10.0 × 1.0–4.5 cm., elliptic to ovate, cuneate to rounded at base, usually acuminate, glabrescent to pubescent above and below, often particularly along main veins; petioles 8–50 mm. long, thickly pubescent all round (not in the Flora Zambesiaca area) to pubescent along upper side only or glabrescent.
    Inflorescences
    Inflorescences to 13 cm. long.
    Flowers
    Flowers glabrous; pedicels to 5 mm. long.
    Bracts
    Bract 1, 1–1.2 mm. long, subulate.
    Bracteoles
    Bracteoles 2, appressed to sepals, triangular, 0.2–0.25 × 0.2–0.25 mm.
    Calyx
    Sepals c. 2 × 0.8–1 mm., elliptic, rounded to slightly acute at apex, spreading at anthesis, green, white or pinkish.
    Stamens
    Stamens 1.5–1.8 mm. long; filaments often persistent in fruiting; anthers 0.4–0.5 mm. long.
    Ovary
    Ovary 0.5–0.6 mm. in diam.
    Style
    Style 0.4–0.5 mm. long, filiform.
    Fruits
    Fruit 2.5–3 mm. in diam.
    Seeds
    Seed c. 2 mm. in diam.
    [FTEA]

    Phytolaccaceae, R. M. Polhill. Flora of Tropical East Africa. 1971

    Habit
    Erect herb 3–10 dm. tall, glabrous to densely hairy on the branchlets, but commonly with short weak bristly hairs only on the ridges of the youngest parts.
    Leaves
    Leaf-blades lanceolate to elliptic-ovate or ovate, mostly 5–12(–17) cm. long, 2.5–6(–8) cm. wide, acuminate, broadly cuneate to rounded or cordate and unequal-sided at the base, glabrous to densely hairy, but commonly with small hairs only on the midrib above and on the nerves beneath; crystals often visible as short raised lines beneath; primary lateral nerves 6–12 on either side; petiole slender, 1.5–4.5(–6) cm. long, adaxially pubescent.
    Inflorescences
    Racemes 3–5(–11 in fruit) cm. long, sublaxly many-flowered; bracts caudate from a broader basal part, ± 1 mm. long, caducous; pedicels slender, becoming recurved, 1.5–3(–5 in fruit) mm. long, with minute bracteoles on the upper part.
    Calyx
    Sepals green, white or pinkish, ± 2 mm. long, spreading at anthesis.
    Stamens
    Stamens a little shorter than the sepals.
    Ovary
    Ovary a little shorter than the sepals.
    Fruits
    Berry 3–4 mm. in diameter, ripening red.
    Seeds
    Seeds 2–2.4 mm. across, usually shortly hairy.
    Figures
    Fig. 3.
    Habitat
    In riverine forest and grassland; 550 m.
    Distribution
    T3 native of tropical and subtropical America, locally naturalized in the Old World tropics

    Images

    Distribution

    Native to:

    Argentina Northeast, Argentina Northwest, Arizona, Arkansas, Aruba, Bahamas, Belize, Bolivia, Brazil North, Brazil Northeast, Brazil South, Brazil Southeast, Brazil West-Central, Cayman Is., Colombia, Costa Rica, Cuba, Dominican Republic, El Salvador, Florida, French Guiana, Guatemala, Guyana, Haiti, Honduras, Jamaica, Leeward Is., Louisiana, Mexico Central, Mexico Gulf, Mexico Northeast, Mexico Northwest, Mexico Southeast, Mexico Southwest, Netherlands Antilles, New Mexico, Nicaragua, Oklahoma, Panamá, Paraguay, Peru, Puerto Rico, Southwest Caribbean, Suriname, Texas, Trinidad-Tobago, Turks-Caicos Is., Uruguay, Venezuela, Venezuelan Antilles, Windward Is.

    Introduced into:

    Bangladesh, Bermuda, Cape Provinces, Cape Verde, Chagos Archipelago, China Southeast, Cocos (Keeling) Is., Fiji, Galápagos, India, Ivory Coast, Jawa, KwaZulu-Natal, Malaya, Mauritius, Mozambique, New Caledonia, Norfolk Is., Northern Provinces, Ogasawara-shoto, Réunion, Seychelles, Somalia, Sri Lanka, Sumatera, Taiwan, Tanzania, Thailand, Tonga, Tuvalu, Vanuatu, Zaïre, Zimbabwe

    Rivina humilis L. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Chase, M.W. [10978] K000696072
    Ruiz, H. [10/74], Peru Rivina humilis var. glabra K000601219

    First published in Sp. Pl.: 121 (1753)

    Accepted by

    • Villaseñor, J.L. (2016). Checklist of the native vascular plants of Mexico Revista Mexicana de Biodiversidad 87: 559-902.
    • Bernal, R., Gradstein, R.S. & Celis, M. (eds.) (2016). Catálogo de plantas y líquenes de Colombia 1-2: 1-3068. Libro impreso.
    • Kiew, R. & al. (eds.) (2015). Flora of Peninsular Malaysia. Series II: Seed Plants, volume 5 Malayan Forest Records 49: 1-319.
    • Sikarwar, R.L.S. (2014). Angiosperm diversity assessment of Chitrakootthe legendary place of Vindhyan range, India Journal of Economic and Taxonomic Botany 38: 563-619.
    • Jørgensen, P.M., Nee, M.H. & Beck., S.G. (eds.) (2013). Catálogo de las plantas vasculares de Bolivia Monographs in Systematic Botany from the Missouri Botanical Garden 127: 1-1741. Missouri Botanical Garden.
    • Mostaph, M.K. & Uddin, S.B. (2013). Dictionary of plant names of Bangladesh, Vasc. Pl.: 1-434. Janokalyan Prokashani, Chittagong, Bangladesh.
    • Allred, K.W. (2012). Flora Neomexicana, ed. 2, 1: 1-599. Range Science Herbarium, Las Cruces, New Mexico.
    • López Patiño, E.J., Szeszko, D.R., Rascala Pérez, J. & Beltrán Retis, A.S. (2012). The flora of the Tenacingo-Malinalco-Zumpahuacán protected natural area, state of Mexico, Mexico Harvard Papers in Botany 17: 65-167.
    • Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies Smithsonian Contributions to Botany 98: 1-1192.
    • Garcia-Mendoza, A.J. & Meave, J.A. (eds.) (2012). Diversidad florística de Oaxaca: de musgos a angiospermas (colecciones y listas de especies), ed. 2: 1-351. Instituto de Biología, Universidad Nacional Autónoma de México.
    • van Proosdij, A.S.J. (2012). Arnoldo's Zakflora, ed. 4: 1-318. Walburg Pers, Zutphen.
    • Idárraga-Piedrahita, A., Ortiz, R.D.C., Callejas Posada, R. & Merello, M. (eds.) (2011). Flora de Antioquia: Catálogo de las Plantas Vasculares 2: 1-939. Universidad de Antioquia, Medellín.
    • Paña-Chocarro, M.C., Espada-Mateos, C., Vera, M., Céspedes, G. & Knapp, S. (2010). Updated checklist of vascular plants of the Mbaracayú Forest Nature Reserve (Reserva Natural del Bosque Mbaracayú), Paraguay Phytotaxa 12: 1-224.
    • Vikraman, R/R/, Pandurangan, A.G. & Thulasidas, G. (2008). A study on the garden escaped exotics of Thiruvananthapuram district, Kerala Journal of East African Natural History 32: 765-781.
    • Zuloaga, F.O., Morrone, O. , Belgrano, M.J., Marticorena, C. & Marchesi, E. (eds.) (2008). Catálogo de las Plantas Vasculares del Cono Sur Monographs in Systematic Botany from the Missouri Botanical Garden 107: 1-3348. Missouri Botanical Garden.
    • Tseng, Y.H., Wang, C.C. & Chen, Y.T. (2008). Rivina humilis L. (Phytolaccaceae), a newly naturalised plant in Taiwan Taiwania 53: 417-419.
    • Hokche, O., Berry, P.E. & Huber, O. (eds.) (2008). Nuevo Catálogo de la Flora Vascular de Venezuela: 1-859. Fundación Instituto Botánico de Venezuela.
    • Nelson Sutherland, C.H. (2008). Catálogo de las plantes vasculares de Honduras. Espermatofitas: 1-1576. SERNA/Guaymuras, Tegucigalpa, Honduras.
    • Sant'Ana Melhem, T., das Graças Lapa Wanderley, M., Ehlin Martins, S., Jung-Mendaçolli, S.L., Shepherd, G.J. & Kirizawa, M. (eds.) (2007). Flora Fanerogâmica do Estado de São Paulo 5: 1-476. Instituto de Botânica, São Paulo.
    • Hammel, B.E. & al. (eds.) (2007). Manual de Plantas de Costa Rica, volumen VI, Dicotyledóneas (Haloragaceae-Phytolaccaceae) Monographs in Systematic Botany from the Missouri Botanical Garden 111: 1-933. Missouri Botanical Garden.
    • Correa A., Mireya D. Galdames, Carmen Correa A., M. D., C. Galdames & M. S. de Stapf (2004). Catálogo de las Plantas Vasculares de Panamá: 1-599. Smithsonian Tropical Research Institute.
    • Germishuizen, G. & Meyer, N.L. (eds.) (2003). Plants of Southern Africa an annotated checklist Strelitzia 14: 1-1231. National Botanical Institute, Pretoria.
    • Flora of North America Editorial Committee (2003). Flora of North America North of Mexico 4: 1-559. Oxford University Press, New York, Oxford.
    • Aké Assi, L. (2002). Flore de la Côte-d'Ivoire: catalogue systématique, biogéographie et écologie. II Boissiera 58: 1-401.
    • Stevens, W.D., Ulloa U., C., Pool, A. & Montiel, O.M. (2001). Flora de Nicaragua Monographs in Systematic Botany from the Missouri Botanical Garden 85: i-xlii, 1-2666. Missouri Botanical Garden.
    • Balick, M. J., M. H. Nee & D.E. Atha (2000). Checklist of the vascular plants of Belize Memoirs of the New York Botanical Garden 85: 1-246.
    • Boggan, J. Funck, V. & Kelloff, C. (1997). Checklist of the Plants of the Guianas (Guyana, Surinam, Franch Guiana) ed. 2: 1-238. University of Guyana, Georgetown.
    • Gonzalez, F., Nelson Diaz, J. & Lowry, P. (1995). Flora Illustrada de San Andrés y Providencia: 1-281. Universidad Nacional de Colombia, Instituto de Ciencias Naturales, Colombia.
    • Cirilo, N. & Proctor, G.R. (1994). Vascular plants of the Caribbean Swan islands of Honduras Brenesia 41-42: 73-80.
    • Orchard, A.E. (ed.) (1994). Oceanic Islands 1 Flora of Australia 49: 1-681. Australian Government Publishing Service, Canberra.
    • Bosser, J. & al. (eds.) (1994). Flore des Mascareignes 136-148: 1. IRD Éditions, MSIRI, RBG-Kew, Paris.
    • MacKee, H.S. (1994). Catalogue des plantes introduites et cultivées en Nouvelle-Calédonie, ed. 2: 1-164. Museum national d'histoire naturelle, Paris.
    • George, A.S., Orchard, A.E. & Hewson, H.J. (eds.) (1993). Oceanic islands 2 Flora of Australia 50: 1-606. Australian Government Publishing Service, Canberra.
    • Brako, L. & Zarucchi, J.L. (1993). Catalogue of the Flowering Plants and Gymnosperms of Peru Monographs in Systematic Botany from the Missouri Botanical Garden 45: i-xl, 1-1286. Missouri Botanical Garden.
    • Hansen, A. & Sunding, P. (1993). Flora of Macaronesia. Checklist of vascular plants. 4. revised edition Sommerfeltia 17: 1-295.
    • Thulin, M. (ed.) (1993). Flora of Somalia 1: 1-493. The Royal Botanic Gardens, Kew.
    • Launert, E. (ed.) (1988). Flora Zambesiaca 9(1): 1-179. Royal Botanic Gardens, Kew.
    • Smith, A.C. (1981). Flora Vitiensis Nova. A new flora for Fiji (Spermatophytes only) 2: 1-810. Pacific Tropical Botanical Garden, Lawai.
    • Polhill, R.M. (1971). Flora of Tropical East Africa, Phytolaccaceae: 1-8.

    Literature

    Kew Backbone Distributions
    • Jaramillo Díaz, P. & Guézou, A. (2017). CDF Checklist of Galapagos Vascular Plants - FCD Lista de especies de Plantas Vasculares de Galápagos http://www.darwinfoundation.org/datazone/checklists/vascular-plants/.
    • Forzza, R.C., Zappi, D. & Souza, V.C. (2016-continuously updated). Flora do Brasil 2020 em construção http://reflora.jbrj.gov.br/reflora/listaBrasil/ConsultaPublicaUC/ResultadoDaConsultaNovaConsulta.do.
    • Villaseñor, J.L. (2016). Checklist of the native vascular plants of Mexico Revista Mexicana de Biodiversidad 87: 559-902.
    • Kiew, R. & al. (eds.) (2015). Flora of Peninsular Malaysia. Series II: Seed Plants, volume 5 Malayan Forest Records 49: 1-319.
    • Jørgensen, P.M., Nee, M.H. & Beck., S.G. (eds.) (2013). Catálogo de las plantas vasculares de Bolivia Monographs in Systematic Botany from the Missouri Botanical Garden 127: 1-1741. Missouri Botanical Garden.
    • Mostaph, M.K. & Uddin, S.B. (2013). Dictionary of plant names of Bangladesh, Vasc. Pl.: 1-434. Janokalyan Prokashani, Chittagong, Bangladesh.
    • López Patiño, E.J., Szeszko, D.R., Rascala Pérez, J. & Beltrán Retis, A.S. (2012). The flora of the Tenacingo-Malinalco-Zumpahuacán protected natural area, state of Mexico, Mexico Harvard Papers in Botany 17: 65-167.
    • Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies Smithsonian Contributions to Botany 98: 1-1192.
    • Garcia-Mendoza, A.J. & Meave, J.A. (eds.) (2012). Diversidad florística de Oaxaca: de musgos a angiospermas (colecciones y listas de especies), ed. 2: 1-351. Instituto de Biología, Universidad Nacional Autónoma de México.
    • van Proosdij, A.S.J. (2012). Arnoldo's Zakflora, ed. 4: 1-318. Walburg Pers, Zutphen.
    • Idárraga-Piedrahita, A., Ortiz, R.D.C., Callejas Posada, R. & Merello, M. (eds.) (2011). Flora de Antioquia: Catálogo de las Plantas Vasculares 2: 1-939. Universidad de Antioquia, Medellín.
    • Paña-Chocarro, M.C., Espada-Mateos, C., Vera, M., Céspedes, G. & Knapp, S. (2010). Updated checklist of vascular plants of the Mbaracayú Forest Nature Reserve (Reserva Natural del Bosque Mbaracayú), Paraguay Phytotaxa 12: 1-224.
    • Lejoy, J., Ndjele, M.-B. & Geerinck, D. (2010). Catalogue-flore des plantes vasculaires des districts de Kisangani et de la Tshopo (RD Congo) Taxonomania 30: 1-307.
    • GBIF (2008-continuously updated). Global Biodiversity Information Facility http://www.gbif.org/.
    • Vikraman, R/R/, Pandurangan, A.G. & Thulasidas, G. (2008). A study on the garden escaped exotics of Thiruvananthapuram district, Kerala Journal of East African Natural History 32: 765-781.
    • Zuloaga, F.O., Morrone, O. , Belgrano, M.J., Marticorena, C. & Marchesi, E. (eds.) (2008). Catálogo de las Plantas Vasculares del Cono Sur Monographs in Systematic Botany from the Missouri Botanical Garden 107: 1-3348. Missouri Botanical Garden.
    • Tseng, Y.H., Wang, C.C. & Chen, Y.T. (2008). Rivina humilis L. (Phytolaccaceae), a newly naturalised plant in Taiwan Taiwania 53: 417-419.
    • Hokche, O., Berry, P.E. & Huber, O. (eds.) (2008). Nuevo Catálogo de la Flora Vascular de Venezuela: 1-859. Fundación Instituto Botánico de Venezuela.
    • Nelson Sutherland, C.H. (2008). Catálogo de las plantes vasculares de Honduras. Espermatofitas: 1-1576. SERNA/Guaymuras, Tegucigalpa, Honduras.
    • Sant'Ana Melhem, T., das Graças Lapa Wanderley, M., Ehlin Martins, S., Jung-Mendaçolli, S.L., Shepherd, G.J. & Kirizawa, M. (eds.) (2007). Flora Fanerogâmica do Estado de São Paulo 5: 1-476. Instituto de Botânica, São Paulo.
    • Hammel, B.E. & al. (eds.) (2007). Manual de Plantas de Costa Rica, volumen VI, Dicotyledóneas (Haloragaceae-Phytolaccaceae) Monographs in Systematic Botany from the Missouri Botanical Garden 111: 1-933. Missouri Botanical Garden.
    • Correa A., Mireya D. Galdames, Carmen Correa A., M. D., C. Galdames & M. S. de Stapf (2004). Catálogo de las Plantas Vasculares de Panamá: 1-599. Smithsonian Tropical Research Institute.
    • Germishuizen, G. & Meyer, N.L. (eds.) (2003). Plants of Southern Africa an annotated checklist Strelitzia 14: 1-1231. National Botanical Institute, Pretoria.
    • Aké Assi, L. (2002). Flore de la Côte-d'Ivoire: catalogue systématique, biogéographie et écologie. II Boissiera 58: 1-401.
    • Stevens, W.D., Ulloa U., C., Pool, A. & Montiel, O.M. (2001). Flora de Nicaragua Monographs in Systematic Botany from the Missouri Botanical Garden 85: i-xlii, 1-2666. Missouri Botanical Garden.
    • Balick, M. J., M. H. Nee & D.E. Atha (2000). Checklist of the vascular plants of Belize Memoirs of the New York Botanical Garden 85: 1-246.
    • Sheppard, C.R.C. & Seaward, M.R.D. (eds.) (1999). Ecology of the Chagos archipelago: 1-350. Westbury Academic & Scientific Publishing, Otley.
    • Boggan, J. Funck, V. & Kelloff, C. (1997). Checklist of the Plants of the Guianas (Guyana, Surinam, Franch Guiana) ed. 2: 1-238. University of Guyana, Georgetown.
    • Cirilo, N. & Proctor, G.R. (1994). Vascular plants of the Caribbean Swan islands of Honduras Brenesia 41-42: 73-80.
    • Orchard, A.E. (ed.) (1994). Oceanic Islands 1 Flora of Australia 49: 1-681. Australian Government Publishing Service, Canberra.
    • Bosser, J. & al. (eds.) (1994). Flore des Mascareignes 136-148: 1. IRD Éditions, MSIRI, RBG-Kew, Paris.
    • Castle, G.E. (1994). Flore des Seychelles Dicotylédones: 1-663. Orstom Editions.
    • George, A.S., Orchard, A.E. & Hewson, H.J. (eds.) (1993). Oceanic islands 2 Flora of Australia 50: 1-606. Australian Government Publishing Service, Canberra.
    • Brako, L. & Zarucchi, J.L. (1993). Catalogue of the Flowering Plants and Gymnosperms of Peru Monographs in Systematic Botany from the Missouri Botanical Garden 45: i-xl, 1-1286. Missouri Botanical Garden.
    • Hansen, A. & Sunding, P. (1993). Flora of Macaronesia. Checklist of vascular plants. 4. revised edition Sommerfeltia 17: 1-295.
    • Thulin, M. (ed.) (1993). Flora of Somalia 1: 1-493. The Royal Botanic Gardens, Kew.
    • Launert, E. (ed.) (1988). Flora Zambesiaca 9(1): 1-179. Royal Botanic Gardens, Kew.
    • Kobayashi, S. & Ono, M. (1987). A Revised List of Vascular Plants Indigenous and Introduced to the Bonin (Ogasawara) and the Volcano (Kazan) Islands Ogasawara Research 13: 1-55.
    • Correll, D.S. & Correll, H.B. (1982). Flora of the Bahama Archipelago: 1-1692. J.Cramer, Vaduz.
    • Smith, A.C. (1981). Flora Vitiensis Nova. A new flora for Fiji (Spermatophytes only) 2: 1-810. Pacific Tropical Botanical Garden, Lawai.
    • Polhill, R.M. (1971). Flora of Tropical East Africa, Phytolaccaceae: 1-8.
    Flora of Tropical East Africa
    • Cavaco in Fl. Madag., Phytolaccac.: 2, fig. 1/4, 5 (1954).
    • Backer in Fl. Males., ser. 1, 4: 229, fig. 1 (1951).
    • H. Walt. in A. Engler, Das Pflanzenreich IV. 83: 102, t. 30 (1909).
    • L., Sp. Pl.: 121 (1753).

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2018. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2018. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0