1. Family: Fabaceae Lindl.
    1. Ptycholobium Harms

      1. This genus is accepted, and is native to Africa and Yemen..

    [FZ]

    Flora Zambesiaca Leguminosae subfamily Papillionoideae by R.K. Brummitt

    Habit
     Undershrubs up to 40 cm high. Undershrubs up to 40 cm high.
    Indumentum
    Young stems variously densely pubescent.
    Ovary
    Ovary pubescent; style glabrous.
    Stem
    Young stems variously densely pubescent.
    Leaves
    Leaves digitately trifoliolate (i.e. the terminal one sessile, lacking a rachis) or sometimes unifoliolate, rarely with occasional leaves 5-foliolate with a short rachis; petiole short, up to 7(9) mm long; leaflets ± oblanceolate, usually glabrous above and appressed-pubescent beneath; stipules 1–4 mm long, linear-triangular. Leaves digitately trifoliolate (i.e. the terminal one sessile, lacking a rachis) or sometimes unifoliolate, rarely with occasional leaves 5-foliolate with a short rachis; petiole short, up to 7(9) mm long; leaflets ± oblanceolate, usually glabrous above and appressed-pubescent beneath; stipules 1–4 mm long, linear-triangular.
    Flowers
    Flowers in groups of (1)2–4(7) in axils of foliage leaves; pedicels not exceeding 2 mm; bracteoles absent. Flowers in groups of (1)2–4(7) in axils of foliage leaves; pedicels not exceeding 2 mm; bracteoles absent.
    Calyx
    Calyx 3–5 mm long, tubular to funnel-shaped with 5 subequal acute teeth slightly shorter than the tube, the two upper ones slightly connate at the base, pubescent. Calyx 3–5 mm long, tubular to funnel-shaped with 5 subequal acute teeth slightly shorter than the tube, the two upper ones slightly connate at the base, pubescent.
    Corolla
    Petals up to 7 mm long, usually bluish or purplish; standard spathulate, oblong cuneate or subpandurate, gradually narrowed to the claw, hairy outside; wings with a very slender claw, the limb ± oblong; keel petals with a very slender claw equalling those of the wings, the limb shorter than the claw, ½ to ⅔ as long as the wing limb. Petals up to 7 mm long, usually bluish or purplish; standard spathulate, oblong cuneate or subpandurate, gradually narrowed to the claw, hairy outside; wings with a very slender claw, the limb ± oblong; keel petals with a very slender claw equalling those of the wings, the limb shorter than the claw, ½ to ? as long as the wing limb.
    Stamens
    Stamens united into a tube, the upper one easily detached. Stamens united into a tube, the upper one easily detached.
    Pistil
    Ovary pubescent; style glabrous.
    Fruits
    Pod thin-walled, variously contorted, indehiscent, variously pubescent. Pod thin-walled, variously contorted, indehiscent, variously pubescent.
    [LOWO]

    Legumes of the World. Edited by G. Lewis, B. Schrire, B. MacKinder & M. Lock. Royal Botanic Gardens, Kew. (2005)

    Note

    Relationships among genera of Millettieae have been notoriously difficult to unravel based on traditional morphological evidence and this is exemplified by the alphabetical arrangement of genera in the tribal treatments of Geesink (1981; 1984) and Polhill (1994). Geesink (1981) recognised 44 genera and c. 870 species in tribe Millettieae (as ‘Tephrosieae’) while 43 genera were accounted for in Geesink (1984) and Polhill (1994). The genera recognised, however, varied considerably with only 33 genera in common to both treatments of Geesink, while the list of Polhill (1994) combined elements of Geesink (1981, 1984) with new data accumulated since then. Tephrosia has traditionally comprised some 400 species but this is re-estimated at c. 350 species here.

    The traditional circumscription of the predominantly pantropical and subtropical tribe Millettieae is followed here (Fig. 45), with 45 genera and (904)–909–(914) species being recognised, (i.e. excluding the two genera and 11 species transferred to Brongniartieae, see Table 8), although the concept of what comprises Millettieae sens. strict. is changing rapidly based on evidence from molecular phylogenies. Sequence data for millettioid genera comes from the plastid rbcL gene (Doyle et al., 1997; 2000; Kajita et al., 2001; Hu & Chang, 2003), phytochrome nucleotide genes (Lavin et al., 1998), the plastid trnK-matK region (Hu et al., 2000) and the nuclear ITS region (Hu, 2000; Hu et al., 2002). Molecular data, together with reinterpreted evidence based on chemistry (Evans et al., 1985) and wood anatomy (Gasson et al., 2004), have been the basis for recognising a number of informal suprageneric groupings and for transferring Cyclolobium and Poecilanthe to tribe Brongniartieae (Table 8; Fig. 45).

     The most far-reaching result of the above molecular analyses was that a substantial part of the traditionally circumscribed tribe Phaseoleae is more closely allied to the core-Millettieae than to the Phaseoleae sens. lat. clade (see page 393). Circumscription of a revised tribe Millettieae is not possible at present until genera are more comprehensively sampled; however, a Millettioid sens. strict. group might be expected to include some genera in the basal millettioid and phaseoloid group, Phaseoleae subtribes Diocleinae, Ophrestiinae and in small part the Erythrininae, tribe Abreae and the core-Millettieae (Fig. 45). The basal millettioid and phaseoloid group comprises 17 genera (94 species) that may belong either in the Millettioids sens. strict. or Phaseoleae sens. lat., or to a clade sister to both these groups (e.g., Kajita et al., 2001). The core-Millettieae clade comprises c. 22 genera and c. 777 spp., with some additional generic segregates being necessary within the ‘canavanine group’ (Evans et al., 1985), to accommodate species of Millettia sens. lat. and Fordia sens. lat., which on the basis of molecular and chemical evidence are excluded from Millettia and Fordia sens. strict.

    Relationships between the major groups of genera centred on Lonchocarpus, Derris, Millettia and Tephrosia remain obscure, and still reflect a geographical bias in segregating them, i.e. distributions are limited largely to the New World in the Lonchocarpus group, and the Old World in the other groups. The suggestion that the Andean South American genus Apurimacia might be sister to the largely Old World Tephrosia rather than to Lonchocarpus (e.g., Kajita et al., 2001) is possibly indicative of other Old World–New World sister groups yet to be found. Further molecular evidence will probably result in an overall reduction in the number of genera recognised, particularly in the Tephrosia and Lonchocarpus groups where various small or monotypic ‘one-organ’ genera may be better placed within larger genera. Ptycholobium, Requienia and Paratephrosia, for example, are difficult to distinguish from Tephrosia, but for the emphasis traditionally placed on their atypical pods.

    Further evidence may place this as a lineage within Tephrosia
    Habit
    Shrubs or herbs
    Ecology
    Seasonally dry tropical woodland, wooded grassland and shrubland, on sand
    Distribution
    NE Africa and Arabia (Somalia-Masai region); southern Africa (S Zambezian and Kalahari-Highveld regions)

    Images

    Distribution

    Native to:

    Angola, Botswana, Cape Provinces, Chad, Djibouti, Ethiopia, Mali, Mozambique, Namibia, Northern Provinces, Swaziland, Yemen, Zimbabwe

    Ptycholobium Harms appears in other Kew resources:

    First published in H.G.A.Engler & O.Drude, Veg. Erde 9(III 1): 59 (1915)

    Literature

    Flora Zambesiaca
    • Brummitt in Kew Bull. 35: 460–464 (1980).
    • —Brummitt in Kew Bull. 35: 460–464 (1980).
    • in Engler, Pflanzenw. Afrikas 3(1): 591 (1915).

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2018. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2018. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Legumes of the World Online
    http://creativecommons.org/licenses/by-nc-sa/3.0