1. Family: Scrophulariaceae Juss.
    1. Buddleja Houst. ex L.

      1. This genus is accepted, and its native range is Tropical & Subtropical America, NE. to S. Africa, Madagascar, Asia.

    [FZ]

    Loganiaceae, A. J. M. Leeuwenberg. Flora Zambesiaca 7:1. 1983

    Calyx
    Calyx green, campanulate or nearly so, less often cup–shaped or obconical; lobes usually subequal, from slightly longer to much shorter than the tube, erect, entire.
    Corolla
    Corolla cup–shaped, campanulate, funnel– or salver–shaped, variously coloured, from white to orange or to dark violet, or purple, often with an orange throat, sometimes darker at anthesis, outside with an indumentum of stellate and/or glandular hairs, but glabrous at the base, which is included in the calyx tube, and at the apices of the lobes, less often entirely glabrous or hairy on the entire outer side of the lobes; lobes usually shorter than the tube, imbricate in African species, elliptic or suborbicular, at the apex rounded, obtuse, or emarginate, entire to crenate, spreading or erect.
    Stamens
    Stamens inserted in the corolla, from the base of the tube to the mouth, with a certain degree of variation in a single species; filaments mostly short or very short, long only if stamens well–exserted (section Chilianthus); anthers oblong or less often subcircular in outline, mostly glabrous, apiculate to emarginate at the apex, mostly deeply cordate at the base, introrse; cells parallel or sometimes slightly divergent at the base, discrete, longitudinally dehiscent throughout.
    Pistil
    Pistil: ovary superior, usually 2–celled. Style short or long, included or exserted; stigma often large, clavate, capitate, or less often bilobate.
    Placenta
    In each cell one axile placenta with several to many ovules.
    Fruits
    Capsule bivalved, septicidal; valves mostly splitting up to the middle. Fruit a capsule or sometimes a berry, subglobose to narrowly ellipsoid, surrounded or subtended by the persistent calyx and often at the same time by the persistent corolla.
    Habit
    Shrubs, less often trees or suffrutescent herbs, 0·25–30 m. high, mostly with a white or pale grey indumentum of stellate hairs which dries usually rusty on the branchlets, the lower side of the leaves, and the inflorescences.
    Colleters
    Colleters absent.
    Stem
    Branchlets terete to quadrangular, in the latter case sometimes narrowly winged. Bark rough, fibrous, longitudinally fissured.
    Leaves
    Leaves opposite and those of a pair equal, less often subopposite, or in a few species alternate, petiolate, sessile, or — if opposite — sometimes connate–perfoliate.
    Stipules
    Stipules leafy, reduced to a line, or absent.
    Leaf lamina
    Lamina circular to very narrowly elliptic, entire, crenate to dentate, or sometimes evenlobed.
    Inflorescences
    Inflorescence terminal and/or axillary, thyrsoid, paniculate, or variously reduced even to a single globose head.
    Flowers
    Flowers 4–merous, actinomorphic.
    Seeds
    Seeds pale to dark brown, obliquely fusiform or obliquely polyhedral, winged or not, laterally compressed or not, reticulate or less often smooth or ridged.
    [FTEA]

    Loganiaceae, E. A. Bruce and J. Lewis. Flora of Tropical East Africa. 1960

    Habit
    Shrubs or small trees, hairy and glandular
    Leaves
    Leaves rather thick, often wrinkled, frequently dentate or crenate; stipules interpetiolar or reduced to a line
    Inflorescences
    Cymes many-flowered, variously grouped into terminal and lateral panicles
    Calyx
    Calyx 4-lobed or 4-fid
    Corolla
    Corolla-tube ± cylindrical; lobes spreading, imbricate in bud
    Stamens
    Stamens 4, inserted in the mouth of the corolla, not exserted (except in B. dysophylla)
    Ovary
    Ovary 2-locular; ovules numerous in rows on axile placentas
    Fruits
    Capsule ± oblong, septicidally dehiscent and the valves also at least partially halving from above
    Seeds
    Seeds small and numerous, sometimes winged or tailed; endosperm fleshy; embryo straight.
    [FTEA]

    Loganiaceae, E. A. Bruce and J. Lewis. Flora of Tropical East Africa. 1960

    Habit
    Shrubs with angled branches and usually large interpetiolar stipules
    Leaves
    Leaves elliptic or narrowly elliptic, papery, often glandular
    Inflorescences
    Inflorescences of few-flowered whorls or clusters of usually simple single axillary racemes or spikes
    Flowers
    Flowers similar to those of Buddleia
    Fruits
    Fruit a “drupe”.

    Images

    Distribution

    Native to:

    Afghanistan, Angola, Argentina Northeast, Argentina Northeast, Argentina Northwest, Argentina Northwest, Argentina South, Arizona, Assam, Bangladesh, Belize, Bolivia, Borneo, Brazil Northeast, Brazil South, Brazil Southeast, Brazil West-Central, California, Cambodia, Cape Provinces, Chile Central, Chile North, Chile South, China North-Central, China South-Central, China Southeast, Colombia, Comoros, Costa Rica, Cuba, Dominican Republic, East Himalaya, Ecuador, El Salvador, Eritrea, Ethiopia, Free State, Guatemala, Haiti, Honduras, India, Iran, Jamaica, Japan, Jawa, Kenya, KwaZulu-Natal, Laos, Lesotho, Lesser Sunda Is., Madagascar, Malawi, Malaya, Mauritius, Mexico Central, Mexico Gulf, Mexico Northeast, Mexico Northwest, Mexico Southeast, Mexico Southwest, Mozambique, Myanmar, Nansei-shoto, Nepal, Nevada, New Guinea, New Mexico, Nicaragua, Northern Provinces, Panamá, Paraguay, Peru, Philippines, Rodrigues, Réunion, Saudi Arabia, Somalia, Sudan, Sulawesi, Swaziland, Taiwan, Tanzania, Texas, Thailand, Tibet, Uganda, Uruguay, Utah, Venezuela, Vietnam, West Himalaya, Windward Is., Yemen, Zambia, Zaïre, Zimbabwe

    Introduced into:

    Alabama, Ascension, Belgium, Bermuda, Botswana, Bulgaria, Burundi, Cameroon, Corse, Czechoslovakia, France, Galápagos, Germany, Great Britain, Greece, Hawaii, Illinois, Ireland, Italy, Juan Fernández Is., Kentucky, Masachusettes, Netherlands, New Caledonia, New Zealand North, New Zealand South, North Carolina, Oregon, Puerto Rico, Queensland, Rwanda, Spain, St.Helena, Switzerland, Tennessee, Transcaucasus, Trinidad-Tobago, Washington

    Buddleja Houst. ex L. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Rico, L. [1312], Bolivia K000295147

    First published in Sp. Pl.: 112 (1753)

    Accepted by

    • Govaerts, R. (1996). World Checklist of Seed Plants 2(1, 2): 1-492. MIM, Deurne.

    Literature

    Flora Zambesiaca
    • Leeuwenberg in Meded. Landb. Wag. 79–6: 1–163(1979).
    • Sp. Pl. 1: 112. (1753).
    Flora of Tropical East Africa
    • Marquand in K.B. 1930: 177 (1930)
    • L., Gen. Pl., ed. 5: 51 (1754)
    • Sp. Pl.: 112 (1753)

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2018. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2018. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0