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This genus is accepted, and its native range is Tropical & Subtropical coasts.
A specimen from Kew's Herbarium


Cymodoceaceae, Henk Beentje. Flora of Tropical East Africa. 2002

Morphology General Habit
Submerse aquatic herbs; rhizome creeping, monopodial, with 2 vascular bundles; at each node 1-several roots and a short erect stem bearing 1– 4 leaves
Morphology Leaves
Leaves with sheath 1– 6 cm long, semi-persistent, when shed leaving circular scar; blade linear, entire, narrowed at base, with 3 veins and conspicuous midrib which is widened or furcate at apex
Morphology Reproductive morphology Flowers
Flowers solitary and terminal, enclosed in a leaf similar to others; bud in axil of penultimate leaf develops into prolongation of main axis giving rise to a sympodium; staminate flower subsessile, of 2 anthers attached at different levels, joined basally; carpellate flowers subsessile, of 2 free ovaries, each with a long style
Morphology Reproductive morphology Fruits
Fruit subglobose-ovoid, shortly beaked, ± compressed, with stony pericarp.


M. Thulin et al. Flora of Somalia, Vol. 1-4 [updated 2008]

Vegetative Multiplication Rhizomes
Rhizome with 1–6 unbranched roots and distichously arranged foliage leaves at each node, producing lateral branches but without consistent differentiation between long and short shoots; rhizome scales 2, ovate or elliptic
Morphology Leaves
Leaf sheath 1–6 cm long, scarious, persisting longer than the blade, when shed leaving an annular scar on stem; mouth of the sheath with two lateral rounded auricles; ligule a narrow apical ridge; blade dark green, flat, often narrowed at the base, 3-veined, with prominent midrib, widened or forked at the apex, and inconspicuous lateral nerves; apex very variable in outline, tridentate or bicuspidate, rounded and serrulate or emarginate
Morphology Reproductive morphology Inflorescences
Flowers solitary, terminal on a short erect shoot
Morphology Reproductive morphology Flowers
Male flower stalked; anthers facing opposite directions, joined below, one slightly above the other Female flower subsessile, with obovoid or globose carpels, each with a long simple style
Morphology Reproductive morphology Fruits
Fruit with a stony pericarp, ovoid, somewhat compressed, with a short subapical beak.
Genus of six species, pantropical.

Native to:

Alabama, Aldabra, Andaman Is., Angola, Bahamas, Bangladesh, Belize, Bermuda, Borneo, Brazil Northeast, Brazil Southeast, Caroline Is., Cayman Is., Colombia, Costa Rica, Cuba, Dominican Republic, Egypt, Eritrea, Fiji, Florida, Gabon, Guatemala, Gulf of Guinea Is., Gulf States, Hainan, Haiti, Honduras, India, Iran, Jamaica, Jawa, Kenya, KwaZulu-Natal, Leeward Is., Lesser Sunda Is., Louisiana, Madagascar, Malaya, Maluku, Marianas, Mauritania, Mauritius, Mexico Gulf, Mexico Northeast, Mexico Northwest, Mexico Southeast, Mississippi, Mozambique, Mozambique Channel I, Myanmar, Nansei-shoto, Netherlands Antilles, New Caledonia, New Guinea, Nicaragua, Nicobar Is., North Carolina, Oman, Palestine, Panamá, Philippines, Puerto Rico, Queensland, Saudi Arabia, Senegal, Seychelles, Socotra, Somalia, South China Sea, Sri Lanka, Sudan, Sulawesi, Taiwan, Tanzania, Thailand, Tonga, Trinidad-Tobago, Turks-Caicos Is., Venezuela, Vietnam, Western Australia, Windward Is., Yemen

Halodule Endl. appears in other Kew resources:

First published in Gen. Pl., Suppl. 1: 1368 (1841)

Accepted by

  • Govaerts, R.H.A. (2011). World checklist of selected plant families published update Facilitated by the Trustees of the Royal Botanic Gardens, Kew.


Flora of West Tropical Africa

  • Gen. Pl. Suppl. 1: 1368 (1841)
  • den Hartog in Blumea 12: 289-312 (1964).

Flora of Somalia

  • Flora Somalia, Vol 4, (1995) Author: by G. Sartoni [updated by M. Thulin 2008]

Flora of Tropical East Africa

  • Asch. in Linnaea 35: 163, 187 (1868)
  • Gen. Pl. suppl. 1: 1368 (1841)
  • Hartog in Blumea 12: 289– 382 (1964)
  • Sea-grasses of the World: 146 (1970)

Flora of Somalia
Flora of Somalia

Flora of Tropical East Africa
Flora of Tropical East Africa

Kew Backbone Distributions
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at and
© Copyright 2017 World Checklist of Selected Plant Families.

Kew Names and Taxonomic Backbone
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at and
© Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families.