1. Family: Lentibulariaceae Rich.
    1. Utricularia L.

      1. This genus is accepted, and its native range is Cosmopolitan.

    [FTEA]

    Lentibulariaceae, P. Taylor. Flora of Tropical East Africa. 1973

    Habit
    Herbs, annual or perennial, terrestrial, epiphytic or aquatic
    Roots
    Root-like organs (rhizoids) usually descending from the base of the inflorescence, usually filiform
    Stem
    Stem-like organs (stolons) arising with the rhizoids at the inflorescence base; in the terrestrial and epiphytic species usually short and delicate but sometimes developing into fleshy tubers; in the aquatic species usually more robust and longer Vegetative parts not clearly differentiated but consisting of stems modified to function as roots, stems, leaves and specialized organs (traps) for the capture of small organisms
    Leaves
    Foliar organs (leaves) either rosulate at the inflorescence base or alternate, opposite or verticillate on the stolons; in the terrestrial and epiphytic species entire, erect or thalloid, capillary, linear, orbicular or peltate; in the aquatic species ± dichotomously divided into capillary segments
    Traps
    Traps hollow, globose or ovoid, usually stalked, with a mouth which may be basal (adjacent to the stalk), lateral or terminal (opposite to the stalk); mouth provided externally with 2 lips which bear very diverse appendages
    Inflorescences
    Inflorescence racemose, bracteate; scape usually simple, sometimes branched, usually filiform, erect or twining, usually glabrous, sometimes papillose, glandular or hairy, provided (especially in the terrestrial and epiphytic species) with sterile bracts (scales); raceme usually elongated, rarely short and subcapitate; pedicels usually short, terete, flattened or ± winged, often reflexed or recurved in fruit; bracts persistent, basifixed or produced below the point of insertion or peltate; bracteoles 2 or absent or sometimes ± fused with the bract, inserted with the bract at the base of the pedicel or rarely with the calyx-lobes at the apex of the pedicel
    Calyx
    Calyx-lobes 2, ± equal or sometimes very unequal, usually free, sometimes ± united at the base, persistent and usually accrescent, sometimes very markedly so; upper lobe usually entire, lower lobe usually emarginate or bidentate, rarely both lobes dentate or fimbriate
    Corolla
    Corolla bilabiate, glabrous, glandular or pubescent; throat closed or sometimes open; upper lip usually ± erect, limb entire, emarginate or bilobed; lower lip usually larger, spurred or rarely saccate at the base, palate usually raised and gibbous, limb spreading or deflexed, entire, emarginate or ± deeply 2–5-lobed; spur ± parallel to the lower lip or divergent at an acute or obtuse angle or rarely in the same plane
    Stamens
    Stamens 2, inserted at the base of the corolla; filaments straight or curved, usually twisted, sometimes winged; anthers dorsifixed, ± ellipsoid, the thecae ± confluent
    Pistil
    Ovary globose or ovoid, unilocular; style usually short, often indistinct, persistent; stigma bilabiate, lower lip usually much larger than the upper which may be obscure or obsolete; ovules 2-many, sessile on a ± fleshy basal or free central placenta, anatropous
    Fruits
    Fruit a capsule, ± globose, dehiscing by longitudinal slits or by pores or circumscissile or indehiscent
    Seeds
    Embryo undifferentiated. Seeds 1–many, usually small or very small, globose, ovoid, truncate conical, narrowly cylindrical, fusiform, lenticular or prismatic, smooth, verrucose, reticulate, glochidiate or variously winged
    [FZ]

    Lentibulariaceae, P. Taylor. Flora Zambesiaca 8:3. 1988

    Habit
    Vegetative parts not clearly differentiated but consisting of stems modified to function as roots, stems, leaves and specialized organs (traps) for the capture of small organisms. Herbs, annual or perennial, terrestrial, epiphytic or aquatic.
    Roots
    Root-like organs (rhizoids) usually descending from the base of the inflorescence, usually filiform.
    Stem
    Stem-like organs (stolons) arising with the rhizoids at the inflorescence base; in the terrestrial and epiphytic species usually short and delicate but sometimes developing into fleshy tubers; in the aquatic species usually more robust and longer.
    Leaves
    Foliar organs (leaves) either rosulate at the inflorescence base or alternate, opposite or verticillate on the stolons; in the terrestrial and epiphytic species mostly entire, erect or thalloid, capillary, linear, circular or peltate; in the aquatic species usually more or less dichotomously divided into capillary segments.
    Traps
    Traps rosulate, or lateral on the rhizoids, stolons or leaves or rarely terminal on the leaves, hollow, globose or ovoid, usually stalked, with a mouth which may be basal (adjacent to the stalk), lateral or terminal (opposite to the stalk); mouth usually provided externally with very diverse appendages.
    Inflorescences
    Inflorescence racemose, (but sometimes with a single flower) bracteate; peduncle usually simple, sometimes branched, usually filiform, erect or twining, usually glabrous, sometimes papillose, glandular or hairy, often provided (especially in the terrestrial species) with sterile bracts (scales); raceme usually elongated, rarely short and subcapitate; pedicels usually short, terete, flattened or more or less winged, often deflexed or decurved in fruit; bracts persistent, basifixed or produced below the point of insertion or peltate; bracteoles 2 or absent or sometimes more or less fused with the bract, inserted with the bract at the base of the pedicel or rarely with the calyx lobes at the apex of the pedicel.
    Calyx
    Calyx lobes 2, or rarely 4 in 2 decussate pairs, more or less equal or sometimes very unequal, usually free, sometimes more or less united at the base, persistent and usually accrescent, sometimes very markedly so; upper lobe usually entire, inferior lobe usually emarginate or bidentate, rarely both lobes dentate or fimbriate.
    Corolla
    Corolla bilabiate, glabrous, glandular or pubescent; throat closed or sometimes open; superior lip usually more or less erect, limb entire, emarginate or bilobed; inferior lip usually larger, spurred or rarely saccate at the base, palate usually raised and gibbous, limb spreading or deflexed, entire, emarginate or more or less deeply 2–5-lobed; spur more or less parallel to the inferior lip or divergent at an acute or obtuse angle or rarely in the same plane.
    Stamens
    Stamens 2, inserted at the base of the corolla; filaments straight or curved, usually twisted, sometimes winged; anthers dorsifixed, more or less ellipsoid, the thecae usually more or less confluent.
    Pistil
    Ovary globose or ovoid, unilocular; style usually short, often indistinct, persistent; stigma bilabiate, inferior lip usually much larger than the upper which may be obscure or obsolete; ovules 2-many, sessile on a more or less fleshy free basal placenta, anatropous.
    Fruits
    Fruit a capsule, more or less globose or ovoid, dehiscing by longitudinal slits or by pores or circumscissile or indehiscent.
    Seeds
    Seeds 1-many, usually small or very small, globose, ovoid, truncate conical, narrowly cylindrical, fusiform, lenticular or prismatic, smooth, verrucose, reticulate, glochidiate, papillose or comose or variously winged.

    Images

    Distribution

    Native to:

    Afghanistan, Alabama, Alaska, Albania, Alberta, Aleutian Is., Algeria, Altay, Amur, Andaman Is., Angola, Argentina Northeast, Argentina Northwest, Argentina South, Arizona, Arkansas, Assam, Austria, Bahamas, Baltic States, Bangladesh, Belarus, Belgium, Belize, Benin, Bolivia, Borneo, Botswana, Brazil North, Brazil Northeast, Brazil South, Brazil Southeast, Brazil West-Central, British Columbia, Bulgaria, Burkina, Burundi, Buryatiya, Cabinda, California, Cambodia, Cameroon, Cape Provinces, Caprivi Strip, Caroline Is., Central African Repu, Central European Rus, Chad, China North-Central, China South-Central, China Southeast, Chita, Colombia, Colorado, Comoros, Congo, Connecticut, Costa Rica, Cuba, Czechoslovakia, Delaware, Denmark, District of Columbia, Dominican Republic, East European Russia, East Himalaya, Ecuador, Egypt, El Salvador, Ethiopia, Finland, Florida, France, Free State, French Guiana, Føroyar, Gabon, Galápagos, Gambia, Georgia, Germany, Ghana, Great Britain, Greece, Greenland, Guatemala, Guinea, Guinea-Bissau, Gulf of Guinea Is., Guyana, Hainan, Haiti, Hawaii, Honduras, Hungary, Iceland, Idaho, Illinois, India, Indiana, Inner Mongolia, Iowa, Iran, Iraq, Ireland, Irkutsk, Italy, Ivory Coast, Jamaica, Japan, Jawa, Kamchatka, Kansas, Kazakhstan, Kentucky, Kenya, Khabarovsk, Kirgizstan, Korea, Krasnoyarsk, Kriti, Krym, Kuril Is., KwaZulu-Natal, Labrador, Laos, Lebanon-Syria, Leeward Is., Lesotho, Lesser Sunda Is., Liberia, Libya, Louisiana, Madagascar, Magadan, Maine, Malawi, Malaya, Mali, Maluku, Manchuria, Manitoba, Marianas, Maryland, Masachusettes, Mauritania, Mauritius, Mexico Central, Mexico Gulf, Mexico Northeast, Mexico Northwest, Mexico Southeast, Mexico Southwest, Michigan, Minnesota, Mississippi, Missouri, Mongolia, Montana, Morocco, Mozambique, Myanmar, Namibia, Nansei-shoto, Nebraska, Nepal, Netherlands, Nevada, New Brunswick, New Caledonia, New Guinea, New Hampshire, New Jersey, New Mexico, New South Wales, New York, New Zealand North, New Zealand South, Newfoundland, Nicaragua, Nicobar Is., Niger, Nigeria, North Carolina, North Dakota, North European Russi, Northern Provinces, Northern Territory, Northwest European R, Northwest Territorie, Norway, Nova Scotia, Nunavut, Ohio, Oklahoma, Oman, Ontario, Oregon, Pakistan, Palestine, Panamá, Paraguay, Pennsylvania, Peru, Philippines, Poland, Portugal, Primorye, Prince Edward I., Puerto Rico, Queensland, Québec, Rhode I., Romania, Rwanda, Sakhalin, Saskatchewan, Senegal, Sicilia, Sierra Leone, Socotra, Somalia, South Australia, South Carolina, South Dakota, South European Russi, Spain, Sri Lanka, Sudan, Sulawesi, Sumatera, Suriname, Swaziland, Sweden, Switzerland, Taiwan, Tanzania, Tasmania, Tennessee, Texas, Thailand, Tibet, Togo, Trinidad-Tobago, Tunisia, Turkey, Turkey-in-Europe, Tuva, Uganda, Ukraine, Uruguay, Utah, Venezuela, Vermont, Victoria, Vietnam, Virginia, Washington, West Himalaya, West Siberia, West Virginia, Western Australia, Windward Is., Wisconsin, Wyoming, Yakutskiya, Yemen, Yugoslavia, Yukon, Zambia, Zaïre, Zimbabwe

    Utricularia L. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Jan 1, 1998 Stannard, B. [50857] 74539.000
    Jan 1, 1997 Laessoe, T. [53333] 74540.000
    Venezuela 56423.000
    Taylor, P. [17025], Australia 40518.000
    Taylor, P. [17020], Australia 40519.000
    Taylor, P. [17031], Australia 40521.000
    Taylor, P. [17004], Australia 40523.000
    Taylor, P. [17018], Australia 43273.000
    Taylor, P. [17017], Australia 43275.000
    Taylor, P. [17002], Australia 43276.000
    Taylor, P. [17024], Australia 43284.000
    Taylor, P. [17026], Australia 43288.000
    Melville, R. [4476], Australia 25426.000
    Harley, R.M. [25211], Brazil 54539.000
    Harley, R.M. [24927], Brazil 54730.000
    Harley, R.M. [26557], Brazil 54732.000
    Harley, R.M. [H52068], Brazil 57905.000
    Harley, R.M. [H52064], Brazil 57981.000
    Harley, R.M. [H 52094], Brazil 58312.000
    Harley, R.M. [H50605], Brazil 58990.000
    Harley, R.M. [27643], Brazil 60213.000
    Harley, R.M. [24768], French Guiana 63228.000
    Harley, R.M. [27636], Brazil 63233.000
    Harley, R.M. [27636], Brazil 63239.000
    Sands, M.J.S. [5665], Brunei 59618.000
    Sands, M.J.S. [5664], Brunei 62696.000
    Sands, M.J.S. [5433], Brunei 62701.000
    George, A.S. [9707], Australia 18739.004
    George, A.S. [10388], Australia 18739.005
    George, A.S. [9512], Australia 3360.006
    Erickson, R., Australia 25881.000
    Erickson, R., Australia 30591.000
    Stannard, B. [H52826], Brazil 57904.000
    Stannard, B. [H52787], Brazil 57910.000
    Stannard, B. [H52794], Brazil 57912.000
    Stannard, B. [H52788], Brazil 57915.000
    Stannard, B. [H51699], Brazil 57917.000
    Stannard, B. [H52790], Brazil 57929.000
    Stannard, B. [H51700], Brazil 57959.000
    Stannard, B. [H50857], Brazil 57966.000
    Stannard, B. [H50857], Brazil 57976.000
    Stannard, B. [H52806], Brazil 57977.000
    Edwards, P.J. [4137], New Guinea 70465.000
    Johns, R.J. [9415], New Guinea 62484.000
    Johns, R.J. [9633], New Guinea 62513.000
    Johns, R.J. [9663], New Guinea 62526.000
    Johns, R.J. [9633], New Guinea 62527.000
    Johns, R.J. [10424], New Guinea 70636.000
    Johns, R.J. [10693], Indonesia 72686.000
    Johns, R.J. [10676], Indonesia 73282.000
    Baxell, D.F. [W75/173], Australia 37221.000
    Weston, A. [17000], Australia 40524.000
    Clarkson, J.R. [9939], Australia 74532.000
    Hind, D.J.N. [H50491], Brazil 57911.000
    Hind, D.J.N. [H50491], Brazil 57914.000
    Hind, D.J.N. [H50491], Brazil 57918.000
    Hind, D.J.N. [H50063], Brazil 57945.000
    Hind, D.J.N. [H50491], Brazil 57958.000
    Hind, D.J.N. [H50491], Brazil 57980.000
    Nic Lughadha, E. [H53336], Brazil 57913.000
    Nic Lughadha, E. [H51117], Brazil 57916.000
    Nic Lughadha, E. [H51117], Brazil 58890.000
    Laessoe, T. [H52526], Brazil 57947.000
    Laessoe, T. [H52545], Brazil 57960.000
    Laessoe, T. [H53333], Brazil 57969.000
    Laessoe, T. [H53334], Brazil 57970.000
    Laessoe, T. [H53322], Brazil 57974.000
    Laessoe, T. [H50896], Brazil 58310.000
    Sano, P.T. [H50983], Brazil 58991.000
    Thiv, Colombia 58010.000
    Willis, F.R. [91], Indonesia 71464.000
    Willis, F.R. [161], Indonesia 72302.000
    s.coll. [Cat. no. s.n.], Myanmar K001132502

    First published in Sp. Pl.: 18 (1753)

    Accepted by

    • Taylor, P. (1989). The Genus Utricularia: a taxonomic monograph: 1-724. Her Majesty's stationery office, London.
    • Tutin, T.G. & al. (eds.) (1972). Flora Europaea 3: 1-370. Cambridge University Press.

    Literature

    Flora of West Tropical Africa
    • P. Taylor in Kew Bull. 18 (1) (1963).
    • Pellegr. in Bull. Soc. Bot. Fr. 61: 13—21 (1914)
    • —F.T.A. 4, 2: 469
    Flora Zambesiaca
    • in F.T.E.A., Lentibulariaceae: 1 (1973).
    • in Fl. Afr. Centr., Lentibulariaceae: 2 (1972)
    • P. Taylor in Kew Bull. 18: 1 (1964)
    • Sp. Pl: 18 (1753) & Gen. Pl., ed. 5: 11 (1754).
    Flora of Tropical East Africa
    • in Fl. Afr. Centr., Lentib.: 2 (1972)
    • P. Taylor in K.B. 18: 1 (1964)
    • Stapf in F.T.A. 4 (2): 469 (1906)
    • L., Gen. Pl., ed. 5: 11 (1754)
    • Sp. Pl.: 18 (1753)

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Interactive Key to Seed Plants of Malesia and Indo-China
    The Malesian Key Group (2010) Interactive Key to Seed Plants of Malesia and Indo-China (Version 2.0, 28 Jul 2010) The Nationaal Herbarium Nederland Leiden and The Royal Botanic Gardens Kew

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Neotropikey
    Milliken, W., Klitgard, B. and Baracat, A. (2009 onwards), Neotropikey - Interactive key and information resources for flowering plants of the Neotropics.
    http://creativecommons.org/licenses/by/3.0

    Plants and People Africa
    Common Names from Plants and People Africa http://www.plantsandpeopleafrica.com/
    © Plants and People Africa http://www.plantsandpeopleafrica.com http://creativecommons.org/licenses/by/3.0/