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  1. Family: Rhamnaceae Juss.
    1. Gouania Jacq.

      1. This genus is accepted, and its native range is Tropics.


    Rhamnaceae, R. B. Drummond. Flora Zambesiaca 2:2. 1966

    Climbing shrubs or lianes; branches provided with coiled tendrils.
    Leaves alternate, petiolate; lamina entire or dentate, penninerved but often 3–5-nerved from the base.
    Stipules small, caducous.
    Inflorescence a paniculate thyrse.
    Flowers usually bisexual.
    Sepals 5.
    Petals 5, cucullate; inserted below the margin of the disk.
    Stamens 5.
    Disk filling the receptacle, 5-lobed.
    Ovary inferior, immersed in the disk; style 3-fid or obscurely 3-lobed.
    Fruit a schizocarp, longitudinally 3-winged (or 3-angled), septicidal, separating into 3 woody or coriaceous indehiscent cocci.
    Seeds compressed or planoconvex.

    Rhamnaceae, Marshall C. Johnston (University of Texas Herbarium). Flora of Tropical East Africa. 1972

    Climbing shrubs or lianes with circinnate tendrils (rarely, or apparently never in Africa, seeming to be mere shrubs)
    Leaves alternate, petiolate; blades usually ovate, serrate (rarely nearly entire), acute or acuminate, at base usually rounded or cordate and often 3–5-nerved but penninerved more distally
    Stipules present, free, usually caducous
    Cymes usually small, but usually aggregated into ample leafless terminal panicles
    Flowers 5-merous, bisexual, epigynous at all stages
    Disk rather massive, epigynous
    Ovary usually urceolate; style trifid or obscurely 3-lobed
    Fruit a dry schizocarpous capsule, late in maturation the endo- and exocarpous tissue near the septa (on both sides of each mericarp) producing 3 broad thin wings (thus each wing is composed of parts of 2 adjacent mericarps; reportedly the wing-production fails in certain Asian and American, but not African, populations, and perhaps the reports are based on observations of immature fruit, for the wing-proliferation is a quite late development).



    Native to:

    Aldabra, Andaman Is., Angola, Argentina Northeast, Argentina Northwest, Assam, Bahamas, Bangladesh, Belize, Benin, Bismarck Archipelago, Bolivia, Borneo, Brazil North, Brazil Northeast, Brazil South, Brazil Southeast, Brazil West-Central, Burundi, Cambodia, Cameroon, Central African Repu, China South-Central, China Southeast, Colombia, Comoros, Congo, Costa Rica, Cuba, Dominican Republic, East Himalaya, Ecuador, El Salvador, Equatorial Guinea, Ethiopia, Fiji, Florida, French Guiana, Gabon, Galápagos, Ghana, Guatemala, Guinea, Gulf of Guinea Is., Guyana, Hainan, Haiti, Hawaii, Honduras, India, Ivory Coast, Jamaica, Kenya, Laos, Leeward Is., Liberia, Madagascar, Malawi, Malaya, Mauritius, Mexico Central, Mexico Gulf, Mexico Northeast, Mexico Northwest, Mexico Southeast, Mexico Southwest, Mozambique, Myanmar, Nepal, New Caledonia, New Guinea, Nicaragua, Nicobar Is., Nigeria, Panamá, Paraguay, Peru, Philippines, Puerto Rico, Queensland, Rodrigues, Rwanda, Réunion, Sierra Leone, Sri Lanka, Sudan, Sulawesi, Sumatera, Suriname, Tanzania, Thailand, Togo, Trinidad-Tobago, Tuamotu, Uganda, Uruguay, Vanuatu, Venezuela, Vietnam, West Himalaya, Windward Is., Zaïre, Zimbabwe

    Accepted Species


    Other Data

    Gouania Jacq. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Jul 1, 2002 Cheek, M. [9360], Cameroon K000027198
    DuPuy, B. [MB 608], Madagascar 63559.000
    Cavalerie, J. [3457], Yunnan K000732120


    First published in Select. Stirp. Amer. Hist.: 263 (1763)

    Accepted by

    • Buerki, S., Phillipson, P.B. & Callmander, M.W. (2011). A taxonomic revision of Gouania (Rhamnaceae) in Madagascar and the other islands of the western Indian Ocean (the Comoro and Mascarene islands and Seychelles) Annals from the Missouri Botanical Garden 98: 157-195. MBG press.
    • Govaerts, R. (2003). World Checklist of Seed Plants Database in ACCESS G: 1-40325.


    Flora of West Tropical Africa

    • Suessenguth in E. & P. Pflanzenfam. 20D: 166 (1953).
    • —F.T.A. 1: 383

    Flora Zambesiaca

    • Select. Stirp. Amer. Hist.: 263 (1763).

    Flora of Tropical East Africa

    • Select. Stirp. Amer. Hist.: 263 (1763)


    Flora Zambesiaca
    Flora Zambesiaca

    Flora of Tropical East Africa
    Flora of Tropical East Africa

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2020. Published on the Internet at and
    © Copyright 2017 World Checklist of Selected Plant Families.

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2020. Published on the Internet at and
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families.