1. Family: Phyllanthaceae Martinov
    1. Leptopus Decne.

      1. This genus is accepted, and its native range is Caucasus to China and Malesia.

    [KBu]

    Vorontsova, M.S. & Hoffmann, P. 2009. Revision of the genus Leptopus (Phyllanthaceae, Euphorbiaceae sensu lato). Kew Bulletin 64: 627. DOI: https://doi.org/10.1007/s12225-009-9155-x

    Type
    Leptopus cordifolius Decne.
    Note
    Synonym: Thelypetalum Gagnep. (1924 publ. 1925: 876). Type: Thelypetalum pierrei Gagnep. Synonym: Hexakistra Hook. f. (1887a: 283), nom. invalid., in synon. Synonym: Arachne Neck. (1790: 348) nom. invalid., opus oppressum (Art. 32.9, App. VI, McNeill et al.2006).
    Habit
    Monoecious (rarely dioecious) Herbs to shrubs, procumbent, ascendant or erect
    Indumentum
    Indumentum simple or 0
    Petiole
    Petioles adaxially grooved
    Stipules
    Stipules persistent, membranaceous, eglandular, margin entire to lobed, glabrous or ciliate
    Leaves
    Leaves alternate, petiolate, entire, penninerved, 1.5 – 5 times longer than wide, membranaceous to coriaceous, eglandular
    Inflorescences
    Inflorescences axillary, solitary or fasciculate, male flowers sometimes on short densely bracteate inflorescences
    Pedicel
    Female pedicels apically dilated, not articulate Male pedicels usually filiform, articulate above base; articulation practically invisible before abscission
    Calyx
    Sepals 5 (6), imbricate, entire, chartaceous, not accrescent to slightly accrescent in fruit. Petals 5 (6), membranaceous, shorter than or as long as the sepals in male flowers, minute and often hidden under disc lobes in female flowers
    Disc
    Disc extrastaminal, chartaceous becoming membranaceous with age, glabrous; disc in male flowers composed of 5 (6) contiguous regular segments bilobed for ⅓ - ⅘ of their length, lobes rounded, truncate or irregular; disc in female flowers annular, regularly divided into 5 emarginate segments, lobes entire to erose
    Stamens
    Stamens 5, oppositisepalous; filaments free, filiform; anthers latrorse in bud, ellipsoid, longitudinally dehiscent
    Pollen
    Pollen prolate to subprolate; exine tectate perforate to microreticulate
    Pistillodes
    Pistillode composed of 3 free segments or 3-lobed
    Ovary
    Ovary 3 (4)-locular (4- or 5-locular in L. fangdingianus), globose, glabrous to pilose
    Ovules
    Ovules 2 per locule, anatropous, straight, adnate to the funicle for ⅘ of the ovule length; obturator translucent covering apices of both ovules
    Style
    Styles free (sometimes fused for up to ½ length in L. pachyphyllus), erect, bifid to base or nearly so, recurved
    Stigma
    Stigmas apically dilated to capitate, smooth
    Fruits
    Fruit 3- (4- or 5)-lobate to subglobose, smooth, sometimes with faint reticulate venation. Columella persistent, 5 – 6 times longer than its narrowest width
    Seeds
    Seeds ecarunculate (fleshy micropylar seed appendages in L. emicans), rounded triquetrous to almost reniform, smooth, rugose or pitted, dull; micropyle minute
    Endosperm
    Endosperm copious, entire to deeply invaginate in cross section
    Embryo
    Embryo longitudinally curved to almost straight
    Cotyledons
    Cotyledons equal, broad, thin, as long as the radicle or longer, much wider than radicle Cotyledons positioned parallel to the pericarp

    Images

    Distribution

    Native to:

    Afghanistan, Andaman Is., Assam, Borneo, China North-Central, China South-Central, China Southeast, East Himalaya, Hainan, India, Inner Mongolia, Iran, Jawa, Lesser Sunda Is., Malaya, Maluku, Manchuria, Myanmar, Nepal, North Caucasus, Pakistan, Philippines, Qinghai, Thailand, Tibet, Transcaucasus, Vietnam, West Himalaya

    Leptopus Decne. appears in other Kew resources:

    First published in V.Jacquemont, Voy. Inde 4: 155 (1844)

    Accepted by

    • Vorontsova, M.S. & Hoffmann, P. (2010). Revision of the genus Leptopus (Phyllanthaceae, Euphorbiaceae sensu lato) Kew Bulletin 64: 627-644.

    Not accepted by

    • Govaerts, R., Frodin, D.G. & Radcliffe-Smith, A. (2000). World Checklist and Bibliography of Euphorbiaceae (and Pandaceae) 1-4: 1-1622. The Board of Trustees of the Royal Botanic Gardens, Kew. [Cited as Andrachne.]

    Sources

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Bulletin
    Kew Bulletin
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0