1. Family: Poaceae Barnhart
    1. Genus: Cenchrus L.
      1. Cenchrus echinatus L.

        This species is accepted, and its native range is Central & S. U.S.A. to Tropical & Subtropical America.

    [CPLC]

    Bernal, R., Gradstein, S.R. & Celis, M. (eds.). 2015. Catálogo de plantas y líquenes de Colombia. Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá. http://catalogoplantasdecolombia.unal.edu.co

    Distribution
    Nativa en Colombia; Alt. 0 - 1500 m.; Andes, Guayana y Serranía de La Macarena, Islas Caribeñas, Llanura del Caribe, Orinoquia, Sierra Nevada de Santa Marta, Valle del Cauca, Valle del Magdalena.
    Habit
    Hierba
    Conservation
    No Evaluada
    [FTEA]

    Gramineae, W. D. Clayton, S. M. Phillips & S. A. Renvoize. Flora of Tropical East Africa. 1974

    Habit
    Coarse annual; culms 15–90 cm. high, ascending.
    Leaves
    Leaf-blades 4–25 cm. long, 3–10 mm. wide.
    Inflorescences
    Panicle 2–10 cm. long; involucres globose, 5–10 mm. long; inner spines flattened, connate for ± 1/2 their length to form a deep cup, pubescent, retrorsely barbellate; outer spines rigid, mostly shorter than the inner, ± divergent.
    Spikelets
    Spikelets 2–3 per burr, 5–7 mm. long.
    Habitat
    Roadside and waste places; 0–900 m.
    Distribution
    now established as a weed throughout the tropics and subtropics K7 T3 T6
    [FWTA]

    Gramineae, W. D. Clayton. Flora of West Tropical Africa 3:2. 1972

    Habit
    Annual 30–90 cm. high.
    [GB]
    Habit
    Annual. Culms geniculately ascending; 15-90 cm long. Ligule a fringe of hairs. Leaf-blades 4-25 cm long; 3-10 mm wide.
    Inflorescences
    Inflorescence a panicle. Panicle spiciform; linear; interrupted (or lax); 2-10 cm long. Primary panicle branches accrescent to a central axis; with sessile scars on axis. Panicle axis angular; bearing deciduous spikelet clusters. Spikelets subtended by an involucre. Fertile spikelets sessile; 2-3 in the cluster. Involucre composed of bristles; connate into a cup below; with 2-5 mm connate; globose; 5-10 mm long; base obconical. Involucral bristles deciduous with the fertile spikelets; with an outer whorl of thinner bristles; inner bristles longer than outer; with longest bristle scarcely emergent; 2-5 mm long; flattened; rigid; retrorsely scaberulous; pubescent; spinose.
    Spikelets
    Spikelets comprising 1 basal sterile florets; 1 fertile florets; without rhachilla extension. Spikelets ovate; dorsally compressed; acuminate; 5-7 mm long; falling entire; deciduous with accessory branch structures.
    Fertile
    Spikelets comprising 1 basal sterile florets; 1 fertile florets; without rhachilla extension. Spikelets ovate; dorsally compressed; acuminate; 5-7 mm long; falling entire; deciduous with accessory branch structures.
    Glume
    Glumes shorter than spikelet; thinner than fertile lemma. Lower glume ovate; 1.3-3.4 mm long; 0.33-0.5 length of spikelet; membranous; without keels; 1 -veined. Lower glume lateral veins absent. Lower glume apex obtuse, or acute. Upper glume ovate; 3.8-5.7 mm long; 0.75 length of spikelet; membranous; without keels; 3-5 -veined. Upper glume surface puberulous. Upper glume apex obtuse, or acute.
    Florets
    Basal sterile florets barren; with palea. Lemma of lower sterile floret ovate; 1 length of spikelet; membranous; 5 -veined; obtuse, or acute. Fertile lemma ovate; 5-7 mm long; coriaceous; much thinner on margins; without keel; 5 -veined. Lemma margins flat. Lemma apex obtuse. Palea coriaceous.
    Fruits
    Caryopsis with adherent pericarp; ellipsoid; dorsally compressed; 3 mm long; truncate.
    Distribution
    Africa: north, Macaronesia, west tropical, west-central tropical, east tropical, and western Indian ocean. Asia-temperate: western Asia, Arabia, China, and eastern Asia. Asia-tropical: India, Indo-China, Malesia, and Papuasia. Australasia: Australia. Pacific: southwestern, south-central, northwestern, and north-central. North America: southwest USA, south-central USA, southeast USA, and Mexico. South America: Mesoamericana, Caribbean, northern South America, western South America, Brazil, and southern South America.
    Reference
    Paniceae. FTEA.
    Diagnostic
    Involucre base 2.2-3.6mm wide.

    Images

    Distribution

    Native to:

    Alabama, Argentina Northeast, Argentina Northwest, Arizona, Arkansas, Aruba, Bahamas, Belize, Bermuda, Bolivia, Brazil North, Brazil Northeast, Brazil South, Brazil Southeast, Brazil West-Central, California, Cayman Is., Central American Pac, Chile North, Chile South, Colombia, Costa Rica, Cuba, Desventurados Is., District of Columbia, Dominican Republic, Ecuador, El Salvador, Florida, French Guiana, Galápagos, Georgia, Guatemala, Guyana, Haiti, Honduras, Jamaica, Kansas, Kentucky, Leeward Is., Louisiana, Maryland, Mexico Central, Mexico Gulf, Mexico Northeast, Mexico Northwest, Mexico Southeast, Mexico Southwest, Mississippi, Netherlands Antilles, Nevada, New Mexico, Nicaragua, North Carolina, Oklahoma, Panamá, Paraguay, Peru, Puerto Rico, South Carolina, Southwest Caribbean, Suriname, Tennessee, Texas, Trinidad-Tobago, Turks-Caicos Is., Uruguay, Venezuela, Venezuelan Antilles, Virginia, Windward Is.

    Introduced into:

    Aldabra, Bangladesh, Borneo, Canary Is., Cape Verde, Caroline Is., Chagos Archipelago, China South-Central, China Southeast, Cocos (Keeling) Is., Cook Is., Czechoslovakia, Easter Is., Egypt, Equatorial Guinea, Fiji, Gabon, Ghana, Gilbert Is., Greece, Gulf States, Hainan, Hawaii, India, Ivory Coast, Japan, Kenya, Lebanon-Syria, Lesser Sunda Is., Line Is., Madagascar, Maine, Malaya, Marianas, Marquesas, Marshall Is., Masachusettes, Mauritius, Mexican Pacific Is., Mozambique Channel I, Nansei-shoto, Nauru, New Caledonia, New Guinea, Nigeria, Niue, Northern Territory, Oman, Palestine, Philippines, Phoenix Is., Rodrigues, Réunion, Samoa, Saudi Arabia, Seychelles, Society Is., Solomon Is., South China Sea, Sri Lanka, Taiwan, Tanzania, Togo, Tokelau-Manihiki, Tonga, Tuamotu, Tubuai Is., Tuvalu, Vanuatu, Vermont, Wake I., Wallis-Futuna Is.

    Cenchrus echinatus L. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Jan 1, 1933 Curtis, A.H. [3619], Florida K001056122
    Hinton, G.B. [4800], México State K000476298
    Hinton, G.B. [12066], Michoacán K000476299
    Hinton, G.B. [12066], Michoacán K000476300
    Hinton, G.B. [1160], México State K000476301
    Philcox, D. [3172], Brazil K001075832
    Spruce, R. [5818], Ecuador K000201726
    Spruce, R. [5818], Ecuador K000201727
    Spruce, R. [5818], Ecuador K000201728
    Irwin, H.S. [5653], Brazil K001075830
    Mueller, F. [473], Brazil K001075829
    Wight, R. [1665], India K000245391 Unknown type material
    Bourgeau, M. [3140], Mexico K000643125
    Hitchcock, A.S. [13801], Hawaii K001056124 Unknown type material
    Angus [168], Brazil K001075826
    Sellow [s.n.], Brazil K001075825
    Laegaard [21791], Thailand K000688373
    Glaziou, A.F.M. [6954], Brazil K001075827
    Harger, E.B. [426], Connecticut K001056123 isolectotype
    s.coll. [Cat. no. 8654] K001131124
    s.coll. [Cat. no. 8654] K001131125
    s.coll. [Cat. no. 8654] K001131126
    s.coll. [Cat. no. 8654], India K001131127
    Dombrowski, L.T.D. [2419], Brazil K001075828
    Lechler, P. [1545], Peru K001044597
    Spruce, R. [s.n.], Brazil K001075831

    First published in Sp. Pl.: 1050 (1753)

    Accepted by

    • Sosef, M.S.M. & al. (2019). Novitates Gabonenses 91: additons to the grass flora of Gabon Plant Ecology and Evolution 152: 99-105.
    • Bosser, J. & Renvoize, S.A. (2018). Flore des Mascareignes 203: 1-276. IRD Éditions, MSIRI, RBG-Kew, Paris.
    • Sykes, W.R. (2016). Flora of the Cook Islands: 1-973. National Tropical Botanical Garden, Hawaii.
    • Veldkamp, J.F. (2014). A revision of Cenchrus incl. Pennisetum (Gramineae) in Malesia with some general nomenclatural notes Blumea 59: 59-75.
    • Patzelt, A., Harrison, T., Knees, S.G. & Hartley, L.A. (2014). Studies in the flora of Arabia: XXXI. New records from the Sultanate of Oman Edinburgh Journal of Botany 71: 161-180.
    • Zuloaga, F.O., Morrone, O. & Pensiero, J.F. (2014). Gramineae VI Flora del Paraguay 45: 1-392. Editions des Conservatoire et Jardin botaniques de la ville de Genève.
    • Verloove, F. (2013). New xenophytes from Gran Canaria (Canary islands, Spain), with emphasis on naturalised and (potentially) invasive species Collectanea Botanica 32: 59-82.
    • Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies Smithsonian Contributions to Botany 98: 1-1192.
    • Giraldo-Cañas, D. (2011). Catálogo de la familia Poaceae en Colombia Darwiniana 49: 139-247.
    • Reddy, G.V.P. (2011). Survey of invasive plants on Guam and identification of the 20 most widespread Micronesica; Journal of the College of Guam 41: 263-274.
    • Dobignard, D. & Chatelain, C. (2010). Index synonymique de la flore d'Afrique du nord 1: 1-455. Éditions des conservatoire et jardin botaniques, Genève.
    • Clayton, W.D. & Snow, N. (2010). A key to Pacific Grasses: 1-107. Kew Publishing, Royal Botanic Gardens, Kew.
    • Ahmed, Z.U. (ed.) (2008). Encyclopedia of Flora and Fauna of Bangladesh 12: 1-505. Asiatic Society of Bangladesh.
    • Zuloaga, F.O., Morrone, O. , Belgrano, M.J., Marticorena, C. & Marchesi, E. (eds.) (2008). Catálogo de las Plantas Vasculares del Cono Sur Monographs in Systematic Botany from the Missouri Botanical Garden 107: 1-3348.
    • Hokche, O., Berry, P.E. & Huber, O. (eds.) (2008). Nuevo Catálogo de la Flora Vascular de Venezuela: 1-859. Fundación Instituto Botánico de Venezuela.
    • Nelson Sutherland, C.H. (2008). Catálogo de las plantes vasculares de Honduras. Espermatofitas: 1-1576. SERNA/Guaymuras, Tegucigalpa, Honduras.
    • Cope, T.A., Knees, S.G. & Miller, A.G. (2007). Flora of the Arabian peninsula and Socotra 5(1): 1-387. Edinburgh University Press.
    • Wu, Z. & Raven, P.H. (eds.) (2006). Poaceae Flora of China 22: 1-733. Missouri Botanical Garden Press, St. Louis.
    • Clayton, W.D., Harman, K.T. & Williamson, H. (2006). World Grass Species - Synonymy database The Board of Trustees of the Royal Botanic Gardens, Kew.
    • Dávila, P., Mejia-Saulés, M.T., Gómez-Sánchez, N., Valdés-Reyna, J., Ortíz, J.J., Morín, C., Castrejón, J. & Ocampo, A. (2006). Catálogo de las Gramíneas de México: 1-671. CONABIO, México city.
    • Boulos, L. (2005). Flora of Egypt 4: 1-617. Al Hadara Publishing, Cairo.
    • Danin, A. (2004). Distribution Atlas of Plants in the Flora Palaestina area: 1-517. The Israel Academy of Sciences and Humanities, Jerusalem.
    • Flora of North America Editorial Committee (2003). Flora of North America North of Mexico 25: 1-781. Oxford University Press, New York, Oxford.
    • Aké Assi, L. (2002). Flore de la Côte-d'Ivoire: catalogue systématique, biogéographie et écologie. II Boissiera 58: 1-401.
    • Jørgensen, P.M. & León-Yánez, S. (eds.) (1999). Catalogue of the Vascular Plants of Ecuador Monographs in Systematic Botany from the Missouri Botanical Garden 75: i-viii, 1-1181.
    • Aedo, C., Tellería, M.T. & Velayos, M. (eds.) (1999). Bases Documentales para la Flora de Guinea Ecuatorial; Plantas vascularis y hongos: 1-414. CSIC, real jardín Botánico, Madrid.
    • Renvoize, S.A. (1998). Gramineas de Bolivia: 1-644. The Royal Botanic Gardens, Kew.
    • Boggan, J. Funck, V. & Kelloff, C. (1997). Checklist of the Plants of the Guianas (Guyana, Surinam, Franch Guiana) ed. 2: 1-238. University of Guyana, Georgetown.
    • Cirilo, N. & Proctor, G.R. (1994). Vascular plants of the Caribbean Swan islands of Honduras Brenesia 41-42: 73-80.
    • Davidse, G. & al. (eds.) (1994). Flora Mesoamericana 6: 1-543. Universidad Nacional Autónoma de México, México, D.F.
    • Dassanayake (ed.) (1994). A Revised Handbook to the Flora of Ceylon 8: 1-458. Oxford & IBH Publishing Co. PVT. LTD., New Delhi, Calcutta.
    • George, A.S., Orchard, A.E. & Hewson, H.J. (eds.) (1993). Oceanic islands 2 Flora of Australia 50: 1-606. Australian Government Publishing Service, Canberra.
    • Brako, L. & Zarucchi, J.L. (1993). Catalogue of the Flowering Plants and Gymnosperms of Peru Monographs in Systematic Botany from the Missouri Botanical Garden 45: i-xl, 1-1286.
    • Karthikeyan, S., Jain, S.K., Nayar, M.P. & Sanjappa, M. (1989). Florae Indicae Enumeratio: Monocotyledonae: 1-435. Botanical Survey of India, Calcutta.
    • Levin, G.A. & Moran, R. (1989). The vascular flora of isla Socorro, Mexico: 1-71. Dept. of Botany, San Diego Natural History Museum, San Diego.
    • Robertson, S.A. (1989). Flowering Plants of Seychelles: 1-327. Royal Botanic Gardens, Kew.
    • Koyama, T. (1987). Grasses of Japan and its neighboring regions: an identification manual: 1-570. Kodansha, Tokyo, Japan.
    • Woodroffe, C.D. (1985). Vegetation and flora of Nui Atoll, Tuvalu Atoll Research Bulletin 283: 1-18.
    • Brunel, J.F., Hiepo, P. & Scholz, H. (eds.) (1984). Flore Analytique du Togo Phanérogames: 1-751. GTZ, Eschborn.
    • Clayton, W.D. & Renvoize, S.A. (1982). Gramineae Flora of Tropical East Africa 3: 451-898.
    • Fosberg, F.R. & Renvoize, S.A. (1980). The Flora of Aldabra and neighbouring islands: 1-358. Crown, London.
    • Walker, E.H. (1976). Flora of Okinawa and the southern Ryukyu islands: 1-1159. Smithsonian Institution Press, Washington D.C., U.S.A.
    • Hepper, F.N. (ed.) (1972). Flora of West Tropical Africa, ed. 2, 3(2): 277-574.
    • Parham, B.E.V. (1971). The Vegetation of the Tokelau Islands with special reference to the plants of Nukunonu Atoll New Zealand Journal of Botany 9: 576-609.
    • Bosser, J. (1969). Gramiées des pasturages et des cultures a Madagascar: 1-440. ORSTOM, Paris.
    • Henty, E.E. (1969). A manual of the grasses of New Guinea: 1-215. Division of Botany, Dpt. of Forests, LAE, New Guinea.

    Literature

    Flora of West Tropical Africa
    • F.T.A. 9: 1082.
    • Sp. Pl. 1050 (1753)
    Kew Backbone Distributions
    • Sosef, M.S.M. & al. (2019). Novitates Gabonenses 91: additons to the grass flora of Gabon Plant Ecology and Evolution 152: 99-105.
    • Veldkamp, J.F. (2014). A revision of Cenchrus incl. Pennisetum (Gramineae) in Malesia with some general nomenclatural notes Blumea 59: 59-75.
    • Patzelt, A., Harrison, T., Knees, S.G. & Hartley, L.A. (2014). Studies in the flora of Arabia: XXXI. New records from the Sultanate of Oman Edinburgh Journal of Botany 71: 161-180.
    • Verloove, F. (2013). New xenophytes from Gran Canaria (Canary islands, Spain), with emphasis on naturalised and (potentially) invasive species Collectanea Botanica 32: 59-82.
    • Forzza, R.C. & al. (2013). Lista de Espécies da Flora do Brasil http://floradobrasil.jbrj.gov.br/2010/.
    • Danihelka, J. Chrtek, J. & Kaplan, Z. (2012). Checklist of vascular plants of the Czech Republic Preslia. Casopsi Ceské Botanické Spolecnosti 84: 647-811.
    • Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies Smithsonian Contributions to Botany 98: 1-1192.
    • Greuter, W. & Raus, T. (eds.) (2012). Med-Checklist Notulae, 31 Willdenowia 42: 287-295.
    • Giraldo-Cañas, D. (2011). Catálogo de la familia Poaceae en Colombia Darwiniana 49: 139-247.
    • Reddy, G.V.P. (2011). Survey of invasive plants on Guam and identification of the 20 most widespread Micronesica; Journal of the College of Guam 41: 263-274.
    • Dobignard, D. & Chatelain, C. (2010). Index synonymique de la flore d'Afrique du nord 1: 1-455. Éditions des conservatoire et jardin botaniques, Genève.
    • Clayton, W.D. & Snow, N. (2010). A key to Pacific Grasses: 1-107. Kew Publishing, Royal Botanic Gardens, Kew.
    • Ahmed, Z.U. (ed.) (2008). Encyclopedia of Flora and Fauna of Bangladesh 12: 1-505. Asiatic Society of Bangladesh.
    • Zuloaga, F.O., Morrone, O. , Belgrano, M.J., Marticorena, C. & Marchesi, E. (eds.) (2008). Catálogo de las Plantas Vasculares del Cono Sur Monographs in Systematic Botany from the Missouri Botanical Garden 107: 1-3348.
    • Cope, T.A., Knees, S.G. & Miller, A.G. (2007). Flora of the Arabian peninsula and Socotra 5(1): 1-387. Edinburgh University Press.
    • Wu, Z. & Raven, P.H. (eds.) (2006). Poaceae Flora of China 22: 1-733. Missouri Botanical Garden Press, St. Louis.
    • Dávila, P., Mejia-Saulés, M.T., Gómez-Sánchez, N., Valdés-Reyna, J., Ortíz, J.J., Morín, C., Castrejón, J. & Ocampo, A. (2006). Catálogo de las Gramíneas de México: 1-671. CONABIO, México city.
    • Danin, A. (2004). Distribution Atlas of Plants in the Flora Palaestina area: 1-517. The Israel Academy of Sciences and Humanities, Jerusalem.
    • Flora of North America Editorial Committee (2003). Flora of North America North of Mexico 25: 1-781. Oxford University Press, New York, Oxford.
    • Aké Assi, L. (2002). Flore de la Côte-d'Ivoire: catalogue systématique, biogéographie et écologie. II Boissiera 58: 1-401.
    • Jørgensen, P.M. & León-Yánez, S. (eds.) (1999). Catalogue of the Vascular Plants of Ecuador Monographs in Systematic Botany from the Missouri Botanical Garden 75: i-viii, 1-1181.
    • Aedo, C., Tellería, M.T. & Velayos, M. (eds.) (1999). Bases Documentales para la Flora de Guinea Ecuatorial; Plantas vascularis y hongos: 1-414. CSIC, real jardín Botánico, Madrid.
    • Sheppard, C.R.C. & Seaward, M.R.D. (eds.) (1999). Ecology of the Chagos archipelago: 1-350. Westbury Academic & Scientific Publishing, Otley.
    • Renvoize, S.A. (1998). Gramineas de Bolivia: 1-644. The Royal Botanic Gardens, Kew.
    • Boggan, J. Funck, V. & Kelloff, C. (1997). Checklist of the Plants of the Guianas (Guyana, Surinam, Franch Guiana) ed. 2: 1-238. University of Guyana, Georgetown.
    • Cirilo, N. & Proctor, G.R. (1994). Vascular plants of the Caribbean Swan islands of Honduras Brenesia 41-42: 73-80.
    • Davidse, G. & al. (eds.) (1994). Flora Mesoamericana 6: 1-543. Universidad Nacional Autónoma de México, México, D.F.
    • Dassanayake (ed.) (1994). A Revised Handbook to the Flora of Ceylon 8: 1-458. Oxford & IBH Publishing Co. PVT. LTD., New Delhi, Calcutta.
    • George, A.S., Orchard, A.E. & Hewson, H.J. (eds.) (1993). Oceanic islands 2 Flora of Australia 50: 1-606. Australian Government Publishing Service, Canberra.
    • Brako, L. & Zarucchi, J.L. (1993). Catalogue of the Flowering Plants and Gymnosperms of Peru Monographs in Systematic Botany from the Missouri Botanical Garden 45: i-xl, 1-1286.
    • Levin, G.A. & Moran, R. (1989). The vascular flora of isla Socorro, Mexico: 1-71. Dept. of Botany, San Diego Natural History Museum, San Diego.
    • Robertson, S.A. (1989). Flowering Plants of Seychelles: 1-327. Royal Botanic Gardens, Kew.
    • Koyama, T. (1987). Grasses of Japan and its neighboring regions: an identification manual: 1-570. Kodansha, Tokyo, Japan.
    • Woodroffe, C.D. (1985). Vegetation and flora of Nui Atoll, Tuvalu Atoll Research Bulletin 283: 1-18.
    • Morat, P. & Veillon, J.-M. (1985). Contributions à la conaissance de la végétation et de la flore de Wallis et Futuna Bulletin du Muséum National d'Histoire Naturelle. Section B, Adansonia 7: 259-329.
    • Brunel, J.F., Hiepo, P. & Scholz, H. (eds.) (1984). Flore Analytique du Togo Phanérogames: 1-751. GTZ, Eschborn.
    • Clayton, W.D. & Renvoize, S.A. (1982). Gramineae Flora of Tropical East Africa 3: 451-898.
    • Correll, D.S. & Correll, H.B. (1982). Flora of the Bahama Archipelago: 1-1692. J.Cramer, Vaduz.
    • Walker, E.H. (1976). Flora of Okinawa and the southern Ryukyu islands: 1-1159. Smithsonian Institution Press, Washington D.C., U.S.A.
    • Lebrun, J.P. (1973). Énumération des plantes vasculaires du Sénégal: 1-209. Maisons Alfort: Institut d'élevage et de médecine vétérinaire des pays tropicaux.
    • Hepper, F.N. (ed.) (1972). Flora of West Tropical Africa, ed. 2, 3(2): 277-574.
    • Parham, B.E.V. (1971). The Vegetation of the Tokelau Islands with special reference to the plants of Nukunonu Atoll New Zealand Journal of Botany 9: 576-609.
    • Bosser, J. (1969). Gramiées des pasturages et des cultures a Madagascar: 1-440. ORSTOM, Paris.
    • Henty, E.E. (1969). A manual of the grasses of New Guinea: 1-215. Division of Botany, Dpt. of Forests, LAE, New Guinea.
    • Masamune, G. (1942). Enumeratio phanerogamarum Bornearum: 1-739. Taihoku, Formosa.
    • Hubbard, C.E. & Vaughan, R.E. (1940). Grasses of Mauritius and Rodriguez: 1-128. Crown Agents for the Colonies, London.
    • Britton, N. (1918). Flora of Bermuda: 1-585. Charles Scribner's Sons, New York.
    • de Cordemoy, E.J. (1895). Flore de l'ile de la Réunion: 1-574. Librairie des sciences naturelles Paul Klincksieck, Paris.
    Flora of Tropical East Africa
    • Clayton in Flora of West Tropical Africa, ed. 2, 3: 464 (1972).
    • Stapf & C. E. Hubbard in Flora of Tropical Africa 9: 1082 (1934).
    • L., Sp. Pl.: 1050 (1753).

    Sources

    Catálogo de Plantas y Líquenes de Colombia
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of West Tropical Africa
    Flora of West Tropical Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    GrassBase - The Online World Grass Flora
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Science Photographs
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/