1. Family: Ebenaceae Gürke
    1. Genus: Diospyros L.
      1. Diospyros verrucosa Hiern

        This species is accepted, and its native range is Tanzania to Mozambique.

    [FTEA]

    Ebenaceae, F. White & B. Verdcourt. Flora of Tropical East Africa. 1996

    Habit
    Shrub or small tree (1-)2-18 m. tall; bark dark grey or black, closely and deeply fissured longitudinally; young branchlets, tomentellous or densely ferruginous pubescent.
    Leaves
    Leaves drying blackish brown above, distinctly paler, usually whitish beneath, ovate or ovate-oblong to lanceolate, 2.5-12.5 cm. long, 1.2-5.5(-7) cm. wide, subacute to shortly acuminate at the apex, rounded at the base, subcoriaceous, sometimes obscurely crenulate, ± shiny and slightly pubescent above particularly on the midrib, soon glabrous, lower surface adpressed sericeous-pilose, somewhat glabrescent except on nerves and epidermis glaucous due to dense minute papilla-like hairs; lateral nerves 5-7 pairs, prominent beneath but tertiary nerves and veins usually indistinct; petiole 3-7 mm. long.
    Flowers
    Male flowers in 3-7-flowered cymules, borne towards the base of the current year’ s growth in the axils of deciduous reduced leaves or the first-formed foliage leaves; peduncle 1-2 mm. long; pedicels 1-2 mm. long; calyx 2.5 mm. long; calyx-lobes 4, triangular, 1.5 mm. long; corolla white or yellowish white, ellipsoid or ovoid-ellipsoid in bud, 6 mm. long; corolla-lobes 4, 1 mm. long; stamens ± 12, inserted on the receptacle, included; anthers linear-lanceolate, apiculate, strigose; rudimentary ovary minute, tomentose. Female flowers usually solitary, rarely in 2-3-flowered cymules, in the axils of leaves or of bracts below the leaves; peduncle together with pedicel 0.3-1.2 cm. long; calyx 3-4 mm. long; calyx-lobes 4, broadly triangular, 2 mm. long; corolla urceolate, 7 mm. long, 3 mm. wide; corolla-lobes 4, under 1 mm. long; ovary 4 mm. long, 2.4 mm. wide; locules 4, uniovulate; style scarcely differentiated from the ovary, under 1 mm. long, bifid at the apex, the branches ending in 2 fleshy irregularly lobed stigmas.
    Male
    Male flowers in 3-7-flowered cymules, borne towards the base of the current year’ s growth in the axils of deciduous reduced leaves or the first-formed foliage leaves; peduncle 1-2 mm. long; pedicels 1-2 mm. long; calyx 2.5 mm. long; calyx-lobes 4, triangular, 1.5 mm. long; corolla white or yellowish white, ellipsoid or ovoid-ellipsoid in bud, 6 mm. long; corolla-lobes 4, 1 mm. long; stamens ± 12, inserted on the receptacle, included; anthers linear-lanceolate, apiculate, strigose; rudimentary ovary minute, tomentose.
    Female
    Female flowers usually solitary, rarely in 2-3-flowered cymules, in the axils of leaves or of bracts below the leaves; peduncle together with pedicel 0.3-1.2 cm. long; calyx 3-4 mm. long; calyx-lobes 4, broadly triangular, 2 mm. long; corolla urceolate, 7 mm. long, 3 mm. wide; corolla-lobes 4, under 1 mm. long; ovary 4 mm. long, 2.4 mm. wide; locules 4, uniovulate; style scarcely differentiated from the ovary, under 1 mm. long, bifid at the apex, the branches ending in 2 fleshy irregularly lobed stigmas.
    Fruits
    Fruit cream at first, orange then blackish, subglobose or broadly ellipsoid, 3-4.5 cm. long, 2-4 cm. wide, verrucose, densely rusty hairy at first, glabrescent, the calyx sometimes with reflected lobes.
    Seeds
    Seeds 4 or fewer, blackish, oblong-trigonous, 1.8-2 cm. long, 0.8-1.2 cm. wide, 5 mm. thick; endosperm ruminate.
    Figures
    Fig. 3/24.
    Habitat
    Evergreen forest, coastal scrub and grassland; 100-600 m.
    Distribution
    T6 T8
    [FZ]

    Ebenaceae, F. White. Flora Zambesiaca 7:1. 1983

    Habit
    Evergreen shrub or small tree usually less than 5 m. high, rarely up to 15 m.
    Bark
    Bark blackish with close longitudinal fissures.
    Branches
    Young branchlets tomentellous.
    Leaves
    Leaves subcoriaceous, drying blackish–brown above, paler beneath; lamina up to 12·5 x 5·5 cm., ovate or ovate–oblong to lanceolate, apex subacute to shortly acuminate, base usually rounded; lower surface appressed sericeous–tomentose, somewhat glabres–cent except on nerves; lower epidermis glaucous, papillose; secondary nerves in 5–7 pairs, prominent beneath, tertiary nerves and veins usually indistinct.
    Stamens
    Stamens c. 12, included, inserted on receptacle; anthers linear–lanceolate, apiculate, strigose.
    Pistillodes
    Pistillode minute, tomentose.
    Male
    Pistillode minute, tomentose. Corolla 0·6 cm. long, ellipsoid or ovoid–ellipsoid in bud, lobes 4, c. 0·1 cm. long. Stamens c. 12, included, inserted on receptacle; anthers linear–lanceolate, apiculate, strigose. Male flowers in 3–7–flowered cymules, borne towards the base of the current year’s growth in the axils of caducous reduced leaves or the first formed foliage leaves; peduncle 0·1–0·2 cm. long; pedicels 0·1–0·2 cm. long. Calyx c. 0·25 cm. long, lobes 4, 0·15 cm. long, deltate.
    Flowers
    Female flowers usually solitary, rarely in 2–3–flowered cymules, in axils of leaves or of bracts below the leaves; peduncle (including pedicel) 0·3–1·2 cm. long. Male flowers in 3–7–flowered cymules, borne towards the base of the current year’s growth in the axils of caducous reduced leaves or the first formed foliage leaves; peduncle 0·1–0·2 cm. long; pedicels 0·1–0·2 cm. long.
    Calyx
    Calyx 0·3–0·4 cm. long, lobes 4, c. 0·2 cm. long, broadly debate. Calyx c. 0·25 cm. long, lobes 4, 0·15 cm. long, deltate.
    Corolla
    Corolla urceolate, 0·7 x 0·3 cm., lobes 4, less than 0–1 cm. long. Corolla 0·6 cm. long, ellipsoid or ovoid–ellipsoid in bud, lobes 4, c. 0·1 cm. long.
    Ovary
    Ovary 0·4 x 0·25 cm.; locules 4, uniovulate; style scarcely differentiated from ovary, less than 0·1 cm. long, bifid at apex, branches ending in 2 fleshy, irregularly lobed stigmata.
    Female
    Ovary 0·4 x 0·25 cm.; locules 4, uniovulate; style scarcely differentiated from ovary, less than 0·1 cm. long, bifid at apex, branches ending in 2 fleshy, irregularly lobed stigmata. Calyx 0·3–0·4 cm. long, lobes 4, c. 0·2 cm. long, broadly debate. Corolla urceolate, 0·7 x 0·3 cm., lobes 4, less than 0–1 cm. long. Female flowers usually solitary, rarely in 2–3–flowered cymules, in axils of leaves or of bracts below the leaves; peduncle (including pedicel) 0·3–1·2 cm. long.
    Fruits
    Fruit up to 4·5 x 4 cm., orange, subglobose or broadly ellipsoid, verrucose, especially when young, glabrescent. Fruiting calyx sometimes with reflexed lobes.
    Seeds
    Seeds 4 or fewer, blackish, endosperm ruminate.
    [FTEA]

    Ebenaceae, F. White & B. Verdcourt. Flora of Tropical East Africa. 1996

    Habit
    Shrub 2 m. tall, with young branchlets fulvous-tomentellous.
    Leaves
    Leaves lanceolate, 12-14 cm. long, 4-5 cm. wide, long acuminate at the apex, acute at the base, glabrous and very shining above save for midrib, fulvous pubescent beneath, not very coriaceous; petiole 5-8 mm. long.
    Flowers
    Male flowers solitary in the axils of the upper leaves, known only as buds; buds 1.6-1.8 cm. long; calyx finely felty, campanulate, 4-lobed, 4-5 mm. long; calyx-lobes triangular, 2.5 mm. long, acute; corolla 4-lobed, covered with short yellow hairs; stamens 12; filaments 0.5 mm. long; anthers 2.5 mm. long, with a tuft of hair at the base.
    Male
    Male flowers solitary in the axils of the upper leaves, known only as buds; buds 1.6-1.8 cm. long; calyx finely felty, campanulate, 4-lobed, 4-5 mm. long; calyx-lobes triangular, 2.5 mm. long, acute; corolla 4-lobed, covered with short yellow hairs; stamens 12; filaments 0.5 mm. long; anthers 2.5 mm. long, with a tuft of hair at the base.
    Habitat
    Dry woodland; ± 250 m.

    Images

    Distribution

    Native to:

    Mozambique, Tanzania

    Diospyros verrucosa Hiern appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Jan 1, 1987 Tanzania K000350761 syntype
    Jan 1, 1987 Kirk, J. [s.n.], Tanzania K000350760 lectotype

    First published in Trans. Cambridge Philos. Soc. 12: 167 (1873)

    Accepted by

    • Govaerts, R. (2000). World Checklist of Seed Plants Database in ACCESS D: 1-30141.

    Literature

    Flora of Tropical East Africa
    • F. White in Bulletin du Jardin Botanique National de Belgique 58: 406 (1988).
    • F. White in Flora Zambesiaca 7(1): 284, t. 65A (1983)
    • J.P.M. Brenan, Check-lists of the Forest Trees and Shrubs of the British Empire no. 5, part II, Tanganyika Territory p. 186 (1949).
    • Gürke in Die Pflanzenwelt Ost-Afrikas und der Nachbargebiete, Theile C: 306 (1895).
    • Hiern in Flora of Tropical Africa 3: 519 (1877).
    • Hiern in Trans. Camb. Phil. Soc. 12: 167 (1873)

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0