- Family: Lamiaceae Martinov
Stachys L.
[LKGF]
Harley, R.M. et al. (2004). Labiatae, in The Families and Genera of Vascular Plants (K. Kubitzki, ed. in chief) VI: 167-275. Springer-Verlag, Berlin, Heidelberg, New York.
- Morphology General Habit
- Perennial or annual herbs or rarely small subshrubs, with simple and/or branched hairs, often strongly aromatic
- Morphology Leaves
- Leaves usually toothed, petiolate or rarely sessile
- Morphology Reproductive morphology Inflorescences
- Inflorescence thyrsoid or racemoid
- Morphology Reproductive morphology Flowers
- Flowers pedicellate, rarely sessile
- Morphology Reproductive morphology Inflorescences Bracts
- Bracts present or absent
- Morphology Reproductive morphology Inflorescences Bracteoles
- Bracteoles present or absent
- Morphology Reproductive morphology Flowers Calyx
- Calyx actinomorphic to weakly 2-lipped, 5-lobed, lobes equal or unequal (3/2), often spinose, tube with throat often barbate
- Morphology Reproductive morphology Flowers Corolla
- Corolla strongly 2-lipped, 4-lobed (1/3), posterior lip long, hooded or sometimes almost flat, externally pubescent, anterior lip with median lobe usually much larger than lateral lobes, corolla-tube annulate or rarely exannulate
- Morphology Reproductive morphology Flowers Androecium Stamens
- Stamens not exserted from corolla, thecae ± distinct, usually divaricate, rarely subparallel to parallel
- Morphology Reproductive morphology Flowers Gynoecium Stigma
- Stigma-lobes subequal or slightly unequal
- Morphology Reproductive morphology Fruits
- Nutlets obovoid to oblong, rarely flattened trigonous, rounded or sometimes winged at apex.
- Note
- 2n =10, 16, 18, 24, 30, 32, 34, 48, 64, 66, 68, 80, 102. About 300 species A cladistic analysis by Lindquist and Albert (2002) suggests that Betonica and other Stachys may have different origins.
- Ecology
- Various habitats
- Distribution
- Cosmopolitan, except for Australia (introduced to New Zealand, Middle Atlantic Ocean, Southwestern and North-Central Pacific. Doubtfully present in Indo-China
[FTEA]
Lamiaceae (Labiatae), A.J. Paton, G. Bramley, O. Ryding, R.M. Polhill, Y.B. Harvey, M. Iwarsson, F. Willis, P.B. Phillipson, K. Balkwill, C.W. Lukhoba, D.F. Otieno, & R.M. Harley. Stachys, Y.B. Harvey. Flora of Tropical East Africa. 2009
- Morphology General Habit
- Perennial scandent or erect herbs, seldom aromatic, with an indumentum of simple, biramous, dendroid or stellate hairs
- Morphology Stem
- Stems quadrangular or roundedquadrangular, sometimes armed with prickles
- Morphology Leaves
- Leaves opposite, distichous or decussate, petiolate or sessile
- Morphology Reproductive morphology Inflorescences
- Inflorescences lax, with verticils variously 2–12(–16)flowered; bracts similar to cauline leaves, but smaller; bracteoles linear or subulate
- Morphology Reproductive morphology Flowers Calyx
- Calyx regularly 5-toothed, campanulate to funnel-shaped, 10-veined, not or enlarging only slightly in fruit
- Morphology Reproductive morphology Flowers Corolla
- Corolla white, pink, purplish or blue, 2-lipped; tube straight with annulus at midpoint; posterior lip flattened or slightly hooded; anterior lip with two small rounded lateral lobes and a large median lobe
- Morphology Reproductive morphology Flowers Androecium Stamens
- Stamens held under posterior lip adnating to just above mid-point of the tube; filaments sometimes pubescent; anthers 2-thecous, divaricate
- Morphology Reproductive morphology Flowers Disc
- Disk equally 4-lobed
- Morphology Reproductive morphology Flowers Gynoecium Ovary
- Ovary deeply 4-lobed; style gynobasic, bifid, branches equal
- Morphology Reproductive morphology Fruits
- Nutlets ovoid, minutely tuberculate and rounded at apex, with a small basal scar.
[FSOM]
M. Thulin et al. Flora of Somalia, Vol. 1-4 [updated 2008] https://plants.jstor.org/collection/FLOS
- Morphology General Habit
- Herbs or shrubs with branched hairs (in Somali species)
- Morphology Reproductive morphology Inflorescences
- Inflorescences thyrse-like, lax or dense; bracts large and leaf-like; cymes 1–10-flowered; bracteoles present (in Somali species)
- Morphology Reproductive morphology Flowers Calyx
- Calyx 5-lobed, with 5–10 ± distinct ribs; lobes subequal or 3 upper slightly different from the 2 lower
- Morphology Reproductive morphology Flowers Corolla
- Corolla 2-lipped; upper lip straight, slightly hooded, entire or emarginate, hairy but not bearded along the margin; lower lip 3-lobed with the midlobe larger, deflexed, markedly longer than the upper lip (in African species)
- Morphology Reproductive morphology Flowers Androecium Stamens
- Stamens directed to the upper lip of the corolla
- Morphology Reproductive morphology Flowers Gynoecium Style
- Style-branches equally long
- Morphology Reproductive morphology Fruits
- Nutlets rounded at apex, glabrous.
- Distribution
- Cosmopolitan genus of about 370 species.
>
[LKGF]
- Use
- Several species are widely cultivated as ornamentals. Stachys affinis Bunge is cultivated for its edible, subterranean tubers
Doubtfully present in:
Senegal
Native to:
Afghanistan, Alabama, Alaska, Albania, Alberta, Algeria, Altay, Amur, Angola, Argentina Northeast, Argentina Northwest, Argentina South, Arizona, Arkansas, Assam, Austria, Azores, Baleares, Baltic States, Bangladesh, Belarus, Belgium, Bolivia, Botswana, Brazil South, Brazil Southeast, British Columbia, Bulgaria, Burundi, Buryatiya, California, Cameroon, Canary Is., Cape Provinces, Cape Verde, Central European Rus, Chile Central, Chile North, Chile South, China North-Central, China South-Central, China Southeast, Chita, Colombia, Colorado, Connecticut, Corse, Costa Rica, Cyprus, Czechoslovakia, Denmark, District of Columbia, East Aegean Is., East European Russia, East Himalaya, Ecuador, Egypt, El Salvador, Eritrea, Ethiopia, Finland, Florida, France, Free State, Georgia, Germany, Great Britain, Greece, Guatemala, Gulf of Guinea Is., Honduras, Hungary, Illinois, Indiana, Inner Mongolia, Iowa, Iran, Iraq, Ireland, Irkutsk, Italy, Japan, Jawa, Kamchatka, Kansas, Kazakhstan, Kentucky, Kenya, Khabarovsk, Kirgizstan, Korea, Krasnoyarsk, Kriti, Krym, Kuril Is., KwaZulu-Natal, Lebanon-Syria, Lesotho, Libya, Louisiana, Madagascar, Madeira, Magadan, Maine, Malawi, Manchuria, Manitoba, Massachusetts, Mexico Central, Mexico Gulf, Mexico Northeast, Mexico Northwest, Mexico Southeast, Mexico Southwest, Michigan, Minnesota, Missouri, Mongolia, Montana, Morocco, Mozambique, Myanmar, Namibia, Nebraska, Nepal, Netherlands, Nevada, New Hampshire, New Jersey, New Mexico, New York, Nicaragua, Nigeria, North Carolina, North Caucasus, North Dakota, North European Russi, Northern Provinces, Northwest European R, Northwest Territorie, Norway, Ohio, Oklahoma, Ontario, Oregon, Pakistan, Palestine, Panamá, Paraguay, Pennsylvania, Peru, Poland, Portugal, Primorye, Qinghai, Québec, Rhode I., Romania, Rwanda, Sakhalin, Sardegna, Saskatchewan, Saudi Arabia, Sicilia, Sinai, Somalia, South Carolina, South Dakota, South European Russi, Spain, Sudan, Swaziland, Sweden, Switzerland, Tadzhikistan, Taiwan, Tanzania, Tennessee, Texas, Thailand, Tibet, Transcaucasus, Tunisia, Turkey, Turkey-in-Europe, Turkmenistan, Tuva, Uganda, Ukraine, Uruguay, Utah, Uzbekistan, Venezuela, Vermont, Vietnam, Virginia, Washington, West Himalaya, West Siberia, West Virginia, Wisconsin, Wyoming, Xinjiang, Yakutskiya, Yemen, Yugoslavia, Yukon, Zambia, Zaïre, Zimbabwe
Introduced into:
Arizona, Ascension, Bermuda, Easter Is., Galápagos, Hawaii, Juan Fernández Is., Leeward Is., Mali, Maryland, Mississippi, New Brunswick, New Caledonia, New Zealand North, New Zealand South, Newfoundland, Norfolk Is., Nova Scotia, Puerto Rico, St.Helena, Tasmania, Tristan da Cunha, Windward Is.
- Stachys acerosa Boiss.
- Stachys aculeolata Hook.f.
- Stachys adulterina Hemsl.
- Stachys aegyptiaca Pers.
- Stachys aethiopica L.
- Stachys affinis Bunge
- Stachys agraria Schltdl. & Cham.
- Stachys ajugoides Benth.
- Stachys alabamica B.R.Keener & L.J.Davenp.
- Stachys albanica Markgr.
- Stachys albens A.Gray
- Stachys albicaulis Lindl.
- Stachys albiflora N.E.Br.
- Stachys albotomentosa Ramamoorthy
- Stachys aleurites Boiss. & Heldr.
- Stachys alpigena T.C.E.Fr.
- Stachys alpina L.
- Stachys amanica P.H.Davis
- Stachys × ambigua Sm.
- Stachys anamurensis Sümbül
- Stachys andongensis Hiern
- Stachys angustifolia M.Bieb.
- Stachys anisochila Vis. & Pancic
- Stachys annua (L.) L.
- Stachys antalyensis Ayasligil & P.H.Davis
- Stachys aperta Epling
- Stachys appalachiana D.B.Poind. & J.B.Nelson
- Stachys arabica Hornem.
- Stachys arachnoidea Codd
- Stachys araucana Phil.
- Stachys arenaria Vahl
- Stachys arenariiformis Rouy
- Stachys argillicola Sebsebe
- Stachys aristata Greenm.
- Stachys arrecta L.H.Bailey
- Stachys arriagana B.L.Turner
- Stachys arvensis (L.) L.
- Stachys aspera Michx.
- Stachys asperata Hedge
- Stachys atherocalyx K.Koch
- Stachys aucheri Benth.
- Stachys aurea Benth.
- Stachys babunensis Micevski
- Stachys bakeri Briq.
- Stachys balansae Boiss. & Kotschy
- Stachys balensis Sebsebe
- Stachys bayburtensis R.Bhattacharjee & Hub.-Mor.
- Stachys baytopiorum Kit Tan & Yildiz
- Stachys beckeana Dörfl. & Hayek
- Stachys benthamiana Boiss.
- Stachys bergii G.A.Mulligan & D.B.Munro
- Stachys biflora Hook. & Arn.
- Stachys bigelovii A.Gray
- Stachys bithynica Boiss.
- Stachys bizensis Schweinf. ex Baker
- Stachys × bodeana Bunge
- Stachys bogotensis Kunth
- Stachys bolusii Skan
- Stachys bombycina Boiss.
- Stachys boraginoides Schltdl. & Cham.
- Stachys brachiata Bojer ex Benth.
- Stachys brachyclada de Noé
- Stachys bridgesii Benth.
- Stachys bullata Benth.
- Stachys burchelliana Launert
- Stachys burgsdorffioides (Benth.) Boiss.
- Stachys × burrii P.H.Davis
- Stachys buttleri R.R.Mill
- Stachys byzantina K.Koch
- Stachys caffra E.Mey. ex Benth.
- Stachys calcicola Epling
- Stachys candida Bory & Chaub.
- Stachys canescens Bory & Chaub.
- Stachys carduchorum (R.Bhattacharjee) Rech.f.
- Stachys caroliniana J.B.Nelson & D.A.Rayner
- Stachys cataonica R.Bhattacharjee & Hub.-Mor.
- Stachys chamissonis Benth.
- Stachys chasmosericea Ayasligil & P.H.Davis
- Stachys chinensis Bunge ex Benth.
- Stachys choruhensis Kit Tan & Sorger
- Stachys chrysantha Boiss. & Heldr.
- Stachys circinata L'Hér.
- Stachys citrina Boiss. & Heldr. ex Benth.
- Stachys clingmanii Small
- Stachys coccinea Ortega
- Stachys collina Brandegee
- Stachys comosa Codd
- Stachys cordata Riddell
- Stachys cordifolia Prain
- Stachys corsica Pers.
- Stachys costaricensis Briq.
- Stachys cretica L.
- Stachys × cryptadenia Rech.f.
- Stachys cuneata Banks ex Benth.
- Stachys cydni Kotschy ex Gemici & Leblebici
- Stachys cymbalaria Briq.
- Stachys darcyana A.Pool
- Stachys debilis Kunth
- Stachys didymantha Brenan
- Stachys × digenea Legué
- Stachys dinteri Launert
- Stachys distans Benth.
- Stachys diversifolia Boiss.
- Stachys dregeana Benth.
- Stachys drummondii Benth.
- Stachys durandiana Coss.
- Stachys duriaei de Noé
- Stachys ehrenbergii Boiss.
- Stachys elliptica Kunth
- Stachys eplingii J.B.Nelson
- Stachys erectiuscula Gürke
- Stachys eremicola Epling
- Stachys eriantha Benth.
- Stachys euadenia P.H.Davis
- Stachys euboica Rech.f.
- Stachys fendleri Briq.
- Stachys filifolia Hedge
- Stachys flavescens Benth.
- Stachys flexuosa Skan
- Stachys floccosa Benth.
- Stachys floridana Shuttlew. ex Benth.
- Stachys fominii Sosn. ex Grossh.
- Stachys fontqueri Pau
- Stachys forsythii Hedge
- Stachys fragillima Bornm.
- Stachys fruticulosa M.Bieb.
- Stachys geobombycis C.Y.Wu
- Stachys germanica L.
- Stachys gilliesii Benth.
- Stachys glandulibracteata Y.B.Harv.
- Stachys glandulifera Post
- Stachys glandulosa Hutch. & E.A.Bruce
- Stachys glandulosissima Floden
- Stachys globosa Epling
- Stachys glutinosa L.
- Stachys gossweileri G.Taylor
- Stachys goulimyi Rech.f.
- Stachys graciliflora C.Presl
- Stachys graeca Boiss. & Heldr.
- Stachys grandidentata Lindl.
- Stachys grandifolia E.Mey.
- Stachys graveolens Nábelek
- Stachys guyoniana de Noé ex Batt.
- Stachys hakkariensis Akçiçek & Firat
- Stachys hamata Epling
- Stachys harkerae J.G.González
- Stachys harleyana A.Pool
- Stachys hebens Epling
- Stachys heraclea All.
- Stachys herrerae Epling
- Stachys herrerana Rzed. & Calderón
- Stachys hians Briq.
- Stachys hildebrandtii Vatke
- Stachys hintoniorum B.L.Turner
- Stachys hispida Pursh
- Stachys hissarica Regel
- Stachys huber-morathii R.Bhattacharjee
- Stachys huetii Boiss.
- Stachys huillensis Hiern
- Stachys humbertii Hedge
- Stachys humifusa Burch. ex Benth.
- Stachys hydrophila Boiss.
- Stachys hyssopifolia Michx.
- Stachys hyssopoides Burch. ex Benth.
- Stachys iberica M.Bieb.
- Stachys iltisii J.B.Nelson
- Stachys inanis Hausskn. & Bornm.
- Stachys inflata Benth.
- Stachys intermedia Aiton
- Stachys ionica Halácsy
- Stachys iraqensis R.Bhattacharjee
- Stachys iva Griseb.
- Stachys jaimehintonii B.L.Turner
- Stachys jijigaensis Sebsebe
- Stachys keerlii Benth.
- Stachys kermanshahensis Rech.f.
- Stachys ketenoglui Kaynak, Daskin & Yilmaz
- Stachys komarovii Knorring
- Stachys kotschyi Boiss. & Hohen.
- Stachys kouyangensis (Vaniot) Dunn
- Stachys kulalensis Sebsebe
- Stachys kuntzei Gürke
- Stachys kurdica Boiss. & Hohen.
- Stachys lamarckii Benth.
- Stachys lamioides Benth.
- Stachys langmaniae Rzed. & Calderón
- Stachys lanigera (Bornm.) Rech.f.
- Stachys latidens Small
- Stachys lavandulifolia Vahl
- Stachys laxa Boiss. & Buhse
- Stachys leucoglossa Griseb.
- Stachys × leucomalla Bornm. & Gauba
- Stachys libanotica Benth.
- Stachys lindenii Benth.
- Stachys linearis Burch. ex Benth.
- Stachys longiflora Boiss. & Balansa
- Stachys longispicata Boiss. & Kotschy
- Stachys lurestanica Jamzad
- Stachys lyallii Benth.
- Stachys macraei Benth.
- Stachys macrotricha Rech.f. & Goulimy
- Stachys manantlanensis B.L.Turner
- Stachys mandoniana Briq.
- Stachys marashica Ilçim, Çenet & Dadandi
- Stachys mardinensis (Post) R.R.Mill
- Stachys maritima Gouan
- Stachys marrubiifolia Viv.
- Stachys matthewsii G.P.Fleming, J.B.Nelson & J.F.Towns.
- Stachys × medebachensis Feld & O.Koenen
- Stachys megalodonta Hausskn. & Bornm. ex P.H.Davis
- Stachys melampyroides Hand.-Mazz.
- Stachys melissifolia Benth.
- Stachys menthifolia Vis.
- Stachys menthoides Kotschy & Boiss.
- Stachys mexicana Benth.
- Stachys mialhesii de Noé
- Stachys milanii Petrov ex Magnier
- Stachys minor (Boiss.) Akçiçek & Dirmenci
- Stachys × mirabilis Rouy
- Stachys mohinora B.L.Turner
- Stachys mollissima Willd.
- Stachys moorei B.L.Turner
- Stachys mouretii Batt. & Pit.
- Stachys mucronata Sieber ex Spreng.
- Stachys multicaulis Benth.
- Stachys munzurdagensis R.Bhattacharjee
- Stachys musili Velen.
- Stachys namazdaghensis Yild.
- Stachys natalensis Hochst.
- Stachys nelsonii B.R.Keener & L.J.Davenp.
- Stachys nemorivaga Briq.
- Stachys nepetifolia Desf.
- Stachys nephrophylla Rech.f.
- Stachys neurocalycina Boiss.
- Stachys nigricans Benth.
- Stachys nivea Labill.
- Stachys nubilorum Epling
- Stachys obliqua Waldst. & Kit.
- Stachys oblongifolia Wall. ex Benth.
- Stachys obscura Boiss. & Balansa
- Stachys obtusicrena Boiss.
- Stachys obtusifolia MacOwan
- Stachys ochroleuca Phil.
- Stachys ocymastrum (L.) Briq.
- Stachys oligantha Baker
- Stachys oreophila Hedge
- Stachys palaestina L.
- Stachys palustris L.
- Stachys paneiana Mouterde
- Stachys pannosa Phil.
- Stachys parolinii Vis.
- Stachys pauli Grossh.
- Stachys penanevada B.L.Turner
- Stachys peruviana Dombey ex Benth.
- Stachys petrokosmos Rech.f.
- Stachys philippiana Vatke
- Stachys pilifera Benth.
- Stachys pilosa Nutt.
- Stachys pilosissima M.Martens & Galeotti
- Stachys pinardii Boiss.
- Stachys pinetorum Boiss. & Balansa
- Stachys pittieri Briq.
- Stachys plumosa Griseb.
- Stachys pringlei Greenm.
- Stachys pseudohumifusa Sebsebe
- Stachys pseudonigricans Gürke
- Stachys pseudophlomis C.Y.Wu
- Stachys pseudopinardii R.Bhattacharjee & Hub.-Mor.
- Stachys pumila Banks & Sol.
- Stachys pusilla (Wedd.) Briq.
- Stachys pycnantha Benth.
- Stachys pyramidalis J.K.Morton
- Stachys radicans Epling
- Stachys ramosissima Montbret & Aucher ex Benth.
- Stachys recta L.
- Stachys rehmannii Skan
- Stachys reptans Hedge
- Stachys reticulata Codd
- Stachys riederi Cham.
- Stachys rigida Nutt. ex Benth.
- Stachys riparia A.Pool
- Stachys rivularis J.M.Wood & M.S.Evans
- Stachys rizeensis R.Bhattacharjee
- Stachys rosea (Desf.) Boiss.
- Stachys rothrockii A.Gray
- Stachys rotundifolia Moc. & Sessé ex Benth.
- Stachys rubella Hedge
- Stachys rudatisii Skan
- Stachys rugosa Aiton
- Stachys rupestris Montbret & Aucher ex Benth.
- Stachys salisii Jord. & Fourr.
- Stachys sanchezii Rzed. & A.García
- Stachys sandersii B.L.Turner
- Stachys saturejoides Montbret & Aucher ex Benth.
- Stachys saxicola Coss. & Balansa
- Stachys scaberula Vatke
- Stachys scabrida Skan
- Stachys schimperi Vatke
- Stachys serbica Pancic
- Stachys sericantha P.H.Davis
- Stachys sericea Cav.
- Stachys sericophylla Halácsy
- Stachys sessilifolia E.Mey.
- Stachys sessilis Gürke
- Stachys setifera C.A.Mey.
- Stachys simplex Schltr.
- Stachys sintenisii Gand.
- Stachys sivasica Kit Tan & Yildiz
- Stachys sparsipilosa R.Bhattacharjee & Hub.-Mor.
- Stachys spathulata Burch. ex Benth.
- Stachys spectabilis Choisy ex DC.
- Stachys speluncarum Contandr. & Quézel
- Stachys sphaerodonta Baker
- Stachys spinosa L.
- Stachys spinulosa Sm.
- Stachys spreitzenhoferi Heldr.
- Stachys sprucei Briq.
- Stachys spruneri Boiss.
- Stachys stebbinsii G.A.Mulligan & D.B.Munro
- Stachys stricta Greene
- Stachys strictiflora C.Y.Wu
- Stachys subaphylla Rech.f.
- Stachys sublobata Skan
- Stachys subnuda Montbret & Aucher ex Benth.
- Stachys swainsonii Benth.
- Stachys sylvatica L.
- Stachys taliensis C.Y.Wu
- Stachys talyschensis Kapeller
- Stachys tamaulipana B.L.Turner
- Stachys tenerrima Epling
- Stachys tenuifolia Willd.
- Stachys tetragona Boiss. & Heldr.
- Stachys thirkei K.Koch
- Stachys thracica Davidov
- Stachys thunbergii Benth.
- Stachys tlaxiacana B.L.Turner
- Stachys tmolea Boiss.
- Stachys torresii B.L.Turner
- Stachys tournefortii Poir.
- Stachys trichophylla Baker
- Stachys trinervis Aitch. & Hemsl.
- Stachys truncata Kunze ex Benth.
- Stachys tubulosa MacOwan
- Stachys tundjeliensis Kit Tan & Sorger
- Stachys turcomanica Trautv.
- Stachys turkestanica (Regel) Popov ex Knorring
- Stachys turneri Rzed. & Calderón
- Stachys tymphaea Hausskn.
- Stachys tysonii Skan
- Stachys uniflora A.Pool
- Stachys urticoides Epling
- Stachys venezuelana Briq.
- Stachys venulosa Greene
- Stachys veroniciformis Rech.f.
- Stachys virgata Bory & Chaub.
- Stachys viscosa Montbret & Aucher ex Benth.
- Stachys viticina Boiss.
- Stachys vulnerabilis Rzed. & Calderón
- Stachys vuralii Yildiz, Dirmenci & Akçiçek
- Stachys willemsei Kit Tan & Hedge
- Stachys woronowii (Schischk. ex Grossh.) R.R.Mill
- Stachys xanthantha C.Y.Wu
- Stachys yemenensis Hedge
- Stachys yildirimlii Dinç
- Stachys zepcensis Formánek
- Stachys zeyheri Skan
- Stachys zoharyana Eig
Stachys L. appears in other Kew resources:
Date | Reference | Identified As | Barcode | Type Status |
---|---|---|---|---|
Ash [1024], Ethiopia | 35506.000 | |||
Sebsebe, D. [2312], Ethiopia | 54597.000 | |||
Balansa, B. [2927], Vietnam | K000853945 | |||
Balansa, B. [2928], Vietnam | K000853946 | |||
Smiles, H.H. [s.n.], Thailand | K000853944 |
First published in Sp. Pl.: 580 (1753)
Accepted by
- Krestovskaya, T. (2011). The critical review of species of the genus Stachys L. section Eriostemum Hoffmanns. et Link (Labiatae) Novosti Sistematiki Vysshikh Rastenii 42: 198-221.
- Govaerts, R. (2003). World Checklist of Selected Plant Families Database in ACCESS: 1-216203. The Board of Trustees of the Royal Botanic Gardens, Kew.
Literature
Flora of West Tropical Africa
- —F.T.A. 5: 465.
Flora of Somalia
- Flora Somalia, (2000) Author: by O. Ryding [updated by M. Thulin 2008]
- Sebsebe Demissew in Kew Bull. 48: 327–341 (1993).
Flora of Tropical East Africa
- Kubitzki, Fam. Gen. Vasc. Pl. 7: 223 (2004)
- K.B. 51: 433–454 (1996)
- K.B. 48: 327–341 (1993)
- Norw. Journ. Bot. 18: 121–137 (1971)
- N.B.G.B. 8: 625–646 (1923)
- Gen. Pl., ed. 5: 253 (1754)
- Sp. Pl. 580 (1753)
Lamiaceae Key Genus Fact Sheets
- Harley RM, Atkins S, Budantsev AL, Cantino PD, Conn BJ, Grayer R, Harley MM, de Kok RPJ, Krestovskaja T, Morales R, Paton AJ, Ryding O, and Upson T. 2004. Labiatae, in The Families and Genera of Vascular Plants (K. Kubitzki, ed. in chief) VI: 167-275. Sp
- Bhattacharjee, Notes Roy. Bot. Gard. Edinburgh 38: 65-96 (1980).
- Menitskia (Krestovsk.) Krestovsk., Bot. Zhurn. (Moscow & Leningrad) 91: 1893 (2006).
- Stachyus St.-Lag., Ann. Soc. Bot. Lyon 7: 109 (1880).
- Aspasia E.Mey. ex Pfeiff., Nomencl. Bot. 1: 295 (1873).
- Ortostachys Fourr., Ann. Soc. Linn. Lyon, n.s., 17: 136 (1869).
- Trixella Fourr., Ann. Soc. Linn. Lyon, n.s., 17: 135 (1869).
- Olisia Spach, Hist. Nat. Vég. 9: 166 (1840).
- Eriostemum Steud., Nomencl. Bot., ed. 2, 1: 588 (1840).
- Eriostomum Hoffmanns. & Link, Fl. Portug. 1: 105 (1809).
- Tetrahitum Hoffmanns. & Link, Fl. Portug. 1: 103 (1809).
- Galeopsis Moench, Methodus: 397 (1794), nom. illeg.
- Bonamya Neck., Elem. Bot. 1: 316 (1790), opus utique oppr.
- Trixago Haller, Hist. Stirp. Helv. 1: 101 (1768).
- Zietenia Gled., Syst. Pl. Stamin. Situ: 185 (1764).
- Tetrahit Gérard (1761).
- Galeopsis Hill, Brit. Herb.: 359 (1756), nom. illeg.
- Betonica L., Sp. Pl.: 573 (1753).
- Stachys L., Sp. Pl. 2: 580 (1753)
- Lamiostachys Krestovsk., Bot. Zhurn. (Moscow & Leningrad) 91: 1258 (2006).
Flora of Somalia
Flora of Somalia
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Flora of Tropical East Africa
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Herbarium Catalogue Specimens
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Kew Backbone Distributions
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Lamiaceae Key Genus Fact Sheets
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