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  1. Family: Polypodiaceae J.Presl & C.Presl
    1. Dryopteris Adans.

      1. This genus is accepted, and its native range is Cosmopolitan.


    Dryopteridaceae, JP Roux, M Shaffer-Fehre, B Verdcourt. Flora of Tropical East Africa. 2007

    Plants terrestrial or epilithic
    Rhizome erect to short-decumbent, simple or sparsely branched, densely scaled
    Fronds closely spaced, axes shallowly sulcate, sulci confluent or not.
    Leaf lamina
    Lamina anadromous, catadromous towards apex, herbaceous to firmly herbaceous, 2–4-pinnate-pinnatifid, basal pinna pair mostly basiscopically developed, adaxially proliferous along rachis or not.
    Leaf veins
    Venation free, lateral veins in lobes simple, forked, or pinnately branched, vein branches end in teeth near margin, endings often slightly enlarged.
    Indumentum composed of unicellular glands, uniseriate pluricellular, isocytic (all cells similar) or moniliform hairs, and scales occurring on rhizome, frond axes and lamina
    Sori circular, medial to inframedial on predominantly anadromous (often modified) vein branches; sporangium stalk simple, glandular and/ or haired; sori exindusiate or indusiate, indusium firmly herbaceous, deltoid to reniform, entire to erose, often glandular along margin and on surface
    Spores ellipsoidal, monolete, pale to dark brown, perispore folded to form reticulate ridges or echinulate.
    A genus of about 150 species with a nearly world-wide distribution.

    Dryopteridaceae, JP Roux, M Shaffer-Fehre, B Verdcourt. Flora of Tropical East Africa. 2007

    Medium-sized terrestrial ferns, rhizome erect or ascending, covered in dark, entire scales
    Fronds in tufts
    Stipes densely covered in scales, adaxial groove present, grooves of axes of higher order dwindling at base
    Leaf lamina
    Lamina deltoid-ovate, acuminate, tripinnatifid; lowest pinna basiscopically produced, anadromous, those above catadromous.
    Pinna bases equal, margin entire, rarely serrate.
    Leaf veins
    Veins dark, free, not reaching margin, apexes terminate in hydathodes
    Sori round, indusiate.
    A genus of five species from southern China, Taiwan, northern India, South Africa and the Madagascan region.



    Native to:

    Afghanistan, Alabama, Alaska, Albania, Alberta, Algeria, Altay, Amsterdam-St.Paul Is, Amur, Angola, Argentina Northwest, Arizona, Arkansas, Ascension, Assam, Austria, Azores, Baleares, Baltic States, Bangladesh, Belarus, Belgium, Bermuda, Bismarck Archipelago, Bolivia, Borneo, Brazil North, Brazil South, Brazil Southeast, Brazil West-Central, British Columbia, Bulgaria, Burundi, Buryatiya, California, Cambodia, Cameroon, Canary Is., Cape Provinces, Cape Verde, Caroline Is., Central African Repu, Central European Rus, China North-Central, China South-Central, China Southeast, Chita, Colombia, Colorado, Comoros, Connecticut, Cook Is., Corse, Costa Rica, Cuba, Cyprus, Czechoslovakia, Delaware, Denmark, District of Columbia, Dominican Republic, East European Russia, East Himalaya, Easter Is., Ecuador, El Salvador, Equatorial Guinea, Eritrea, Ethiopia, Fiji, Finland, Florida, France, Free State, Føroyar, Gabon, Galápagos, Georgia, Germany, Great Britain, Greece, Greenland, Guatemala, Guinea, Guinea-Bissau, Gulf of Guinea Is., Hainan, Haiti, Hawaii, Honduras, Hungary, Iceland, Idaho, Illinois, India, Indiana, Inner Mongolia, Iowa, Iran, Ireland, Irkutsk, Italy, Ivory Coast, Jamaica, Japan, Jawa, Kamchatka, Kansas, Kazakhstan, Kazan-retto, Kentucky, Kenya, Khabarovsk, Kirgizstan, Korea, Krasnoyarsk, Kriti, Krym, Kuril Is., KwaZulu-Natal, Labrador, Laos, Lebanon-Syria, Lesotho, Lesser Sunda Is., Liberia, Louisiana, Madagascar, Madeira, Magadan, Maine, Malawi, Malaya, Maluku, Manchuria, Manitoba, Marquesas, Maryland, Masachusettes, Mauritius, Mexico Central, Mexico Gulf, Mexico Northeast, Mexico Northwest, Mexico Southeast, Mexico Southwest, Michigan, Minnesota, Mississippi, Missouri, Mongolia, Montana, Morocco, Mozambique, Myanmar, Nansei-shoto, Nebraska, Nepal, Netherlands, Nevada, New Brunswick, New Guinea, New Hampshire, New Jersey, New Mexico, New York, Newfoundland, Nicaragua, Nigeria, North Carolina, North Caucasus, North Dakota, North European Russi, Northern Provinces, Northwest European R, Northwest Territorie, Norway, Nova Scotia, Nunavut, Ogasawara-shoto, Ohio, Oklahoma, Ontario, Oregon, Pakistan, Palestine, Panamá, Pennsylvania, Peru, Philippines, Poland, Portugal, Primorye, Prince Edward I., Qinghai, Queensland, Québec, Rhode I., Romania, Rwanda, Réunion, Sakhalin, Samoa, Sardegna, Saskatchewan, Seychelles, Sicilia, Sierra Leone, Society Is., Solomon Is., Somalia, South Carolina, South Dakota, Spain, Sri Lanka, St.Helena, Sudan, Sulawesi, Sumatera, Swaziland, Sweden, Switzerland, Tadzhikistan, Taiwan, Tanzania, Tennessee, Texas, Thailand, Tibet, Transcaucasus, Tristan da Cunha, Tunisia, Turkey, Turkey-in-Europe, Tuva, Uganda, Ukraine, Utah, Uzbekistan, Vanuatu, Venezuela, Vermont, Vietnam, Virginia, Washington, West Himalaya, West Siberia, West Virginia, Wisconsin, Wyoming, Xinjiang, Yakutskiya, Yemen, Yugoslavia, Yukon, Zambia, Zaïre, Zimbabwe

    Introduced into:

    Argentina South, Chatham Is., Falkland Is.

    Accepted Species


    Other Data

    Dryopteris Adans. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Aug 1, 2001 Etuge, M. [2062], Cameroon K000108985
    Aug 1, 2001 Etuge, M. [2102], Cameroon K000108986
    Kamau [621], Kenya K000696578
    Kamau [622], Kenya K000696579
    Comins, R.B. [s.n.], Solomon Is. Nephrodium K000636178 Unknown type material


    First published in Fam. Pl. 2: 20, 551 (1763)

    Accepted by

    • Fraser-Jenkins, C.R. & Kandel, D.R. (2019). Ferns and Fern-Allies of Nepal 2: 1-446. Government of Nepal, Ministry of Forests and Environment, Department of Plant Resources.
    • Roskov Y. & al. (eds.) (2018). World Ferns: Checklist of Ferns and Lycophytes of the World Species 2000 & ITIS Catalogue of Life Naturalis, Leiden, the Netherlands.


    Flora of Tropical East Africa

    • Kubitzki et al., Fam. Gen. Vasc. Pl. 1:110–112 (1990)
    • Fam. Pl. 2: 20, 551 (1763);


    Flora of Tropical East Africa
    Flora of Tropical East Africa

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2020. Published on the Internet at and
    © Copyright 2017 World Checklist of Selected Plant Families.

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2020. Published on the Internet at and
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families.