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  1. Family: Nyctaginaceae Juss.
    1. Pisonia Plum. ex L.

      1. This genus is accepted, and is native to Asia-Tropical, Africa, Pacific, Southern America, Australasia, Northern America and Asia-Temperate..


    Nyctaginaceae, B. L. Stannard. Flora Zambesiaca 9:1. 1988

    Scandent or erect shrubs or trees, unarmed or with axillary spines.
    Leaves opposite, alternate or verticillate, petiolate or sessile, simple, entire.
    Inflorescences paniculate cymes, axillary or terminal, dense or lax, subsessile or pedunculate.
    Flowers small, dioecious, rarely monoecious or hermaphrodite, male and female flowers sometimes of different shape; bracts 2–4, not involucrate.
    Perianth infundibuliform, campanulate, urceolate or tubular, 5–10 lobed or toothed; lobes erect, spreading or reflexed.
    Stamens variable in number usually 5–10 mostly exserted, rudimentary in female flowers; filaments connate below.
    Ovary sessile or stipitate, elongate, rudimentary in male flowers; style filiform, included or exserted; stigma lobed-capitate, multifid.
    Fruit enclosed in hardened, elongated, oblong or clavate, smooth or 4–5-angled perianth base (anthocarp); angles sometimes furnished with longitudinal rows of viscid stipitate glands.

    Nyctaginaceae, Christopher Whitehouse. Flora of Tropical East Africa. 1996

    Trees, shrubs or rarely lianas, sometimes with axillary spines
    Leaves opposite or alternate, sometimes clustered towards the ends of branches, entire, sessile or petiolate
    Inflorescences terminal or axillary cymes, clusters or panicles
    Flowers unisexual, usually on separate plants, or bisexual; perianth campanulate, urceolate or funnel-shaped, 5(–10)-lobed, often falling early; bracts 2–3 or absent, not forming an involucre
    Stamens 2–20, exserted; filaments united into a ring
    Ovary elongate-ovoid, sessile; style longer than ovary; stigma capitate, lobed or fimbriate
    Anthocarp oblong, linear or clavate, often 5(–6)-ribbed or sulcate, smooth, tuberculate or with sticky stipitate glands; embryo straight.



    Native to:

    Aldabra, Andaman Is., Argentina Northeast, Argentina Northwest, Arizona, Bahamas, Belize, Bolivia, Borneo, Brazil North, Brazil Northeast, Brazil South, Brazil Southeast, Brazil West-Central, Cambodia, Cayman Is., Colombia, Costa Rica, Cuba, Dominican Republic, Ecuador, El Salvador, Ethiopia, Fiji, Florida, Guatemala, Guyana, Hainan, Haiti, Hawaii, Honduras, India, Jamaica, Jawa, Kenya, KwaZulu-Natal, Laos, Leeward Is., Lesser Sunda Is., Madagascar, Malaya, Maluku, Marianas, Marquesas, Mauritius, Mexico Central, Mexico Gulf, Mexico Northeast, Mexico Northwest, Mexico Southeast, Mexico Southwest, Mozambique, Myanmar, New Caledonia, New Guinea, New South Wales, Nicaragua, Nicobar Is., Northern Territory, Panamá, Paraguay, Peru, Philippines, Pitcairn Is., Puerto Rico, Queensland, Réunion, Samoa, Saudi Arabia, Seychelles, Society Is., Solomon Is., Southwest Caribbean, Sri Lanka, Sulawesi, Sumatera, Taiwan, Tanzania, Texas, Thailand, Tonga, Tuamotu, Tubuai Is., Uganda, Venezuela, Venezuelan Antilles, Vietnam, Western Australia, Windward Is., Yemen

    Introduced into:

    Bangladesh, Benin, Cape Verde, Ghana, Ivory Coast, Nigeria, Sudan, Trinidad-Tobago

    Accepted Species


    Other Data

    Pisonia Plum. ex L. appears in other Kew resources:


    First published in Sp. Pl.: 1026 (1753)


    Flora of West Tropical Africa

    • —F.T.A. 6, 1: 8.

    Flora Zambesiaca

    • in Gen. Pl. ed. 5: 451 (1754).
    • Sp. Pl. 1: 1026 (1753)

    Flora of Tropical East Africa

    • Bittrich & U. Kühn in Kubitzki et al., Fam. Gen. Vasc. Pl. 2: 484 (1993)
    • Stemm. in Blumea 12: 275–284 (1964)
    • L., Gen. Pl., ed. 5: 451 (1754)
    • Sp. Pl.: 1026 (1753)


    Flora Zambesiaca
    Flora Zambesiaca

    Flora of Tropical East Africa
    Flora of Tropical East Africa

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2020. Published on the Internet at and
    © Copyright 2017 World Checklist of Selected Plant Families.

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2020. Published on the Internet at and
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families.

    Milliken, W., Klitgard, B. and Baracat, A. (2009 onwards), Neotropikey - Interactive key and information resources for flowering plants of the Neotropics.