1. Family: Amaranthaceae Juss.
    1. Genus: Gomphrena L.
      1. Gomphrena globosa L.

        This species is accepted, and its native range is Mexico to Brazil. It is used to treat unspecified medicinal disorders and as a soil improver.

    [FWTA]

    Amaranthaceae, Hutchinson and Dalziel. Flora of West Tropical Africa 1:1. 1954

    Note
    A native of S. America, cultivated as an ornamental, also subspontaneous.
    [UPB]

    The Useful Plants of Boyacá project

    Distribution
    Cultivated in Colombia.
    Habit
    Herb.
    Ecology
    Alt. 27 - 2600 m.
    [CPLC]

    Bernal, R., Gradstein, S.R. & Celis, M. (eds.). 2015. Catálogo de plantas y líquenes de Colombia. Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá. http://catalogoplantasdecolombia.unal.edu.co

    Distribution
    Cultivada en Colombia; Alt. 27 - 2600 m.; Amazonia, Andes, Islas Caribeñas, Orinoquia, Valle del Cauca.
    Habit
    Hierba
    [FTEA]

    Amaranthaceae, C.C. Townsend. Flora of Tropical East Africa. 1985

    Habit
    Annual herb, decumbent or erect, branched from the base and also above, ± 15–60 cm.; stem and branches striate or sulcate, ± densely clothed with appressed white hairs at least when young.
    Leaves
    Leaves broadly lanceolate to oblong or elliptic-oblong, 2.5–12(–15) × 2–4(–6) cm., narrowed to an ill-defined petiole below, thinly pilose on both surfaces, the pair of leaves subtending the terminal inflorescence sessile or almost so, broadly ovate to subcordate-ovate.
    Inflorescences
    Inflorescences sessile above the uppermost pair of leaves, usually solitary, globose or depressed-globose, rarely ovoid, ± 2 cm. in diameter, whitish to pinkish or deep red; bracts deltoid-ovate, 3–5 mm., mucronate with the shortly excurrent midrib; bracteoles strongly laterally compressed, navicular, ± 8–12 mm., mucronate with the excurrent midrib, furnished from the apex almost to the base of the dorsal surface of the midrib with a broad irregularly dentate crest.
    Tepal
    Tepals similar to those of G. celosioides but longer (6–6.5 mm.), the outer more lanate and the inner less markedly indurate at the base.
    Stamens
    Staminal tube subequalling the perianth, the 5 teeth deeply bilobed with obtuse lobes subequalling the anthers; pseudostaminodes absent.
    Pistil
    Style and stigmas together ± 2.5 mm.; stigmas divergent, subequalling or slightly longer than the style.
    Fruits
    Capsule oblong-ovoid, compressed, ± 2.5 mm.
    Seeds
    Seeds compressed-ovoid, ± 2 mm., brown, almost smooth, shining.
    [FZ]

    Amaranthaceae, C. C. Townsend. Flora Zambesiaca 9:1. 1988

    Habit
    Annual herb, decumbent or erect, branched from the base and also above, c. 15–60 cm.; stem and branches striate or sulcate, more or less densely clothed with appressed white hairs at least when young.
    Leaves
    Leaves broadly lanceolate to oblong or elliptic-oblong, 2.5–12 (15) × 2–4 (6) cm., narrowed to an ill-defined petiole below, thinly pilose on both surfaces, the pair of leaves subtending the terminal inflorescence sessile or almost so, broadly to subcordate-ovate.
    Inflorescences
    Inflorescences sessile above the uppermost pair of leaves, usually solitary, globose or depressed-globose, c. 2 cm. in diam., pinkish to deep red; bracts deltoid-ovate, 3–5 mm. long, mucronate with the shortly excurrent midrib; bracteoles strongly laterally compressed, navicular, c.8–12 mm., mucronate with the excurrent midrib, furnished from the apex almost to the base of the dorsal surface of the midrib with a broad, irregularly dentate crest.
    Tepal
    Tepals similar to those of G. celosioides but longer (6–6.5 mm.), the outer more lanate and the inner less markedly indurate at the base.
    Stamens
    Staminal tube subequalling the perianth, the 5 teeth deeply bilobed with obtuse lobes subequalling the anthers; pseudostaminodes absent.
    Style
    Style and stigmas together c. 2.5 mm. long, stigmas divergent, subequalling or slightly longer than the style.
    Fruits
    Capsule oblong-ovoid, compressed, c. 2.5 mm. long.
    Seeds
    Seeds compressed-ovoid, c. 2 mm. in diam., brown, almost smooth, shining.
    [UPB]
    Soil Improvers
    Live plant (in situ) - Used for food security, phytoremediation and crop rotation systems in arsenic-contaminated agricultural soils (Signes-Pastor et al. 2015).
    Unspecified Medicinal Disorders
    Medicinal (State of the World's Plants 2016, Instituto Humboldt 2014).

    Images

    Distribution

    Native to:

    Bolivia, Brazil North, Brazil Northeast, Brazil Southeast, Brazil West-Central, Colombia, Ecuador, Guatemala, Guyana, Honduras, Mexico Gulf, Mexico Northeast, Mexico Southeast, Mexico Southwest, Nicaragua, Panamá

    Introduced into:

    Alabama, Angola, Arkansas, Assam, Bahamas, Bangladesh, Benin, Brazil South, Burkina, Cambodia, Cameroon, Cape Verde, Cayman Is., Central African Repu, Chad, China North-Central, China South-Central, China Southeast, Congo, Cook Is., Cuba, Djibouti, Dominican Republic, Ethiopia, Fiji, Gilbert Is., Guinea, Guinea-Bissau, Gulf of Guinea Is., Haiti, India, Jamaica, Japan, Kenya, Korea, Laos, Leeward Is., Louisiana, Malawi, Malaya, Marquesas, Mauritius, Mozambique, Myanmar, New Caledonia, New Jersey, New South Wales, New York, Nicobar Is., Nigeria, Niue, Northern Provinces, Northern Territory, Paraguay, Pennsylvania, Peru, Pitcairn Is., Puerto Rico, Queensland, Samoa, Senegal, Sierra Leone, Society Is., Solomon Is., Somalia, South Carolina, Tadzhikistan, Tanzania, Texas, Tonga, Trinidad-Tobago, Tuamotu, Tubuai Is., Turkmenistan, Uganda, Uzbekistan, Vanuatu, Venezuela, Venezuelan Antilles, Vietnam, Virginia, Western Australia, Windward Is., Xinjiang, Zambia, Zaïre

    Common Names

    English
    Globe amaranth
    Spanish
    Inmortal, perpetuas, siempreviva, inmortales, manto de cristo, para todo, alumbra a las once, dondiego morado, sandiego.

    Gomphrena globosa L. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    s.coll. [Cat. no. 6920] K001126212
    s.coll. [Cat. no. 6920] K001126213
    s.coll. [Cat. no. 6920] K001126214

    First published in Sp. Pl.: 224 (1753)

    Accepted by

    • Pedersen, T.M. (2016). Flora del Paraguay 46: 1-258. Editions des Conservatoire et Jardin Botaniques de la ville de Genève.
    • Baksh-Comeau, Y., Maharaj, S.S., Adams, C.D., Harris, S.A., Filer, D.L. & Hawthorne, W.D. (2016). An annotated checklist of the vascular plants of Trinidad and Tobago with analysis of vegetation types and botanical 'hotspots' Phytotaxa 250: 1-431.
    • Sykes, W.R. (2016). Flora of the Cook Islands: 1-973. National Tropical Botanical Garden, Hawaii.
    • Chang, C.S., Kim, H. & Chang, K.S. (2014). Provisional checklist of vascular plants for the Korea peninsula flora (KPF): 1-660. DESIGNPOST.
    • Serviss, B.E. & Peck, J.H. (2013). Rediscovery of Persea borbonia var. borbonia (Lauraceae), Prosopis glandulosa var. glandulosa (Fabaceae), and Pinus palustris (Pinaceae) in Arkansas Journal of the Botanical Research Institute of Texas 7: 841-845.
    • Sánchez-del Pino, I., Espadas, C. & Pool, R. (2013). Taxonomy and richness of nine genera of Amaranthaceae s.s. (Caryophyllales) in the Yucatan peninsula biotic province Phytotaxa 107: 1-74.
    • Mostaph, M.K. & Uddin, S.B. (2013). Dictionary of plant names of Bangladesh, Vasc. Pl.: 1-434. Janokalyan Prokashani, Chittagong, Bangladesh.
    • Thiombiano, A., Schmidt, M., Dressler, S., Ouédraogo, A., Hahn, K. & Zizka, G. (2012). Catalogue des plantes vasculaires du Burkina Faso Boissiera 65: 1-391.
    • Singh, A. (2012). Exotic flora of the Chandauli district Uttar Pradesh, India: an overview Indian Journal of Forestry 35: 79-84.
    • Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies Smithsonian Contributions to Botany 98: 1-1192.
    • Garcia-Mendoza, A.J. & Meave, J.A. (eds.) (2012). Diversidad florística de Oaxaca: de musgos a angiospermas (colecciones y listas de especies), ed. 2: 1-351. Instituto de Biología, Universidad Nacional Autónoma de México.
    • Figueiredo, E., Paiva, J., Stévart, T., Oliveira, F. & Smith, G.F. (2011). Annotated catalogue of the flowering plants of São Tomé and Príncipe Bothalia, A Journal of Botanical Research 41: 41-82.
    • Idárraga-Piedrahita, A., Ortiz, R.D.C., Callejas Posada, R. & Merello, M. (eds.) (2011). Flora de Antioquia: Catálogo de las Plantas Vasculares 2: 1-939. Universidad de Antioquia, Medellín.
    • Kral, R., Diamond, A.R., Ginzbarg, S.L., Hansen, C.J., Haynes, R.R., Keener, B.R., Lelong, M.G., Spaulding, D.D. & Woods, M. (2011). Annotated checklist of the vascular plants of Alabama: 1-112. Botanical reseach institute of Texas.
    • Onana, J.M. (2011). The vascular plants of Cameroon a taxonomic checklist with IUCN assessments: 1-195. National herbarium of Cameroon, Yaoundé.
    • Ramos Senna, L., Giulietti, A.M. & Rapini, A. (2010). Flora da Bahia: Amaranthaceae - Amaranthoideae e Gomphrenoideae Sitientibus Sérrie Ciências Biológicas 10: 3-73.
    • Lejoy, J., Ndjele, M.-B. & Geerinck, D. (2010). Catalogue-flore des plantes vasculaires des districts de Kisangani et de la Tshopo (RD Congo) Taxonomania 30: 1-307.
    • Lisowski, S. (2009). Flore (Angiospermes) de la République de Guinée Scripta Botanica Belgica 41: 1-517.
    • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
    • Hokche, O., Berry, P.E. & Huber, O. (eds.) (2008). Nuevo Catálogo de la Flora Vascular de Venezuela: 1-859. Fundación Instituto Botánico de Venezuela.
    • Nelson Sutherland, C.H. (2008). Catálogo de las plantes vasculares de Honduras. Espermatofitas: 1-1576. SERNA/Guaymuras, Tegucigalpa, Honduras.
    • Newman, M., Ketphanh, S., Svengsuksa, B., Thomas, P., Sengdala, K., Lamxay, V. & Armstrong, K. (2007). A checklist of the vascular plants of Lao PDR: 1-394. Royal Botanic Gardens, Edinburgh.
    • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
    • Catarino, L., Sampaio Martins, E., Pinto-Basto, M.F. & Diniz, M.A. (2006). Plantas Vasculares e Briófitos da Guiné-Bissau: 1-298. Instituto de investigação científica tropical, Instituto Português de apoio ao desenvolvimento.
    • Sita, P. & Moutsambote, J.-M. (2005). Catalogue des plantes vasculaires du Congo, ed. sept. 2005: 1-158. ORSTOM, Centre de Brazzaville.
    • Correa A., Mireya D. Galdames, Carmen Correa A., M. D., C. Galdames & M. S. de Stapf (2004). Catálogo de las Plantas Vasculares de Panamá: 1-599. Smithsonian Tropical Research Institute.
    • Kress, W.J., DeFilipps, R.A., Farr, E. & Kyi, D.Y.Y. (2003). A Checklist of the Trees, Shrubs, Herbs and Climbers of Myanmar Contributions from the United States National Herbarium 45: 1-590. Smithsonian Institution.
    • Germishuizen, G. & Meyer, N.L. (eds.) (2003). Plants of Southern Africa an annotated checklist Strelitzia 14: 1-1231. National Botanical Institute, Pretoria.
    • Flora of North America Editorial Committee (2003). Flora of North America North of Mexico 4: 1-559. Oxford University Press, New York, Oxford.
    • Govaerts, R. (2003). World Checklist of Seed Plants Database in ACCESS G: 1-40325.
    • Lê, T.C. (2003). Danh l?c các loài th?c v?t Vi?t Nam 2: 1-1203. Hà N?i : Nhà xu?t b?n Nông nghi?p.
    • Edwards, S., Tadesse, M., Demissew, S. & Hedberg, I. (eds.) (2000). Flora of Ethiopia and Eritrea 2(1): 1-532. The National Herbarium, Addis Ababa University, Ethiopia & The Department of Systematic Botany, Upps.
    • Dy Phon, P. (2000). Dictionnaire des plantes utilisées au Cambodge: 1-915. chez l'auteur, Phnom Penh, Cambodia.
    • Bosser, J. & al. (eds.) (1994). Flore des Mascareignes 136-148: 1. IRD Éditions, MSIRI, RBG-Kew, Paris.
    • MacKee, H.S. (1994). Catalogue des plantes introduites et cultivées en Nouvelle-Calédonie, ed. 2: 1-164. Museum national d'histoire naturelle, Paris.
    • Hansen, A. & Sunding, P. (1993). Flora of Macaronesia. Checklist of vascular plants. 4. revised edition Sommerfeltia 17: 1-295.
    • Thulin, M. (ed.) (1993). Flora of Somalia 1: 1-493. The Royal Botanic Gardens, Kew.
    • Launert, E. (ed.) (1988). Flora Zambesiaca 9(1): 1-179. Royal Botanic Gardens, Kew.
    • Hancock, I.R. & Henderson, C.P. (1988). Flora of the Solomon Islands Research Bulletin Dodo Creek Research Station 7: 1-203.
    • Fosberg, F.R. & Sachet, M.-H. (1987). Flora of the Gilbert Island, Kiribati, Checklist Atoll Research Bulletin 295: 1-33.
    • Townsend, C.C. (1985). Flora of Tropical East Africa, Amaranthaceae: 1-136.
    • Boulvert, Y. (1977). Catalogue de la Flore de Centrafrique 2(1): 1-85. ORSTROM, Bangui.
    • Lebrun, J.-P., Audru, J., Gaston, A. & Mosnier, M. (1972). Catalogue des Plantes Vasculaires du Tchad Méridional: 1-289. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
    • Berhaut, J. (1971). Flore illustrée du Sénégal 1: 1-626. Gouvernement du Sénégal, Ministère du développement rural direction des eaux et forêta, Dakar.
    • Sykes, W.R. (1970). Contributions to the flora of Niue Bulletin, New Zealand Department of Scientific and Industrial Research 200: 1-321.
    • Ovczinnikov, P.N. (ed.) (1968). Flora Tadzhikskoi SSR 3: 1-710. Izd-vo Akademii nauk SSSR, Moskva.
    • Hutchinson, J., Dalziel, J.M. & Keay, R.W.J. (1954-1958). Flora of West Tropical Africa, ed. 2, 1: 1-828.
    • Korovin, E.P. & Vvedensky, A.I. (eds.) (1953). Flora Uzbekistana 2: 1-547. Izd-va Akademii nauk Uzbekskoi SSR, Tashkent.
    • B.A.Fedtschenko & al. (1948). Flora Turkmenii 3: 1-280. Turkmenskoe gosudarstvennoe izd., Ashkhabad.

    Literature

    Flora of West Tropical Africa
    • —F.T.A. 6, 1: 75.
    Useful Plants of Boyacá Project
    • Kew Medicinal Plant Names Services (MPNS) https://www.kew.org/science/data-and-resources/tools-and-services/medicinal-plant-names-services in The State of the World’s Plants Report–2016. (2016). Royal Botanic Gardens, Kew https://stateoftheworldsplants.org
    • Signes-Pastor, A. J., Munera-Picazo, S., Burló, F., Cano-Lamadrid, M., & Carbonell-Barrachina, A. A. (2015). Phytoremediation assessment of Gomphrena globosa and Zinnia elegans grown in arsenic-contaminated hydroponic conditions as a safe and feasible alternative to be applied in arsenic-contaminated soils of the Bengal Delta. Environmental monitoring and assessment, 187(6), 387.
    • Instituto de Investigación de Recursos Biológicos Alexander von Humbodlt (2014). Plantas alimenticias y medicinales nativas de Colombia. 2567 registros, aportados por: Castellanos, C. (Contacto del recurso), Valderrama, N. (Creador del recurso, Autor), Castro, C. (Proveedor de metadatos), Bernal, Y. (Autor), García, N. (Autor). Versión 11.0. http://i2d.humboldt.org.co/ceiba/resource.do?r=ls_colombia_magnoliophyta_2014
    Kew Backbone Distributions
    • Forzza, R.C., Zappi, D. & Souza, V.C. (2016-continuously updated). Flora do Brasil 2020 em construção http://reflora.jbrj.gov.br/reflora/listaBrasil/ConsultaPublicaUC/ResultadoDaConsultaNovaConsulta.do.
    • Pedersen, T.M. (2016). Flora del Paraguay 46: 1-258. Editions des Conservatoire et Jardin Botaniques de la ville de Genève.
    • Baksh-Comeau, Y., Maharaj, S.S., Adams, C.D., Harris, S.A., Filer, D.L. & Hawthorne, W.D. (2016). An annotated checklist of the vascular plants of Trinidad and Tobago with analysis of vegetation types and botanical 'hotspots' Phytotaxa 250: 1-431.
    • Sykes, W.R. (2016). Flora of the Cook Islands: 1-973. National Tropical Botanical Garden, Hawaii.
    • Chang, C.S., Kim, H. & Chang, K.S. (2014). Provisional checklist of vascular plants for the Korea peninsula flora (KPF): 1-660. DESIGNPOST.
    • Serviss, B.E. & Peck, J.H. (2013). Rediscovery of Persea borbonia var. borbonia (Lauraceae), Prosopis glandulosa var. glandulosa (Fabaceae), and Pinus palustris (Pinaceae) in Arkansas Journal of the Botanical Research Institute of Texas 7: 841-845.
    • Sánchez-del Pino, I., Espadas, C. & Pool, R. (2013). Taxonomy and richness of nine genera of Amaranthaceae s.s. (Caryophyllales) in the Yucatan peninsula biotic province Phytotaxa 107: 1-74.
    • Mostaph, M.K. & Uddin, S.B. (2013). Dictionary of plant names of Bangladesh, Vasc. Pl.: 1-434. Janokalyan Prokashani, Chittagong, Bangladesh.
    • Thiombiano, A., Schmidt, M., Dressler, S., Ouédraogo, A., Hahn, K. & Zizka, G. (2012). Catalogue des plantes vasculaires du Burkina Faso Boissiera 65: 1-391.
    • Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies Smithsonian Contributions to Botany 98: 1-1192.
    • Figueiredo, E., Paiva, J., Stévart, T., Oliveira, F. & Smith, G.F. (2011). Annotated catalogue of the flowering plants of São Tomé and Príncipe Bothalia, A Journal of Botanical Research 41: 41-82.
    • Kral, R., Diamond, A.R., Ginzbarg, S.L., Hansen, C.J., Haynes, R.R., Keener, B.R., Lelong, M.G., Spaulding, D.D. & Woods, M. (2011). Annotated checklist of the vascular plants of Alabama: 1-112. Botanical reseach institute of Texas.
    • Onana, J.M. (2011). The vascular plants of Cameroon a taxonomic checklist with IUCN assessments: 1-195. National herbarium of Cameroon, Yaoundé.
    • Ramos Senna, L., Giulietti, A.M. & Rapini, A. (2010). Flora da Bahia: Amaranthaceae - Amaranthoideae e Gomphrenoideae Sitientibus Sérrie Ciências Biológicas 10: 3-73.
    • Buragohain, S. & Sarma, G.C. (2009). The exotic weeds of Guwahati, Assam and their role in employment generation Pleione 3(1): 45-49.
    • Lisowski, S. (2009). Flore (Angiospermes) de la République de Guinée Scripta Botanica Belgica 41: 1-517.
    • GBIF (2008-continuously updated). Global Biodiversity Information Facility http://www.gbif.org/.
    • Pandey, R.P. & Dilwakar, P.G. (2008). An integrated check-list flora of Andaman and Nicobar islands, India Journal of Economic and Taxonomic Botany 32: 403-500.
    • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
    • Hokche, O., Berry, P.E. & Huber, O. (eds.) (2008). Nuevo Catálogo de la Flora Vascular de Venezuela: 1-859. Fundación Instituto Botánico de Venezuela.
    • Nelson Sutherland, C.H. (2008). Catálogo de las plantes vasculares de Honduras. Espermatofitas: 1-1576. SERNA/Guaymuras, Tegucigalpa, Honduras.
    • Newman, M., Ketphanh, S., Svengsuksa, B., Thomas, P., Sengdala, K., Lamxay, V. & Armstrong, K. (2007). A checklist of the vascular plants of Lao PDR: 1-394. Royal Botanic Gardens, Edinburgh.
    • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
    • Catarino, L., Sampaio Martins, E., Pinto-Basto, M.F. & Diniz, M.A. (2006). Plantas Vasculares e Briófitos da Guiné-Bissau: 1-298. Instituto de investigação científica tropical, Instituto Português de apoio ao desenvolvimento.
    • Sita, P. & Moutsambote, J.-M. (2005). Catalogue des plantes vasculaires du Congo, ed. sept. 2005: 1-158. ORSTOM, Centre de Brazzaville.
    • Correa A., Mireya D. Galdames, Carmen Correa A., M. D., C. Galdames & M. S. de Stapf (2004). Catálogo de las Plantas Vasculares de Panamá: 1-599. Smithsonian Tropical Research Institute.
    • Kress, W.J., DeFilipps, R.A., Farr, E. & Kyi, D.Y.Y. (2003). A Checklist of the Trees, Shrubs, Herbs and Climbers of Myanmar Contributions from the United States National Herbarium 45: 1-590. Smithsonian Institution.
    • Germishuizen, G. & Meyer, N.L. (eds.) (2003). Plants of Southern Africa an annotated checklist Strelitzia 14: 1-1231. National Botanical Institute, Pretoria.
    • Flora of North America Editorial Committee (2003). Flora of North America North of Mexico 4: 1-559. Oxford University Press, New York, Oxford.
    • Lê, T.C. (2003). Danh l?c các loài th?c v?t Vi?t Nam 2: 1-1203. Hà N?i : Nhà xu?t b?n Nông nghi?p.
    • Edwards, S., Tadesse, M., Demissew, S. & Hedberg, I. (eds.) (2000). Flora of Ethiopia and Eritrea 2(1): 1-532. The National Herbarium, Addis Ababa University, Ethiopia & The Department of Systematic Botany, Upps.
    • Dy Phon, P. (2000). Dictionnaire des plantes utilisées au Cambodge: 1-915. chez l'auteur, Phnom Penh, Cambodia.
    • Bosser, J. & al. (eds.) (1994). Flore des Mascareignes 136-148: 1. IRD Éditions, MSIRI, RBG-Kew, Paris.
    • Audru, J., Cesar, J. & Lebrun, J.-P. (1994). Les Plantes Vasculaires de la République de Djibouti. Flore Illustrée 1: 1-336. CIRAD, Départerment d'Elevage et de Médecine vétérinaire, Djibouti.
    • MacKee, H.S. (1994). Catalogue des plantes introduites et cultivées en Nouvelle-Calédonie, ed. 2: 1-164. Museum national d'histoire naturelle, Paris.
    • Hansen, A. & Sunding, P. (1993). Flora of Macaronesia. Checklist of vascular plants. 4. revised edition Sommerfeltia 17: 1-295.
    • Thulin, M. (ed.) (1993). Flora of Somalia 1: 1-493. The Royal Botanic Gardens, Kew.
    • Launert, E. (ed.) (1988). Flora Zambesiaca 9(1): 1-179. Royal Botanic Gardens, Kew.
    • Hancock, I.R. & Henderson, C.P. (1988). Flora of the Solomon Islands Research Bulletin Dodo Creek Research Station 7: 1-203.
    • Fosberg, F.R. & Sachet, M.-H. (1987). Flora of the Gilbert Island, Kiribati, Checklist Atoll Research Bulletin 295: 1-33.
    • Townsend, C.C. (1985). Flora of Tropical East Africa, Amaranthaceae: 1-136.
    • Boulvert, Y. (1977). Catalogue de la Flore de Centrafrique 2(1): 1-85. ORSTROM, Bangui.
    • Lebrun, J.-P., Audru, J., Gaston, A. & Mosnier, M. (1972). Catalogue des Plantes Vasculaires du Tchad Méridional: 1-289. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
    • Berhaut, J. (1971). Flore illustrée du Sénégal 1: 1-626. Gouvernement du Sénégal, Ministère du développement rural direction des eaux et forêta, Dakar.
    • Sykes, W.R. (1970). Contributions to the flora of Niue Bulletin, New Zealand Department of Scientific and Industrial Research 200: 1-321.
    • Ovczinnikov, P.N. (ed.) (1968). Flora Tadzhikskoi SSR 3: 1-710. Izd-vo Akademii nauk SSSR, Moskva.
    • Yuncker, T.G. (1959). Plants of Tonga Bernice P. Bishop Museum Bulletin 220: 1-283.
    • Hutchinson, J., Dalziel, J.M. & Keay, R.W.J. (1954-1958). Flora of West Tropical Africa, ed. 2, 1: 1-828.
    • Korovin, E.P. & Vvedensky, A.I. (eds.) (1953). Flora Uzbekistana 2: 1-547. Izd-va Akademii nauk Uzbekskoi SSR, Tashkent.
    • Guillaumin, A. (1948). Compendium de la Flore Phanérogamique des Nouvelles Hébrides Annales de l'Institut Botanico-Geologique de Marseille, VI, 5-6: 1-56.
    • B.A.Fedtschenko & al. (1948). Flora Turkmenii 3: 1-280. Turkmenskoe gosudarstvennoe izd., Ashkhabad.
    • Christopherson, E. (1935). Flowering Plants of Samoa Bernice P. Bishop Museum Bulletin 128: 1-221.
    Flora of Tropical East Africa
    • Cavaco in Mém. Mus. Nat. Hist. Nat. Paris, sér. B. 13: 167 (1962).
    • Cufod., Enumeratio Plantarum Aethiopiae Spermatophyta (Supplement in Bull. Jard. Bot. Brux.) p. 75 (1953).
    • Hauman in Flore du Congo Belge et du Ruanda-Urundi, 2: 80 (1951).
    • R. O. Williams, Useful and Ornamental Plants in Zanzibar and Pemba p. 277 (1949).
    • Peter, Flora von Deutsch-Ostafrika 2: 245 (1932).
    • J. G. Baker & C. B. Clarke in Flora of Tropical Africa 6(1): 75 (1909).
    • Gilg in Die Pflanzenwelt Ost-Afrikas und der Nachbargebiete, Theile C: 174 (1895).
    • L., Sp. Pl.: 224 (1753).

    Sources

    Catálogo de Plantas y Líquenes de Colombia
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of West Tropical Africa
    Flora of West Tropical Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Useful Plants of Boyacá Project
    ColPlantA database
    http://creativecommons.org/licenses/by-nc-sa/3.0
    http://creativecommons.org/licenses/by/3.0/