1. Family: Phyllanthaceae Martinov
    1. Margaritaria L.f.

      1. This genus is accepted, and its native range is Tropics & Subtropics.

    [FTEA]

    Euphorbiaceae, A. R.-Smith. Flora of Tropical East Africa. 1987

    Habit
    Dioecious trees or shrubs with a simple indumentum
    Leaves
    Leaves alternate, shortly petiolate, stipulate, simple, entire, penninerved
    Flowers
    Flowers axillary, usually in the proximal axils of leafy branches, the males several, fasciculate, the females 2–3 per axil or often solitary Male flowers: pedicels capillary; calyx-lobes 4, imbricate; petals 0; disc annular, ± entire; stamens 4, free, anthers extrorse, longitudinally dehiscent; pistillode 0 Female flowers: pedicels stouter than in the ♂ flowers; calyx-lobes and disc ± as in the ♂ flowers; ovary smooth, commonly glabrous, (2–)3–4(–5)-locular, with 2 ovules per locule; styles free or united at the base or for 1/2 their length, shortly bifid, rarely bifid
    Male
    Male flowers: pedicels capillary; calyx-lobes 4, imbricate; petals 0; disc annular, ± entire; stamens 4, free, anthers extrorse, longitudinally dehiscent; pistillode 0
    Female
    Female flowers: pedicels stouter than in the ♂ flowers; calyx-lobes and disc ± as in the ♂ flowers; ovary smooth, commonly glabrous, (2–)3–4(–5)-locular, with 2 ovules per locule; styles free or united at the base or for 1/2 their length, shortly bifid, rarely bifid
    Fruits
    Fruit (2–)3–4(–5)-lobed, irregularly dehiscent or subindehiscent; endocarp paleaceous-crustaceous
    Seeds
    Seeds 2 per locule, sometimes 1 by abortion, hemispherical, smooth or rugose, ecarunculate; endosperm copious; embryo ± straight; cotyledons thin, flat, much longer than the radicle.
    [FZ]

    Euphorbiaceae, A. Radcliffe-Smith. Flora Zambesiaca 9:4. 1996

    Habit
    Dioecious, very rarely monoecious, trees or shrubs, rarely scandent.
    Indumentum
    Indumentum simple.
    Leaves
    Leaves distichous, shortly petiolate, stipulate, simple, entire, penninerved, usually deciduous.
    Flowers
    Flowers axillary, usually in the proximal axils of new shoots; the males usually several, fasciculate, often precocious; the females 2–3 per axil or often solitary. Male flowers: pedicels capillary; calyx lobes 4, imbricate; petals absent; disk annular, ± entire; stamens 4, filaments free, anthers extrorse, longitudinally dehiscent; pistillode (non-functional ovary) absent. Female flowers: pedicels stouter than in the male flowers; calyx lobes and disk ± as in the male flowers; ovary smooth, commonly glabrous, (2)3–4(5)-locular, ovules 2 per locule; styles free or variously united, bifid to bipartite.
    Fruits
    Fruit (2)3–4(5)-lobed, irregularly dehiscent or subindehiscent; exocarp green, usually separating from the paleaceous-crustaceous endocarp.
    Seeds
    Seeds 2 per locule, sometimes 1 by abortion, hemispherical, ecarunculate; sarcotesta fleshy, bluish; sclerotesta thick, woody or bony, smooth or rugose, chalaza invaginated; endosperm copious, whitish; embryo ± straight; cotyledons thin, flat, much longer than the radicle.

    Images

    Distribution

    Native to:

    Aldabra, Andaman Is., Angola, Argentina Northeast, Bahamas, Bangladesh, Belize, Benin, Bolivia, Borneo, Botswana, Brazil North, Brazil Northeast, Brazil South, Brazil Southeast, Brazil West-Central, Burkina, Burundi, Cameroon, Caprivi Strip, Cayman Is., Central African Repu, China Southeast, Colombia, Comoros, Congo, Costa Rica, Cuba, Dominican Republic, Ecuador, El Salvador, Ethiopia, French Guiana, Gabon, Galápagos, Gambia, Ghana, Guatemala, Guinea, Guinea-Bissau, Gulf of Guinea Is., Guyana, Haiti, Honduras, India, Ivory Coast, Jamaica, Jawa, Kenya, KwaZulu-Natal, Leeward Is., Lesser Sunda Is., Liberia, Madagascar, Malawi, Malaya, Mali, Mali, Maluku, Mauritius, Mexico Central, Mexico Gulf, Mexico Northeast, Mexico Northwest, Mexico Southeast, Mexico Southwest, Mozambique, Myanmar, New Guinea, Nicaragua, Nicobar Is., Nigeria, Northern Provinces, Panamá, Paraguay, Peru, Philippines, Puerto Rico, Queensland, Senegal, Sierra Leone, Southwest Caribbean, Sri Lanka, Sudan, Sumatera, Suriname, Swaziland, Taiwan, Tanzania, Thailand, Togo, Trinidad-Tobago, Uganda, Venezuela, Venezuelan Antilles, Vietnam, Windward Is., Zambia, Zaïre, Zimbabwe

    Margaritaria L.f. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Milliken, W. [1838], Brazil 59730.000

    First published in Suppl. Pl.: 66 (1782)

    Accepted by

    • Govaerts, R., Frodin, D.G. & Radcliffe-Smith, A. (2000). World Checklist and Bibliography of Euphorbiaceae (and Pandaceae) 1-4: 1-1622. The Board of Trustees of the Royal Botanic Gardens, Kew.

    Literature

    Flora Zambesiaca
    • G.L. Webster in J. Arnold Arbor. 60: 407 (1979).
    • Suppl. Pl.: 66 (1781).
    Flora of Tropical East Africa
    • Webster in Journ. Arn. Arb. 60: 403–444 (1979)
    • Suppl. Pl. Syst. Veg.: 66 (1781)

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0