1. Family: Rubiaceae Juss.
    1. Bertiera Aubl.

      1. This genus is accepted, and its native range is Mexico to Tropical America, Tropical Africa to W. Indian Ocean.

    [FTEA]

    Rubiaceae, B. Verdcourt. Flora of Tropical East Africa. 1976

    Habit
    Small trees or shrubs, rarely lianas
    Leaves
    Leaves shortly petiolate; petioles normally channelled above; blades thin or scarcely coriaceous, mid- and lateral nerves usually prominent beneath; stipules persistent, usually connate above the nodes for a short distance
    Flowers
    Flowers of moderate size (rarely very small), borne in terminal, spicate thyrsoid panicles, scorpioid cymes or spherical compact heads, very occasionally axillary (not in Flora area)
    Calyx
    Calyx with limb produced above the tube, cup-shaped and frequently wider than the tube, truncate or 5-toothed or -lobed
    Corolla
    Corolla with hairy throat, often with the walls of tube toughened or thick; tube exceeding the calyx-limb; lobes (4–)5, usually acuminate or apiculate, contorted in bud
    Anthers
    Anthers (4–)5, subsessile, attached by very short filaments near the base, included, linear, sagittate or bilobed at the base, the connective extended into an apiculate apex
    Pistil
    Ovary 2-locular with the numerous ovules inserted on a thickened placenta; disc glabrous, fleshy, cup-shaped, or annular; stigma borne on style ± club-shaped, formed from 2 flattened appressed branches (seldom separating), with 10 membranous linear vanes which fit between the anthers in the bud
    Fruits
    Fruit indehiscent, ovoid or globose, often coriaceous; calyx persistent; pedicels sometimes accrescent
    Seeds
    Seeds red-brown to black, angular, rugulose or granulose, numerous.
    [FZ]

    Rubiaceae, D.M.Bridson & B.Verdcourt. Flora Zambesiaca 5:3. 2003

    Habit
    Small trees or shrubs, rarely lianes.
    Leaves
    Leaves shortly petiolate; blades thin or scarcely coriaceous, mid and lateral veins usually prominent beneath; petioles normally channelled above.
    Stipules
    Stipules persistent, usually connate above the nodes for a short distance.
    Flowers
    Flowers hermaphrodite, 5-merous, of moderate size (rarely very small), borne in terminal, spicate thyrsoid panicles, scorpioid cymes or spherical compact heads.
    Calyx
    Calyx limb-tube cup-shaped and frequently wider than the tube in flower, but narrower in fruit, truncate or 5-toothed or -lobed.
    Corolla
    Corolla with a hairy throat, often with the walls of tube toughened or thick; tube exceeding the calyx limb; lobes (4)5, usually acuminate or apiculate, contorted to the left in the bud.
    Disc
    Disk cup-shaped or annular, glabrous, fleshy.
    Anthers
    Anthers (4)5, included, subsessile, attached by very short filaments near the base, linear, sagittate or bilobed at the base; connective extended into an apiculate apex.
    Ovary
    Ovary 2-celled with the numerous ovules inserted on a thickened placenta; pollen presenter ± club-shaped, formed from 2 flattened appressed branches (seldom separating), with 10 membranous linear vanes which fit between the anthers in bud.
    Fruits
    Fruit indehiscent, ovoid or globose, often coriaceous; calyx limb persistent and pedicels sometimes accrescent.
    Seeds
    Seeds numerous, red-brown to black, angular, rugulose or granulose.

    Images

    Distribution

    Native to:

    Angola, Belize, Benin, Bolivia, Brazil North, Brazil West-Central, Cabinda, Cameroon, Central African Repu, Central American Pac, Colombia, Congo, Costa Rica, Cuba, Dominican Republic, Ecuador, Equatorial Guinea, French Guiana, Gabon, Ghana, Guatemala, Guinea, Guinea-Bissau, Gulf of Guinea Is., Guyana, Honduras, Ivory Coast, Liberia, Madagascar, Mauritius, Mexico Gulf, Mexico Southeast, Mexico Southwest, Nicaragua, Nigeria, Panamá, Paraguay, Peru, Rwanda, Réunion, Senegal, Sierra Leone, Suriname, Tanzania, Togo, Trinidad-Tobago, Uganda, Venezuela, Zambia, Zaïre

    Bertiera Aubl. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Jan 1, 2008 Zapfack, L. [900], Cameroon K000436860
    Jan 1, 1996 Ekema, N. [1120a], Cameroon K000085330
    Coode, M. [4230], Reunion 33406.000
    Richardson, I. [4036], Mauritius 40027.000
    Davis, A.P. [3017], Cameroon 70351.000
    Sonké, B. [372], Cameroon K000093851

    First published in Hist. Pl. Guiane: 180 (1775)

    Accepted by

    • Govaerts, R. (2003). World Checklist of Selected Plant Families Database in ACCESS: 1-216203. The Board of Trustees of the Royal Botanic Gardens, Kew.
    • Govaerts, R. (1996). World Checklist of Seed Plants 2(1, 2): 1-492. MIM, Deurne.

    Literature

    Flora of West Tropical Africa
    • N. Hallé in Notulae Syst. 16: 280 (1960).
    • Wernham in J. Bot. 50: 110 (1912)
    • —F.T.A. 3: 82
    Flora Zambesiaca
    • Robbrecht, Rohrhofer & Puff in Opera Bot. Belg. 6: 101–141 (1993).
    • in Adansonia, n.s. 4: 457–459 (1964).
    • in Adansonia, n.s. 3: 294–306 (1963)
    • N. Hallé in Notul. Syst. (Paris) 16: 280–292 (1960)
    • Wernham in J. Bot. 50: 110–117 & 156–164 (1912).
    • Hist. Pl. Guiane1: 180, t. 69 (1775).
    Flora of Tropical East Africa
    • in Adansonia sér. 2, 4: 457–459 (1964)
    • in Adansonia sér. 2, 3: 294–306 (1963)
    • Hallé in Not. Syst. 16: 280–292 (1960)
    • Wernham in J.B. 50: 110–117 & 156–164 (1912)
    • Hist. Pl. Guiane Fr. 1: 180, t. 69 (1775)

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0