1. Family: Verbenaceae J.St.-Hil.
    1. Phyla Lour.

      1. This genus is accepted, and its native range is Tropics & Subtropics.

    [FTEA]

    Verbenaceae, B. Verdcourt. Flora of Tropical East Africa. 1992

    Habit
    Trailing perennial herbs with angular stems rooting at the nodes
    Leaves
    Leaves opposite, simple, subentire to toothed, mostly tapering into an obscure petiole; lamina with sparse to dense adpressed medifixed hairs
    Inflorescences
    Inflorescences ± long-pedunculate axillary short ovoid to cylindric very dense bracteate spikes
    Calyx
    Calyx membranous, 2-lobed, compressed
    Corolla
    Corolla with short tube and spreading somewhat irregularly 4-lobed white or purple limb
    Stamens
    Stamens 4, included
    Fruits
    Fruit of 2 nutlets enveloped by the persistent calyx.
    [FZ]

    Flora Zambesiaca. Vol 8, Pt 7. Avicenniaceae, R. Fernandes. Nesogenaceae, M.A. Diniz. Verbenaceae, R. Fernandes. Lamiaceae, R. Fernandes. 2005.

    Habit
    Prostrate perennial herbs with woody taproots; indumentum of sparse to closely spaced appressed medifixed hairs or plants glabrous Prostrate perennial herbs with woody taproots; indumentum of sparse to closely spaced appressed medifixed hairs or plants glabrous.
    Stem
    Stems trailing, rooting at the nodes; branches short, ascending Stems trailing, rooting at the nodes; branches short, ascending.
    Leaves
    Leaves opposite, simple, shortly petiolate or subsessile, serrate Leaves opposite, simple, shortly petiolate or subsessile, serrate.
    Ovary
    Ovary 1-carpellate, carpel 2-locular, each locule 1-ovulate; style short, undivided; stigma thick, obliquely capitate
    Fruits
    Fruit a schizocarp, compressed oblong, separating at maturity into two 1-locular mericarps; mericarps with a dry hard pericarp, semi ovoid, flat on the commissural face, rounded and smooth on the dorsal face, glabrous. Fruit a schizocarp, compressed oblong, separating at maturity into two 1-locular mericarps; mericarps with a dry hard pericarp, semi ovoid, flat on the commissural face, rounded and smooth on the dorsal face, glabrous.
    Distribution
    A genus of c. 10 species in tropical and subtropical America, with 1 species widespread throughout the warmer parts of the world. A genus of c. 10 species in tropical and subtropical America, with 1 species widespread throughout the warmer parts of the world.
    Inflorescences
    Inflorescences axillary, pedunculate, spicate; spikes short, dense, ovoid becoming cylindric after anthesis, with bracts closely imbricate. Inflorescences axillary, pedunculate, spicate; spikes short, dense, ovoid becoming cylindric after anthesis, with bracts closely imbricate
    Flowers
    Flowers small, sessile, each solitary in the axil of a bract. Flowers small, sessile, each solitary in the axil of a bract
    Calyx
    Calyx membranous, dorsiventrally flattened, 2-lobed, persistent. Calyx membranous, dorsiventrally flattened, 2-lobed, persistent
    Corolla
    Corolla usually white, becoming rose-coloured; tube straight or curved, cylindric or widening toward the apex; limb oblique, c. 2-lipped, with the (upper) adaxial lip entire, emarginate or 2-lobed, and the lower (abaxial) lip 3-lobed with the median lobe usually larger than the others. Corolla usually white, becoming rose-coloured; tube straight or curved, cylindric or widening toward the apex; limb oblique, c. 2-lipped, with the (upper) adaxial lip entire, emarginate or 2-lobed, and the lower (abaxial) lip 3-lobed with the median lobe usually larger than the others
    Stamens
    Stamens 4, included or somewhat exserted. Stamens 4, included or somewhat exserted
    Pistil
    Ovary 1-carpellate, carpel 2-locular, each locule 1-ovulate; style short, undivided; stigma thick, obliquely capitate.

    Images

    Distribution

    Native to:

    Alabama, Albania, Aldabra, Algeria, Andaman Is., Angola, Argentina Northeast, Argentina Northwest, Argentina South, Arizona, Arkansas, Aruba, Assam, Bahamas, Baleares, Bangladesh, Belize, Benin, Bermuda, Bolivia, Botswana, Brazil North, Brazil Northeast, Brazil South, Brazil Southeast, Brazil West-Central, Burkina, Burundi, California, Cambodia, Cape Provinces, Caprivi Strip, Caroline Is., Cayman Is., Central African Repu, Chad, Chagos Archipelago, Chile Central, Chile North, China South-Central, China Southeast, Colombia, Colorado, Corse, Costa Rica, Cuba, Cyprus, District of Columbia, Djibouti, Dominican Republic, East Himalaya, Ecuador, Egypt, El Salvador, Eritrea, Ethiopia, Florida, Free State, French Guiana, Galápagos, Georgia, Ghana, Greece, Guatemala, Guinea-Bissau, Gulf States, Guyana, Hainan, Haiti, Honduras, Illinois, India, Indiana, Iowa, Iran, Italy, Jamaica, Japan, Kansas, Kentucky, Kenya, Kriti, KwaZulu-Natal, Laccadive Is., Lebanon-Syria, Leeward Is., Lesotho, Lesser Sunda Is., Liberia, Libya, Louisiana, Madagascar, Malawi, Malaya, Maldives, Mali, Marianas, Maryland, Mauritania, Mexico Central, Mexico Gulf, Mexico Northeast, Mexico Northwest, Mexico Southeast, Mexico Southwest, Michigan, Minnesota, Mississippi, Missouri, Morocco, Mozambique, Myanmar, Namibia, Nansei-shoto, Nebraska, Nepal, Netherlands Antilles, Nevada, New Guinea, New Jersey, New Mexico, New South Wales, New York, Nicaragua, Nicobar Is., Niger, Nigeria, North Carolina, Northern Provinces, Northern Territory, Ohio, Oklahoma, Oman, Ontario, Oregon, Pakistan, Panamá, Paraguay, Pennsylvania, Peru, Philippines, Puerto Rico, Queensland, Samoa, Saudi Arabia, Senegal, Seychelles, Sicilia, Sinai, Socotra, Solomon Is., Somalia, South Carolina, South China Sea, South Dakota, Southwest Caribbean, Spain, Sri Lanka, Sudan, Sulawesi, Suriname, Swaziland, Tanzania, Tennessee, Texas, Thailand, Tibet, Togo, Trinidad-Tobago, Tunisia, Turkey-in-Europe, Turks-Caicos Is., Uganda, Uruguay, Utah, Venezuela, Venezuelan Antilles, Virginia, West Himalaya, West Virginia, Western Australia, Windward Is., Wisconsin, Wyoming, Zambia, Zaïre, Zimbabwe

    Introduced into:

    Canary Is., Cocos (Keeling) Is., East Aegean Is., France, Great Britain, Hawaii, Mauritius, New Caledonia, Portugal, Sardegna, South Australia, St.Helena, Victoria

    Phyla Lour. appears in other Kew resources:

    First published in Fl. Cochinch.: 66 (1790)

    Accepted by

    • O'Leary, N. & Múlgura, M.E. (2011). A taxonomic revision of the genus Phyla (Verbenaceae) Annals of the Missouri Botanical Garden 98: 578-596.
    • Govaerts, R. (2003). World Checklist of Selected Plant Families Database in ACCESS: 1-216203. The Board of Trustees of the Royal Botanic Gardens, Kew.

    Literature

    Flora of West Tropical Africa
    • Greene in Pittonia 4: 48 (1899).
    • Fl. Cochinch. 66 (1790)
    Flora Zambesiaca
    • Atkins in Kadereit (ed.), Fam. Gen. Vasc. Pl. (Kubitzki, ed. in chief) VII: 462 (2004).
    • Kadereit (ed.), Fam. Gen. Vasc. Pl. (Kubitzki, ed. in chief) VII : 462 (2004).
    • Fl. Cochinch.: 66 (1790).
    Flora of Tropical East Africa
    • Greene in Pittonia 4: 45–48 (1899)
    • Fl. Cochinch.: 66 (1790)

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0