1. Family: Euphorbiaceae Juss.
    1. Genus: Acalypha L.
      1. Acalypha nyasica Hutch.

        This species is accepted, and its native range is S. Tanzania to Mozambique.

    [FTEA]

    Euphorbiaceae, A. R.-Smith. Flora of Tropical East Africa. 1987

    Habit
    A prostrate puberulous or pubescent annual herb with branches up to 0.5 m. long.
    Leaves
    Petioles 1–7 mm. long; leaf-blade ovate-lanceolate to lanceolate, (1–)3–6.7 cm. long, (0.4–)1–1.8 cm. wide, subacute or obtuse, cuneate or rounded, closely and rather indistinctly serrate or crenate-serrate, firmly membranous, 3(–5)-nerved at the base, lateral nerves 6–8 pairs, sparingly pubescent above and beneath at first, later becoming glabrous except along the midrib and main nerves, green above, paler beneath.
    Stipules
    Stipules subulate, 1–2 mm. long, sparingly pubescent.
    Inflorescences
    Male racemes solitary, axillary, 4–5 cm. long, long-pedunculate, sometimes replaced by allomorphic ♀ flowers which give rise to a clavate, 1-seeded fruit; bracts 0.5 mm. long. Female inflorescences spicate, solitary, terminal or subterminal, cylindric, 2–4(–6) cm. long, ± 1 cm. wide, subsessile or shortly pedunculate; bracts multifid, the segments linear, (3–)4–5 mm. long, pubescent, eglandular, slightly reddish tinged, 1-flowered.
    Flowers
    Male flowers shortly pedicellate; buds subglobose, 4-gonous, to 0.3 mm. diameter, subglabrous, reddish tinged. Female flowers sessile; sepals 3, triangular, 1 mm. long, acute, ciliate; ovary deeply trilobate, 0.5 mm. diameter, sparingly pubescent at the apex, otherwise ± glabrous; styles free, slender, filiform, 7–8 mm. long, simple, glabrous, red.
    Male
    Male flowers shortly pedicellate; buds subglobose, 4-gonous, to 0.3 mm. diameter, subglabrous, reddish tinged. Male racemes solitary, axillary, 4–5 cm. long, long-pedunculate, sometimes replaced by allomorphic ♀ flowers which give rise to a clavate, 1-seeded fruit; bracts 0.5 mm. long.
    Female
    Female flowers sessile; sepals 3, triangular, 1 mm. long, acute, ciliate; ovary deeply trilobate, 0.5 mm. diameter, sparingly pubescent at the apex, otherwise ± glabrous; styles free, slender, filiform, 7–8 mm. long, simple, glabrous, red. Female inflorescences spicate, solitary, terminal or subterminal, cylindric, 2–4(–6) cm. long, ± 1 cm. wide, subsessile or shortly pedunculate; bracts multifid, the segments linear, (3–)4–5 mm. long, pubescent, eglandular, slightly reddish tinged, 1-flowered.
    Fruits
    Fruits deeply trilobate, 1.5 mm. long, 2.5 mm. diameter, sparingly pubescent at the apex, otherwise glabrous.
    Seeds
    Seeds ovoid-subglobose, 1.3 mm. long, 1 mm. wide, smooth, brownish purple, with a small white velar caruncle perpendicular to the raphe.
    Habitat
    Deciduous woodland and thicket, often in open or rocky places; 240–900 m.
    Distribution
    T8
    [FZ]

    Euphorbiaceae, A. Radcliffe-Smith. Flora Zambesiaca 9:4. 1996

    Habit
    An annual herb with an erect lead shoot to c. 15 cm high and numerous prostrate, decumbent or ascending lateral shoots up to 30 cm long.
    Stem
    Stems puberulous or pubescent.
    Petiole
    Petioles 0.3–1(2) cm long.
    Leaf lamina
    Leaf blades 1–5 × 0.4–2.5 cm, ovate-lanceolate or ovate-trullate, subacute or obtuse at the apex, finely crenulate to subentire on the margin, cuneate or rounded at the base, membranous, sparingly puberulous or pubescent on both surfaces, especially on the midrib and main nerves, or else ± glabrous, 3–5-nerved from the base; lateral nerves in 4–6 pairs.
    Stipules
    Stipules c. 1 mm long, subulate.
    Inflorescences
    Male spikes axillary, up to 13 cm long, with a peduncle up to 9 cm long; bracts minute. Female spikes terminal, or apparently lateral when a lateral shoot arises from beneath one, 1–6 × 0.5–1 cm, cylindric or sometimes ovoid, subsessile or shortly pedunculate; bracts 7 × 8–9 mm, ± reniform, pubescent, eglandular, often reddish-tinged, 1-flowered, 16–20-toothed, the teeth c. 3 mm long, linear. Allomorphic female flowers solitary, axillary.
    Male
    Male spikes axillary, up to 13 cm long, with a peduncle up to 9 cm long; bracts minute. Male flowers sessile; buds subglobose, minute, sparingly pubescent to subglabrous, reddish; anthers creamy-white.
    Flowers
    Female flowers sessile; sepals 3, 1 mm long, triangular, acute, ciliate; ovary 0.5 mm in diameter, deeply 3-lobed, sparingly pubescent at the apex, otherwise ± glabrous; styles free, 7–8 mm long, slender, filiform, simple, glabrous, dark red. Allomorphic female flowers long-pedicellate, the pedicels up to 3 cm long, capillary. Male flowers sessile; buds subglobose, minute, sparingly pubescent to subglabrous, reddish; anthers creamy-white.
    Female
    Female flowers sessile; sepals 3, 1 mm long, triangular, acute, ciliate; ovary 0.5 mm in diameter, deeply 3-lobed, sparingly pubescent at the apex, otherwise ± glabrous; styles free, 7–8 mm long, slender, filiform, simple, glabrous, dark red. Allomorphic female flowers long-pedicellate, the pedicels up to 3 cm long, capillary. Female spikes terminal, or apparently lateral when a lateral shoot arises from beneath one, 1–6 × 0.5–1 cm, cylindric or sometimes ovoid, subsessile or shortly pedunculate; bracts 7 × 8–9 mm, ± reniform, pubescent, eglandular, often reddish-tinged, 1-flowered, 16–20-toothed, the teeth c. 3 mm long, linear. Allomorphic female flowers solitary, axillary.
    Fruits
    Fruits of the allomorphic flowers 2 × 1.5 mm, obovoid, tuberculate, minutely puberulous. Fruits 1.5 × 2.5 mm, deeply 3-lobed, sparingly pubescent at the apex, otherwise glabrous.
    Seeds
    Seeds 1.3 × 1 mm, ovoid-subglobose, smooth, pinkish-brown, with a small white velar caruncle.

    Distribution

    Native to:

    Malawi, Mozambique, Tanzania

    Acalypha nyasica Hutch. appears in other Kew resources:

    First published in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 6(1): 894 (1912)

    Accepted by

    • Govaerts, R., Frodin, D.G. & Radcliffe-Smith, A. (2000). World Checklist and Bibliography of Euphorbiaceae (and Pandaceae) 1-4: 1-1622. The Board of Trustees of the Royal Botanic Gardens, Kew.
    • Govaerts, R. (1995). World Checklist of Seed Plants 1(1, 2): 1-483, 1-529. MIM, Deurne.

    Literature

    Flora of Tropical East Africa
    • Pax in A. Engler, Das Pflanzenreich IV. 147(16): 70 (1924).
    • Hutch. in Flora of Tropical Africa 6(1): 894 (1912).

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0