1. Family: Euphorbiaceae Juss.
    1. Genus: Acalypha L.
      1. Acalypha wilkesiana Müll.Arg.

        This species is accepted, and its native range is Bismarck Archipelago to SW. Pacific.

    [FZ]

    Euphorbiaceae, A. Radcliffe-Smith. Flora Zambesiaca 9:4. 1996

    Habit
    A densely branched rounded shrub mostly 1–2 m tall, monoecious, tomentellous to pubescent.
    Petiole
    Petioles 1–5 cm long.
    Leaf lamina
    Leaf blades usually up to 15 × 10 cm, sometimes larger, elliptic-ovate to broadly ovate, obtusely acuminate at the apex, crenate-dentate on the margins, cuneate or rounded at the base, 5–7-nerved from the base, membranous, sparingly pubescent along the midrib and main veins on both surfaces at first, later glabrescent, green, copper or bronze and variously variegated with purple, red, pink, cream-coloured or white, or else pink-, cream- or white-margined; lateral nerves in 6–10 pairs.
    Stipules
    Stipules 7 mm long, narrowly lanceolate, acutely acuminate.
    Inflorescences
    Inflorescences axillary, usually solitary, spicate, unisexual, on the same or different shoots. Male spikes up to 12 cm long, densely but interruptedly flowered; bracts minute, many-flowered. Female spikes up to 7 cm long, lax flowered; bracts c. 5 × 4 mm, ± ovate, later accrescent, dentate, with 3–6 teeth on each side, sparingly pubescent to subglabrous, 1-flowered.
    Male
    Male spikes up to 12 cm long, densely but interruptedly flowered; bracts minute, many-flowered. Male flowers sessile or ± so; buds tetragonal, subglabrous, reddish; anthers yellowish.
    Flowers
    Female flowers sessile; sepals 3–4, 1 mm long, ovate, subacute; ovary 1.5 mm in diameter, subglobose, tomentose; styles c. 6 mm long, united at the base, deeply laciniate, red. Male flowers sessile or ± so; buds tetragonal, subglabrous, reddish; anthers yellowish.
    Female
    Female flowers sessile; sepals 3–4, 1 mm long, ovate, subacute; ovary 1.5 mm in diameter, subglobose, tomentose; styles c. 6 mm long, united at the base, deeply laciniate, red. Female spikes up to 7 cm long, lax flowered; bracts c. 5 × 4 mm, ± ovate, later accrescent, dentate, with 3–6 teeth on each side, sparingly pubescent to subglabrous, 1-flowered.
    Fruits
    Fruits 1.5–2 × 4 mm, 3-lobed, pubescent.
    Seeds
    Seeds not set in Africa.

    Images

    Distribution

    Native to:

    Bismarck Archipelago, Fiji, Solomon Is., Tuvalu, Vanuatu

    Introduced into:

    Bangladesh, Bolivia, Cook Is., Cuba, Dominican Republic, Ecuador, Florida, Gilbert Is., Guinea, Gulf of Guinea Is., Haiti, Hawaii, Jamaica, Leeward Is., Line Is., Madagascar, Marquesas, Mexico Central, Nicobar Is., Niue, Norfolk Is., Paraguay, Puerto Rico, Rwanda, Society Is., Thailand, Tonga, Trinidad-Tobago, Tuamotu, Tubuai Is., Venezuelan Antilles, Zaïre

    Acalypha wilkesiana Müll.Arg. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Wilkes Expedition [s.n.], Fiji K000959008
    Wilkes Expedition [s.n.], Fiji K000959009
    s.coll. [s.n.] Acalypha godseffiana K000959033
    s.coll. [s.n.], New Guinea Acalypha godseffiana K000959034 Unknown type material

    First published in A.P.de Candolle, Prodr. 15(2): 817 (1866)

    Accepted by

    • Flora of North America Editorial Committee (2016). Flora of North America North of Mexico 12: 1-603. Oxford University Press, New York, Oxford.
    • Cardiel, J.M. & Muñoz Rodríguez, P. (2015). Synopsis of Acalypha (Euphorbiaceae) of Argentina, Paraguay, and Uruguay Annals of the Missouri Botanical Garden 101: 384-405.
    • Jørgensen, P.M., Nee, M.H. & Beck., S.G. (eds.) (2013). Catálogo de las plantas vasculares de Bolivia Monographs in Systematic Botany from the Missouri Botanical Garden 127: 1-1741.
    • Mostaph, M.K. & Uddin, S.B. (2013). Dictionary of plant names of Bangladesh, Vasc. Pl.: 1-434. Janokalyan Prokashani, Chittagong, Bangladesh.
    • López Patiño, E.J., Szeszko, D.R., Rascala Pérez, J. & Beltrán Retis, A.S. (2012). The flora of the Tenacingo-Malinalco-Zumpahuacán protected natural area, state of Mexico, Mexico Harvard Papers in Botany 17: 65-167.
    • Cardiel, J.M. & Muñoz Rodríguez, P. (2012). Synopsis of Acalypha (Euphorbiaceae) of continental Ecuador PhytoKeys 17: 1-17.
    • Sagun, V.G., Levin, G.A. & van Welzen, P.C. (2010). Revision and phylogeny of Acalypha (Euphorbiaceae) in Malesia Blumea 55: 21-60.
    • Lejoy, J., Ndjele, M.-B. & Geerinck, D. (2010). Catalogue-flore des plantes vasculaires des districts de Kisangani et de la Tshopo (RD Congo) Taxonomania. Revue de Taxonomie et de Nomenclature Botaniques 30: 1-307.
    • Lisowski, S. (2009). Flore (Angiospermes) de la République de Guinée Scripta Botanica Belgica 41: 1-517.
    • Nelson Sutherland, C.H. (2008). Catálogo de las plantes vasculares de Honduras. Espermatofitas: 1-1576. SERNA/Guaymuras, Tegucigalpa, Honduras.
    • Chayamarit, K. & Van Welzen, P.C. (2005). Euphorbiaceae (Genera A-F) Flora of Thailand 8(1): 1-303. The Forest Herbarium, National Park, Wildlife and Plant Conservation Department, Bangkok.
    • Govaerts, R. (1995). World Checklist of Seed Plants 1(1, 2): 1-483, 1-529. MIM, Deurne.
    • Orchard, A.E. (ed.) (1994). Oceanic Islands 1 Flora of Australia 49: 1-681. Australian Government Publishing Service, Canberra.
    • Smith, A.C. (1981). Flora Vitiensis Nova. A new flora for Fiji (Spermatophytes only) 2: 1-810. Pacific Tropical Botanical Garden, Lawai.
    • Sykes, W.R. (1970). Contributions to the flora of Niue Bulletin, New Zealand Department of Scientific and Industrial Research 200: 1-321.

    Not accepted by

    • Balakrishnan, N.P. & Chakrabarty, T. (2007). The family Euphorbiaceae in India. A synopsis of its profile, taxonomy and bibliography: 1-500. Bishen Singh Mahendra Pal Singh, Dehra Dun. [Cited as Acalypha amentacea subsp. wilkesiana.]
    • Govaerts, R., Frodin, D.G. & Radcliffe-Smith, A. (2000). World Checklist and Bibliography of Euphorbiaceae (and Pandaceae) 1-4: 1-1622. The Board of Trustees of the Royal Botanic Gardens, Kew. [Cited as Acalypha amentacea subsp. wilkesiana.]
    • Welsh, S.L. (1998). Flora Societensis: 1-420. E.P.S. Inc. Utah. [Cited as Acalypha amentacea subsp. wilkesiana.]
    • Brako, L. & Zarucchi, J.L. (1993). Catalogue of the Flowering Plants and Gymnosperms of Peru Monographs in Systematic Botany from the Missouri Botanical Garden 45: i-xl, 1-1286. [Cited as Acalypha amentacea subsp. wilkesiana.]

    Literature

    Kew Backbone Distributions
    • Baksh-Comeau, Y., Maharaj, S.S., Adams, C.D., Harris, S.A., Filer, D.L. & Hawthorne, W.D. (2016). An annotated checklist of the vascular plants of Trinidad and Tobago with analysis of vegetation types and botanical 'hotspots' Phytotaxa 250: 1-431.
    • Flora of North America Editorial Committee (2016). Flora of North America North of Mexico 12: 1-603. Oxford University Press, New York, Oxford.
    • Cardiel, J.M. & Muñoz Rodríguez, P. (2015). Synopsis of Acalypha (Euphorbiaceae) of Argentina, Paraguay, and Uruguay Annals of the Missouri Botanical Garden 101: 384-405.
    • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
    • Jørgensen, P.M., Nee, M.H. & Beck., S.G. (eds.) (2013). Catálogo de las plantas vasculares de Bolivia Monographs in Systematic Botany from the Missouri Botanical Garden 127: 1-1741.
    • Mostaph, M.K. & Uddin, S.B. (2013). Dictionary of plant names of Bangladesh, Vasc. Pl.: 1-434. Janokalyan Prokashani, Chittagong, Bangladesh.
    • López Patiño, E.J., Szeszko, D.R., Rascala Pérez, J. & Beltrán Retis, A.S. (2012). The flora of the Tenacingo-Malinalco-Zumpahuacán protected natural area, state of Mexico, Mexico Harvard Papers in Botany 17: 65-167.
    • Cardiel, J.M. & Muñoz Rodríguez, P. (2012). Synopsis of Acalypha (Euphorbiaceae) of continental Ecuador PhytoKeys 17: 1-17.
    • Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies Smithsonian Contributions to Botany 98: 1-1192.
    • Figueiredo, E., Paiva, J., Stévart, T., Oliveira, F. & Smith, G.F. (2011). Annotated catalogue of the flowering plants of São Tomé and Príncipe Bothalia 41: 41-82.
    • Lejoy, J., Ndjele, M.-B. & Geerinck, D. (2010). Catalogue-flore des plantes vasculaires des districts de Kisangani et de la Tshopo (RD Congo) Taxonomania. Revue de Taxonomie et de Nomenclature Botaniques 30: 1-307.
    • Lisowski, S. (2009). Flore (Angiospermes) de la République de Guinée Scripta Botanica Belgica 41: 1-517.
    • Pandey, R.P. & Dilwakar, P.G. (2008). An integrated check-list flora of Andaman and Nicobar islands, India Journal of Economic and Taxonomic Botany 32: 403-500.
    • Chayamarit, K. & Van Welzen, P.C. (2005). Euphorbiaceae (Genera A-F) Flora of Thailand 8(1): 1-303. The Forest Herbarium, National Park, Wildlife and Plant Conservation Department, Bangkok.
    • Welsh, S.L. (1998). Flora Societensis: 1-420. E.P.S. Inc. Utah.
    • Orchard, A.E. (ed.) (1994). Oceanic Islands 1 Flora of Australia 49: 1-681. Australian Government Publishing Service, Canberra.
    • Fosberg, F.R. & Sachet, M.-H. (1987). Flora of the Gilbert Island, Kiribati, Checklist Atoll Research Bulletin 295: 1-33.
    • Wester, L. (1985). Checklist of the vasular plants of the Northern Line Islands Atoll Research Bulletin 287: 1-38.
    • Woodroffe, C.D. (1985). Vegetation and flora of Nui Atoll, Tuvalu Atoll Research Bulletin 283: 1-18.
    • Fosberg, F.R. (1975). Vascular plants of Aitutaki Atoll Research Bulletin 190: 73-84.
    • Sykes, W.R. (1970). Contributions to the flora of Niue Bulletin, New Zealand Department of Scientific and Industrial Research 200: 1-321.

    Sources

    Art and Illustrations in Digifolia
    Digital Image © Board of Trustees, RBG Kew

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0