1. Family: Rubiaceae Juss.
    1. Rubia L.

      1. This genus is accepted, and its native range is Old World.

    [FTEA]

    Rubiaceae, B. Verdcourt. Flora of Tropical East Africa. 1976

    Habit
    Herbs, the stems sometimes woody towards the base, often stiffly hairy or with tiny prickles
    Leaves
    Leaves in whorls of 4–6(–8), some of which may be stipular in origin; blades linear, lanceolate, obovate or less often cordate, often prickly
    Flowers
    Flowers small, in axillary and/or terminal cymes, hermaphrodite
    Calyx
    Calyx-tube ovoid or globose, the limb obsolete
    Corolla
    Corolla rotate or subcampanulate, (4–)5(–6)-merous
    Stamens
    Stamens inserted in the corolla-tube, the anthers exserted
    Disc
    Disc small, swollen
    Pistil
    Ovary 1–2-locular; ovules solitary in each locule, erect, affixed to the septum, amphitropous; style 2-fid or styles 2, short; stigmas capitate
    Fruits
    Fruit fleshy, of 2 parts or rarely 1-locular
    Seeds
    Seeds suberect, with plane or concave ventral face, horny endosperm and membranous testa, adhering to the pericarp.
    [FZ]

    Rubiaceae, B. Verdcourt. Flora Zambesiaca 5:1. 1989

    Habit
    Scrambling, creeping or climbing perennial herbs, somewhat woody near the base; roots quite woody, reddish (yielding a red dye).
    Stem
    Stems branched, 4-angled, ± brittle, mostly beset with recurved prickles.
    Leaves
    Leaves in whorls of (3)4–8(12), petiolate (in the Flora Zambesiaca area), blades cordate or broadly ovate, ovate-lanceolate to linear, rounded to ± cordate at base or narrowed to base, with 1–5(7) prominent veins.
    Flowers
    Flowers in axillary and/or terminal cymes, hermaphrodite, 5 (rarely 4 or 6) -merous.
    Calyx
    Calyx lobes obsolete.
    Corolla
    Corolla rotate to subcampanulate, greenish, yellowish green to yellow; lobes ± triangular, (long) acuminate.
    Anthers
    Anthers exserted.
    Pistil
    Ovary bilocular, one ovule in each chamber, crowned by a minute disk; style branches 2, joined below; stigmas capitate.
    Fruits
    Fruit fleshy, glabrous, consisting of 2 round mericarps, each with a single seed, round on dorsal side, plane to convex on ventral side; one mericarp often aborted.

    Images

    Distribution

    Native to:

    Afghanistan, Albania, Algeria, Altay, Amur, Angola, Assam, Azores, Baleares, Bangladesh, Borneo, Botswana, Burundi, Buryatiya, Canary Is., Cape Provinces, China North-Central, China South-Central, China Southeast, Chita, Corse, Cyprus, East Aegean Is., East European Russia, East Himalaya, Eritrea, Ethiopia, France, Free State, Great Britain, Greece, Hainan, India, Inner Mongolia, Iran, Iraq, Ireland, Italy, Japan, Jawa, Kazakhstan, Kenya, Khabarovsk, Kirgizstan, Korea, Kriti, Krym, Kuril Is., KwaZulu-Natal, Lebanon-Syria, Lesotho, Libya, Madeira, Malawi, Malaya, Manchuria, Mongolia, Morocco, Mozambique, Myanmar, Namibia, Nepal, North Caucasus, Northern Provinces, Oman, Pakistan, Palestine, Philippines, Portugal, Primorye, Qinghai, Rwanda, Sakhalin, Sardegna, Selvagens, Sicilia, Sinai, Somalia, South European Russi, Spain, Sri Lanka, Sudan, Sumatera, Swaziland, Tadzhikistan, Taiwan, Tanzania, Thailand, Tibet, Transcaucasus, Tunisia, Turkey, Turkey-in-Europe, Turkmenistan, Uganda, Ukraine, Uzbekistan, Vietnam, West Himalaya, West Siberia, Xinjiang, Yugoslavia, Zambia, Zaïre, Zimbabwe

    Introduced into:

    Argentina Northwest, Austria, Bulgaria, Chile Central, Czechoslovakia, Germany, Hungary, Netherlands, Switzerland

    Rubia L. appears in other Kew resources:

    First published in Sp. Pl.: 109 (1753)

    Accepted by

    • Govaerts, R. (2003). World Checklist of Selected Plant Families Database in ACCESS: 1-216203. The Board of Trustees of the Royal Botanic Gardens, Kew.

    Literature

    Flora Zambesiaca
    • Gen. Pl. ed. 5: 47 (1754).
    • Sp. Pl.1:109 (1753)
    Flora of Tropical East Africa
    • L., Gen. Pl., ed. 5: 47 (1754)
    • Sp. Pl.: 109 (1753)

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2018. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2018. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0