1. Family: Poaceae Barnhart
    1. Genus: Anthoxanthum L.
      1. Anthoxanthum odoratum L.

        This species is accepted, and its native range is Macaronesia, Europe to Mongolia, NW. Africa.

    [GB]
    Habit
    Perennial; caespitose. Butt sheaths papery; withering. Culms erect, or geniculately ascending; 10-75(-100) cm long; 1-3 -noded. Lateral branches lacking. Leaves mostly basal. Ligule an eciliate membrane; 1-5(-7) mm long. Leaf-blades 1-12(-30) cm long; 1.5-5(-9) mm wide; aromatic. Leaf-blade surface glabrous, or pubescent; hairy on both sides. Leaf-blade apex acuminate.
    Inflorescences
    Inflorescence a panicle. Panicle spiciform; oblong, or ovate; continuous, or interrupted; 1-12 cm long; 0.6-1.5 cm wide. Spikelets solitary. Fertile spikelets pedicelled. Pedicels oblong; 0.5-1 mm long; pubescent.
    Spikelets
    Spikelets comprising 2 basal sterile florets; 1 fertile florets; without rhachilla extension. Spikelets lanceolate; laterally compressed; 7-9.5 mm long; breaking up at maturity; disarticulating below each fertile floret.
    Fertile
    Spikelets comprising 2 basal sterile florets; 1 fertile florets; without rhachilla extension. Spikelets lanceolate; laterally compressed; 7-9.5 mm long; breaking up at maturity; disarticulating below each fertile floret.
    Glume
    Glumes persistent; dissimilar; exceeding apex of florets; thinner than fertile lemma; shiny. Lower glume ovate; 3.7-5.2 mm long; 0.5-0.6 length of upper glume; membranous; 1-keeled; 1 -veined. Lower glume primary vein smooth. Lower glume lateral veins absent. Lower glume surface pubescent. Lower glume apex acuminate. Upper glume elliptic; 7-9.5 mm long; 3-5 length of adjacent fertile lemma; membranous; with hyaline margins; 1-keeled; 3 -veined. Upper glume primary vein smooth. Upper glume surface pubescent. Upper glume apex acuminate.
    Florets
    Basal sterile florets dissimilar; barren; without significant palea; attached to and deciduous with the fertile. Lemma of lower sterile floret oblong; 2.8-3.6 mm long; 1.5-2 length of fertile lemma; membranous; without keels; 4-5 -veined; pilose; emarginate; awned. Awn of lower sterile floret dorsal; arising 0.6 way up back of lemma; 2-4 mm long. Lemma of upper sterile floret oblong; 3-3.5 mm long; 1 length of lower sterile floret; membranous; pilose; emarginate; awned. Awn of upper sterile floret dorsal; arising 0.1-0.2 way up back of lemma; geniculate; 6-9 mm long. Fertile lemma orbicular; 1.7-2.4 mm long; cartilaginous; shiny; without keel; 5 -veined. Lemma lateral veins obscure. Lemma margins convolute; covering most of palea. Lemma apex obtuse. Palea oblong; 1 length of lemma; 1 -veined; without keels.
    Flowers
    Lodicules absent. Anthers 2; (2.9-)3.5-4.8(-5.5) mm long. Stigmas 2; protogynous; pubescent.
    Fruits
    Caryopsis with adherent pericarp; ellipsoid; 2 mm long. Embryo 0.33 length of caryopsis. Hilum punctiform.
    Distribution
    Europe: northern, central, southwestern, southeastern, and eastern. Africa: north, Macaronesia, south, middle Atlantic ocean, and western Indian ocean. Asia-temperate: Siberia, Soviet far east, Soviet Middle Asia, Caucasus, western Asia, China, Mongolia, and eastern Asia. Asia-tropical: India. Australasia: Australia and New Zealand. Pacific: south-central and north-central. North America: Subarctic, western Canada, eastern Canada, northwest USA, north-central USA, northeast USA, southwest USA, southeast USA, and Mexico. South America: Mesoamericana, Caribbean, northern South America, western South America, Brazil, and southern South America. Antarctic: Subantarctic islands.
    Reference
    Aveneae. TAC.
    [KBu]

    Mashau A. C. 2016. A synopsis of Anthoxanthum (Poaceae: Pooideae: Poeae) in southern Africa and description of a new subspecies. Kew Bulletin 71:18. DOI 10.1007/S12225-016-9629-6

    Type
    Type: Europe. Sweet vernal grass.
    Habit
    Loosely or densely tufted perennial 300 – 600 (– 1000) mm high
    Leaves
    Leaf blade 150 – 300 × 2 – 8 mm, expanded, apex acute to acuminate
    Inflorescences
    Inflorescence spike-like, 10 – 90 mm long
    Spikelets
    Spikelet 7 – 10 mm long; glumes acute, hyaline, glabrous or minutely hairy on the back, keel scaberulous; lower glume up to 5 mm long, 1-nerved; upper glume 7 – 10 mm long, longer than and enfolding the spikelet; lower lemma 3.0 – 3.5 mm long, always dark brown, densely hairy; upper floret lemma 1.5 – 2.0 mm long, glabrous, awnless; anther 4.0 – 4.5 mm long.
    Distribution
    frequency in southern africa. locally common. naturalised from europe. introduced in north and south america, australia, tasmania and new zealand. south africa: kwazulu-natal and eastern cape.
    Conservation
    Not Evaluated.
    Note
    Flowering Oct. to Feb. Anthoxanthum odoratum resembles A. ecklonii, which has the lower lemma longer (5 mm long), lighter coloured and sparsely hairy.
    Ecology
    Humic soils in moist, swampy areas.
    [KBu]
    Use
    Introduced as a fodder grass to some countries. Fresh plant smells of coumarin.

    Images

    Distribution

    Native to:

    Albania, Algeria, Austria, Azores, Baleares, Baltic States, Belarus, Belgium, Bulgaria, Buryatiya, Canary Is., Central European Rus, Corse, Czechoslovakia, Denmark, East Aegean Is., East European Russia, Finland, France, Føroyar, Germany, Great Britain, Greece, Hungary, Iceland, Ireland, Italy, Kazakhstan, Kirgizstan, Kriti, Krym, Lebanon-Syria, Madeira, Mongolia, Morocco, Netherlands, North Caucasus, North European Russi, Northwest European R, Norway, Poland, Portugal, Romania, Sardegna, Sicilia, South European Russi, Spain, Sweden, Switzerland, Transcaucasus, Tunisia, Turkey, Turkey-in-Europe, Ukraine, West Siberia, Yugoslavia

    Introduced into:

    Alabama, Alaska, Aleutian Is., Amsterdam-St.Paul Is, Antipodean Is., Argentina Northeast, Argentina South, Arkansas, Assam, Bolivia, Brazil South, Brazil Southeast, British Columbia, California, Cape Provinces, Chatham Is., Chile Central, Chile South, China Southeast, Colombia, Colorado, Connecticut, Costa Rica, Delaware, District of Columbia, Dominican Republic, East Himalaya, Ecuador, Falkland Is., Georgia, Greenland, Haiti, Hawaii, Idaho, Illinois, Indiana, Jamaica, Japan, Juan Fernández Is., Kentucky, Kerguelen, Kermadec Is., Korea, Kuril Is., KwaZulu-Natal, Louisiana, Macquarie Is., Madagascar, Maine, Marquesas, Maryland, Masachusettes, Mexico Central, Mexico Northeast, Mexico Southeast, Mexico Southwest, Michigan, Mississippi, Missouri, New Brunswick, New Hampshire, New Jersey, New Mexico, New South Wales, New York, New Zealand North, New Zealand South, Newfoundland, Norfolk Is., North Carolina, Northern Provinces, Nova Scotia, Ohio, Ontario, Oregon, Pakistan, Pennsylvania, Peru, Prince Edward I., Queensland, Québec, Rhode I., Réunion, Sakhalin, South Australia, South Carolina, South Georgia, Sri Lanka, St.Helena, Taiwan, Tasmania, Tennessee, Texas, Tristan da Cunha, Uruguay, Venezuela, Vermont, Victoria, Virginia, Washington, West Himalaya, West Virginia, Western Australia, Wisconsin

    Anthoxanthum odoratum L. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Eiten, G. [6630], Brazil K001101038
    Eiten, G. [6622], Brazil K001101039
    Castellanos, A. [s.n.], Brazil K001101040
    Baron, R. [2050], Madagascar K000345008
    Baron, R. [2050], Madagascar K000345009
    Baron, R. [4098], Madagascar K000345010
    Reverchon, E. [403], Corse K000838020
    Cope, T.A. [RBG 43], United Kingdom K000914532
    Carauta, J.P.P. [525], Brazil K001101044
    Fay, M.F. [MFF204], United Kingdom K000696478
    Dombrowski, L.T. [2019], Brazil K001101036
    Glaziou, A.F.M. [16618] K001101042
    Glaziou, A. [16618], Brazil K001101041
    Glaziou, A. [20093], Brazil K001101043
    Love, A. [373], Sweden K000838021 isotype
    Grazziotin, G. [s.n.], Brazil K001101035
    Grazziotin, G. [s.n.], Brazil K001101037

    First published in Sp. Pl.: 28 (1753)

    Accepted by

    • Mashau, A.C. (2016). A synopsis of Anthoxanthum (Poaceae: Pooideae: Poeae) in southern Africa and description of a new subspecies Kew Bulletin 71(18): 1-5.
    • Chang, C.S., Kim, H. & Chang, K.S. (2014). Provisional checklist of vascular plants for the Korea peninsula flora (KPF): 1-660. DESIGNPOST.
    • Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies Smithsonian Contributions to Botany 98: 1-1192.
    • Snow, N. & Davidse, G. (2011). Notes on grasses (Poaceae) in Hawai'i: 3 Bishop Museum Occasional Papers 110: 17-22.
    • Giraldo-Cañas, D. (2011). Catálogo de la familia Poaceae en Colombia Darwiniana 49: 139-247.
    • Meyer, F.K. (2011). Beiträge zur Flora von Albanien Haussknechtia, Beih. 15: 1-220.
    • Edgar, E & Connor, H.E. (2010). Flora of New Zealand, ed. 2, 5: 1-650. R.E.Owen, Government Printer, Wellington.
    • Dobignard, D. & Chatelain, C. (2010). Index synonymique de la flore d'Afrique du nord 1: 1-455. Éditions des conservatoire et jardin botaniques, Genève.
    • Clayton, W.D. & Snow, N. (2010). A key to Pacific Grasses: 1-107. Kew Publishing, Royal Botanic Gardens, Kew.
    • Wilson, A. (ed.) (2009). Flora of Australia 44A: 1-410. Australian Government Publishing Service, Canberra.
    • Kandwal, M.K. & Gupta, B.K. (2009). An update on grass flora of Uttarkhand Indian Journal of Forestry 32: 657-668.
    • Gremmen, N. & Halbertsma, R.L. (2009). Alien plants and their impact on Tristan da Cunha 2: 1-307. Overseas Territories Environment Programme (OTEP).
    • Zuloaga, F.O., Morrone, O. , Belgrano, M.J., Marticorena, C. & Marchesi, E. (eds.) (2008). Catálogo de las Plantas Vasculares del Cono Sur Monographs in Systematic Botany from the Missouri Botanical Garden 107: 1-3348.
    • Grubov, V.I. (2008). Key to the vascular plants of Mongolia (with an atlas) 2: 1-503. Academy of Sciences, Mongolian People's Republic, Ulaan Bator.
    • Hokche, O., Berry, P.E. & Huber, O. (eds.) (2008). Nuevo Catálogo de la Flora Vascular de Venezuela: 1-859. Fundación Instituto Botánico de Venezuela.
    • Flora of North America Editorial Committee (2007). Flora of North America North of Mexico 24: 1-908. Oxford University Press, New York, Oxford.
    • Wu, Z. & Raven, P.H. (eds.) (2006). Poaceae Flora of China 22: 1-733. Missouri Botanical Garden Press, St. Louis.
    • Clayton, W.D., Harman, K.T. & Williamson, H. (2006). World Grass Species - Synonymy database The Board of Trustees of the Royal Botanic Gardens, Kew.
    • Dávila, P., Mejia-Saulés, M.T., Gómez-Sánchez, N., Valdés-Reyna, J., Ortíz, J.J., Morín, C., Castrejón, J. & Ocampo, A. (2006). Catálogo de las Gramíneas de México: 1-671. CONABIO, México city.
    • Takhtajan, A.L. (ed.) (2006). Conspectus Florae Caucasi 2: 1-466. Editio Universitatis Petropolitanae.
    • Germishuizen, G. & Meyer, N.L. (eds.) (2003). Plants of Southern Africa: an annotated checklist. Strelitzia 14.: i-vi, 1-1231. National Botanical Institute, Pretoria.
    • Broughton, D.A. & McAdam, J.H. (2002). The non-native vascular flora of the Falkland islands Botanical Journal of Scotland 54: 153-190.
    • Malyschev, L.I. & Peschkova, G.A. (eds.) (2001). Flora of Siberia 2: 1-362. Scientific Publishers, Inc., Enfield, Plymouth.
    • Noltie, H.J. (2000). Flora of Bhutan 3(2): 457-883. Royal Botanic Gardens, Edinburgh.
    • Fedorov, A.A. (ed.) (1999). Flora of Russia. The European part and bordering regions 1: 1-546. A.A. Balkema, Rotterdam, Broekfield.
    • Jørgensen, P.M. & León-Yánez, S. (eds.) (1999). Catalogue of the Vascular Plants of Ecuador Monographs in Systematic Botany from the Missouri Botanical Garden 75: i-viii, 1-1181.
    • Renvoize, S.A. (1998). Gramineas de Bolivia: 1-644. The Royal Botanic Gardens, Kew.
    • Davidse, G. & al. (eds.) (1994). Flora Mesoamericana 6: 1-543. Universidad Nacional Autónoma de México, México, D.F.
    • Orchard, A.E. (ed.) (1994). Oceanic Islands 1 Flora of Australia 49: 1-681. Australian Government Publishing Service, Canberra.
    • Dassanayake (ed.) (1994). A Revised Handbook to the Flora of Ceylon 8: 1-458. Oxford & IBH Publishing Co. PVT. LTD., New Delhi, Calcutta.
    • George, A.S., Orchard, A.E. & Hewson, H.J. (eds.) (1993). Oceanic islands 2 Flora of Australia 50: 1-606. Australian Government Publishing Service, Canberra.
    • Brako, L. & Zarucchi, J.L. (1993). Catalogue of the Flowering Plants and Gymnosperms of Peru Monographs in Systematic Botany from the Missouri Botanical Garden 45: i-xl, 1-1286.
    • Karthikeyan, S., Jain, S.K., Nayar, M.P. & Sanjappa, M. (1989). Florae Indicae Enumeratio: Monocotyledonae: 1-435. Botanical Survey of India, Calcutta.
    • Koyama, T. (1987). Grasses of Japan and its neighboring regions: an identification manual: 1-570. Kodansha, Tokyo, Japan.
    • Davis, P.H. (ed.) (1985). Flora of Turkey and the East Aegean Islands 9: 1-724. Edinburgh University Press, Edinburgh.
    • Kharkevich, S.S., Probatova, N.S. & Novikov, V.S. (1985). Sosudistye rasteniia sovetskogo Dal’nego Vostoka 1: 1-383. Izd-vo "Nauka," Leningradskoe otd-nie, Leningrad.
    • Cope, T.A. (1982). Flora of Pakistan 143: 1-678. Department of Botany, University of Karachi, Karachi.
    • Tutin, T.G. & al. (eds.) (1980). Flora Europaea 5: 1-452. Cambridge University Press.
    • Bor, N.L. (1970). Flora Iranica 70: 1-573. Naturhistorisches Museums Wien.
    • Bosser, J. (1969). Gramiées des pasturages et des cultures a Madagascar: 1-440. ORSTOM, Paris.

    Literature

    Kew Bulletin
    • Victor, J. E., Fish, L. & Ellis, R. P. (2007). Anthoxanthum brevifolium Stapf. National Assessment: Red List of South African Plants version 2014.1. Accessed on 17 March 2015.Google Scholar
    • Mucina, L. & Rutherford, M. C. (2006). The vegetation of South Africa, Lesotho and Swaziland. Strelitzia 19.Google Scholar
    • Fish, L. & Victor, J. E. (2005). Anthoxanthum dregeanum (Nees) Stapf. National Assessment: Red List of South African Plants version 2014.1. Accessed on 17 March 2015.Google Scholar
    • Fish, L. (2000). Poaceae (Gramineae). In: O. A. Leistner, Seeds plants of southern Africa: families and genera, Strelitzia 10: 659 – 726.Google Scholar
    • Gibbs Russell, G. E., Watson, L., Koekemoer, M., Smook, L., Barker, N. P., Anderson, H. M. & Dallwitz, M. J. (1990). Grasses of southern Africa, Mem. Bot. Surv. South Africa 58.Google Scholar
    • Stapf, O. (1910). Diagnoses africanae: XXXIV. Bull. Misc. Inform., Kew 1910(2): 59.Google Scholar
    • Stapf, O. (1899). Gramineae. Flora capensis 7: 465 – 468. Lovell Reeve, London.Google Scholar
    • Nees ab Esenbeck, C. G. D. (1841). Florae Africanae australioris illustrationes monographicae. Prausnitz, Glogau.Google Scholar
    • Trinius, C. B. (1839). Phalaridea. Mém. Acad. Imp. Sci. Saint-Pétersbourg, Sér. 6, Sci. Math., Seconde Pt. Sci. Nat. 5, 3 (3): 78 – 84.Google Scholar
    • Linnaeus 1753: 28
    • Linnaeus, C. (1753). Species plantarum. Laurentius Salvius, Stockholm.Google Scholar
    Kew Backbone Distributions
    • Chang, C.S., Kim, H. & Chang, K.S. (2014). Provisional checklist of vascular plants for the Korea peninsula flora (KPF): 1-660. DESIGNPOST.
    • Forzza, R.C. & al. (2013). Lista de Espécies da Flora do Brasil http://floradobrasil.jbrj.gov.br/2010/.
    • Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies Smithsonian Contributions to Botany 98: 1-1192.
    • Marhold, K. (ed.) (2012). IAPT/IOPB chromosome data 13 Taxon 61: 889-902.
    • Hassler, M. (2012). Flora of Rhodes. Systematic list of flora of Rhodes http://www.rz.uni-karlsruhe.de/~db111/flora/rhodos/list.php.
    • Giraldo-Cañas, D. (2011). Catálogo de la familia Poaceae en Colombia Darwiniana 49: 139-247.
    • Edgar, E & Connor, H.E. (2010). Flora of New Zealand, ed. 2, 5: 1-650. R.E.Owen, Government Printer, Wellington.
    • Dobignard, D. & Chatelain, C. (2010). Index synonymique de la flore d'Afrique du nord 1: 1-455. Éditions des conservatoire et jardin botaniques, Genève.
    • Clayton, W.D. & Snow, N. (2010). A key to Pacific Grasses: 1-107. Kew Publishing, Royal Botanic Gardens, Kew.
    • Wilson, A. (ed.) (2009). Flora of Australia 44A: 1-410. Australian Government Publishing Service, Canberra.
    • Kandwal, M.K. & Gupta, B.K. (2009). An update on grass flora of Uttarkhand Indian Journal of Forestry 32: 657-668.
    • Gremmen, N. & Halbertsma, R.L. (2009). Alien plants and their impact on Tristan da Cunha 2: 1-307. Overseas Territories Environment Programme (OTEP).
    • Zuloaga, F.O., Morrone, O. , Belgrano, M.J., Marticorena, C. & Marchesi, E. (eds.) (2008). Catálogo de las Plantas Vasculares del Cono Sur Monographs in Systematic Botany from the Missouri Botanical Garden 107: 1-3348.
    • Grubov, V.I. (2008). Key to the vascular plants of Mongolia (with an atlas) 2: 1-503. Academy of Sciences, Mongolian People's Republic, Ulaan Bator.
    • Hokche, O., Berry, P.E. & Huber, O. (eds.) (2008). Nuevo Catálogo de la Flora Vascular de Venezuela: 1-859. Fundación Instituto Botánico de Venezuela.
    • Flora of North America Editorial Committee (2007). Flora of North America North of Mexico 24: 1-908. Oxford University Press, New York, Oxford.
    • Wu, Z. & Raven, P.H. (eds.) (2006). Poaceae Flora of China 22: 1-733. Missouri Botanical Garden Press, St. Louis.
    • Dávila, P., Mejia-Saulés, M.T., Gómez-Sánchez, N., Valdés-Reyna, J., Ortíz, J.J., Morín, C., Castrejón, J. & Ocampo, A. (2006). Catálogo de las Gramíneas de México: 1-671. CONABIO, México city.
    • Takhtajan, A.L. (ed.) (2006). Conspectus Florae Caucasi 2: 1-466. Editio Universitatis Petropolitanae.
    • Frenot, Y., Chown, S.L., Whinam, J., Selkirk, P.M., Convey, P., Skotnicki, M. & Bergstrom,D.M. (2005). Biological invasions in the Antarctic: extent, impacts and implications Biological Reviews of the Cambridge Philosophical Society 80: 45-72.
    • Germishuizen, G. & Meyer, N.L. (eds.) (2003). Plants of Southern Africa: an annotated checklist. Strelitzia 14.: i-vi, 1-1231. National Botanical Institute, Pretoria.
    • Broughton, D.A. & McAdam, J.H. (2002). The non-native vascular flora of the Falkland islands Botanical Journal of Scotland 54: 153-190.
    • Malyschev, L.I. & Peschkova, G.A. (eds.) (2001). Flora of Siberia 2: 1-362. Scientific Publishers, Inc., Enfield, Plymouth.
    • Noltie, H.J. (2000). Flora of Bhutan 3(2): 457-883. Royal Botanic Gardens, Edinburgh.
    • Fedorov, A.A. (ed.) (1999). Flora of Russia. The European part and bordering regions 1: 1-546. A.A. Balkema, Rotterdam, Broekfield.
    • Jørgensen, P.M. & León-Yánez, S. (eds.) (1999). Catalogue of the Vascular Plants of Ecuador Monographs in Systematic Botany from the Missouri Botanical Garden 75: i-viii, 1-1181.
    • Renvoize, S.A. (1998). Gramineas de Bolivia: 1-644. The Royal Botanic Gardens, Kew.
    • Davidse, G. & al. (eds.) (1994). Flora Mesoamericana 6: 1-543. Universidad Nacional Autónoma de México, México, D.F.
    • Orchard, A.E. (ed.) (1994). Oceanic Islands 1 Flora of Australia 49: 1-681. Australian Government Publishing Service, Canberra.
    • Dassanayake (ed.) (1994). A Revised Handbook to the Flora of Ceylon 8: 1-458. Oxford & IBH Publishing Co. PVT. LTD., New Delhi, Calcutta.
    • George, A.S., Orchard, A.E. & Hewson, H.J. (eds.) (1993). Oceanic islands 2 Flora of Australia 50: 1-606. Australian Government Publishing Service, Canberra.
    • Brako, L. & Zarucchi, J.L. (1993). Catalogue of the Flowering Plants and Gymnosperms of Peru Monographs in Systematic Botany from the Missouri Botanical Garden 45: i-xl, 1-1286.
    • Karthikeyan, S., Jain, S.K., Nayar, M.P. & Sanjappa, M. (1989). Florae Indicae Enumeratio: Monocotyledonae: 1-435. Botanical Survey of India, Calcutta.
    • Koyama, T. (1987). Grasses of Japan and its neighboring regions: an identification manual: 1-570. Kodansha, Tokyo, Japan.
    • Davis, P.H. (ed.) (1985). Flora of Turkey and the East Aegean Islands 9: 1-724. Edinburgh University Press, Edinburgh.
    • Kharkevich, S.S., Probatova, N.S. & Novikov, V.S. (1985). Sosudistye rasteniia sovetskogo Dal’nego Vostoka 1: 1-383. Izd-vo "Nauka," Leningradskoe otd-nie, Leningrad.
    • Brown, L.C. (1982). The Flora and Fauna of St Helena: 1-88. Land Resources Development Centre, Surbiton, England.
    • Tutin, T.G. & al. (eds.) (1980). Flora Europaea 5: 1-452. Cambridge University Press.
    • Bor, N.L. (1970). Flora Iranica 70: 1-573. Naturhistorisches Museums Wien.
    • Bosser, J. (1969). Gramiées des pasturages et des cultures a Madagascar: 1-440. ORSTOM, Paris.
    • Mouterde, P. (1966). Nouvelle Flore du Liban et de la Syrie 1(Texte): 1-563. Éditions de l'imprimerie Catholique, Beyrouth.
    • Hultén, E.O.G. (1960). Flora of the Aleutian Islands and westernmost Alaska Peninsula: with notes on the flora of Commander Islands, ed. 2: 1-376. Weinheim : J. Cramer ; New York : Hafner Pub. Co.
    • Pavlov, N.V. (ed.) (1956). Flora Kazakhstana 1: 1-354. Alma-Ata, Izd-vo Akademii nauk Kazakhskoi SSR.
    • Roshevitz, R.J. & al. (eds.) (1950). Flora Kirgizskoi SSR 2: 1-315. Frunze : Izd-vo KirgizFAN SSSR.
    • de Cordemoy, E.J. (1895). Flore de l'ile de la Réunion: 1-574. Librairie des sciences naturelles Paul Klincksieck, Paris.

    Sources

    GrassBase - The Online World Grass Flora
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2018. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Bulletin
    Kew Bulletin
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2018. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0