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  1. Family: Lamiaceae Martinov
    1. Genus: Stachys L.
      1. Stachys pseudonigricans Gürke

        This species is accepted, and its native range is Tanzania to S. Tropical Africa.

    [FTEA]

    Lamiaceae (Labiatae), A.J. Paton, G. Bramley, O. Ryding, R.M. Polhill, Y.B. Harvey, M. Iwarsson, F. Willis, P.B. Phillipson, K. Balkwill, C.W. Lukhoba, D.F. Otieno, & R.M. Harley. Stachys, Y.B. Harvey. Flora of Tropical East Africa. 2009

    Type
    Type: Malawi, Shire Highlands, Buchanan 28 (K!, holo.)
    Habit
    Perennial, faintly aromatic, erect or procumbent, lax or bushy herb, 0.1–0.8(–1) m tall
    Stem
    Stems quadrangular to rounded, branching throughout length, densely to laxly covered with a combination of dendroid, stellate and/or biramous (and occasionally simple) hairs
    Leaves
    Leaves linear-lanceolate to broadly obvate, all ± similar or narrower above, (1.2–)2.4–8(–13) × (0.1–)0.5–3.8(–4.5) cm, crenate to laxly toothed, apex acute to acuminate or rounded, base usually rounded to truncate, sometimes attenuate or cordate, laxly to densely pubescent, upper surface with mostly biramous, stellate and/or dendroid hairs, lower surface with mostly biramous, stellate and/or dendroid hairs, some simple hairs sometimes present; sessile or petiolate, petiole 1–10 mm long, laxly to densely pubescent with biramous, stellate and/or dendroid hairs
    Inflorescences
    Inflorescence lax, with (4–)6–8(–16)-flowers per verticil, verticils < 5–70(–105) mm apart; bracts similar to cauline leaves, 4–90 × 1–28 mm, pubescence as of cauline leaves; bracteoles narrowly linear, 1–8(–12) mm long, pubescence of simple, biramous, stellate and/or dendroid hairs; pedicels (0.5–)1–4 mm long, pubescence of simple, biramous, stellate and/or dendroid hairs
    Calyx
    Calyx ascending, funnel-shaped to campanulate, 5.5–13 mm long at anthesis, extending to 16 mm long in fruit, outer surface pubescent, with simple, biramous, stellate and/or dendroid hairs, glabrous within, with or without long simple hairs between lobes, broadening in fruit; teeth triangular, sometimes narrowly so, (1.5–)2–4(–7) mm long, acuminate, erect to sometimes recurved
    Corolla
    Corolla white to pink to pale lavender or mauve, with or without darker flecks, 9–16 mm long; tube (5–)6–10 mm long, ± shorter than the calyx, outer surface ± glabrous, and pubescent at apex; anterior lip 3.5–8 mm long, outer surface pubescent, with simple, biramous, stellate and/or dendroid hairs, glabrous within; posterior lip 2–5(–6) mm long, outer surface densely pubescent with simple, biramous, stellate and/or dendroid hairs
    Fruits
    Nutlets dark brown, broadly ovoid, 1.5–3 mm long, minutely tuberculate, with pubescent apex when immature, glabrous when mature.
    Ecology
    Grassland, bushland, marshes and Brachystegia woodland in rich loam soil; 950–2150 m
    Conservation
    Least concern; widely distributed,.
    Note
    The remaining group tends to have much narrower leaves and indumentum which lacks dendroid hairs. There are two extremes, one with tall plants with very narrow leaves and another with shorter plants with narrow leaves, and distinctly long bracts at the apices of the branches. Intermediates linking these two extremes with the type form have both narrower leaves and slightly broader lanceolate leaves. There is also a much larger intermediate with less of a difference between the leaf types. The taller of the extremes is known from Tanzania (Rungwe and Mbeya District). The intermediate with pronounced leaf differences is known from Tanzania (Mbeya District) and Malawi (Chitipa and Rungwe Districts). The other intermediate is known from Tanzania (Mpanda, Mbeya, Ufipa and Songea Districts), Malawi (Nyika District) and Zambia (Chati, Ndola and Kitwe Districts). These biramous and stellate plants also show similar morphologies to the dendroid haired plants with both smaller ovate and narrower leaves in the second group detailed above. The second group with dendroid hairs are large plants with ovate-lanceolate leaves. Intermediates between this and the type form are still as large but have more lanceolate leaves. The extreme form is known from Tanzania (Mufindi, Iringa and Ufipa District) and the intermediate form from Tanzania (Mbeya, Chunya and Ufipa District) and Zambia (Mongu, Mazabuka, Kalambo Falls and Chilongowelo). The final dendroid-haired extreme is smaller than the type form and has very densely pubescent ovate leaves. Intermediates between this and the type are altogether larger, and have a laxer pubescence or have both ovate in addition to narrowly ovate-lanceolate leaves. The extreme form is known form Tanzania (Iringa and Ufipa Districts). The larger ovate intermediate is known from Mozambique (Niassa and an unknown locality) and Zambia (Kasama and Isoka Districts). The intermediate with two leaf types is known from Tanzania (Iringa District and Songea District), Mozambique (Niassa), Zambia (Mbala District) and Malawi (Nkhata Bay District). The type form has lanceolate leaves (24–80 × 5–38 mm) throughout the plant that are laxly pubescent with (simple to) biramous, stellate and/or dendroid hairs on the upper surface and biramous, stellate and/or dendroid hairs on the lower surface, and has an acute apex and a rounded to truncate base. It has lax dendroid, dendroid and stellate, and occasionally stellate pubescence on its stems. The material is mostly petiolate (1–10 mm long) but is occasionally sessile; has 6–12 flowers per verticil; bracteoles 1–8 mm long; corolla 9–16 mm long and white to pink to pale lavender/mauve in colour (with or without darker flecks). The type form is known from Malawi, Zimbabwe, Tanzania (Mbeya District, Iringa-Songea District, Njombe District, Ludewa District) and Zambia. The first of the extremes has narrow lanceolate leaves densely covered with stellate and dendroid hairs. The leaves can be as narrow as 6 mm wide. Intermediates between this and the type form have broader leaves, with a laxer pubescence. The extreme form is known from Tanzania (Ufipa District) and the intermediates from Zambia (Mbala and Mpika Districts). The Southern Highlands in Tanzania appears to be the centre for morphological diversity of this species, all extreme forms are known from only this region. Intermediates and the type form occur in this region in addition to other areas. The type form has (simple to) biramous, stellate and/or dendroid hairs. For this reason the last group cannot justifiably be separated from the other forms for lacking dendroid hairs. All extremes have intermediates linking them to the type form and so also cannot justifiably be placed into separate taxa. Stachys pseudonigricans is a polymorphic species comprised of a “central” type form, from which four morphological extremes radiate. Each of the extremes has intermediates linking them to the type. The type form and three of the groups are petiolate and have dendroid (and stellate) hairs. The remaining group has biramous and stellate hairs and has ± sessile leaves.
    Distribution
    Range: Zambia, Malawi, Mozambique and Zimbabwe Flora districts: T4 T7 T8

    Images

    Distribution

    Native to:

    Angola, Malawi, Mozambique, Tanzania, Zambia, Zimbabwe

    Other Data

    Stachys pseudonigricans Gürke appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Sep 1, 1995 Buchanan, J. [28] K000193108 Unknown type material
    Sep 1, 1995 Buchanan, J. [1], Malawi K000414654
    Gossweiler, J. [9985], Angola K001008811
    Henriques, C. [90], Angola K001008813
    Barbosa, G. [9684], Angola K001008812

    Bibliography

    First published in Bot. Jahrb. Syst. 28: 315 (1900)

    Accepted by

    • Paton, A.J., Bramley, G., Ryding, O., Polhill, R.M., Harvey, Y.B., Iwarsson, M., Otieno, D., Balkwill, K., Phillipson, P.B., Harley, R.M. & Willis, F. (2013). Flora Zambesiaca 8(8): 1-346. Royal Botanic Gardens, Kew.
    • Paton, A.J., Bramley, G., Ryding, O., Polhill, R., Harvey, Y., Iwarsson, M., Willis, F., Phillipson, P., Balkwill, K., Lukhoba, C., Otiend, D & Harley (2009). Lamiaceae (Labiatae) Flora of Tropical East Africa: 1-430.
    • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
    • Krestovskaya, T.V. (2005). On species of the genus Stachys (Lamiaceae) section Ambleia from Africa Botanicheskii Zhurnal. Moscow & Leningrad 90: 1445-1462.
    • Govaerts, R. (2003). World Checklist of Selected Plant Families Database in ACCESS: 1-216203. The Board of Trustees of the Royal Botanic Gardens, Kew.

    Literature

    Kew Backbone Distributions

    • Paton, A.J., Bramley, G., Ryding, O., Polhill, R.M., Harvey, Y.B., Iwarsson, M., Otieno, D., Balkwill, K., Phillipson, P.B., Harley, R.M. & Willis, F. (2013). Flora Zambesiaca 8(8): 1-346. Royal Botanic Gardens, Kew.
    • Paton, A.J., Bramley, G., Ryding, O., Polhill, R., Harvey, Y., Iwarsson, M., Willis, F., Phillipson, P., Balkwill, K., Lukhoba, C., Otiend, D & Harley (2009). Lamiaceae (Labiatae) Flora of Tropical East Africa: 1-430.
    • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
    • Krestovskaya, T.V. (2005). On species of the genus Stachys (Lamiaceae) section Ambleia from Africa Botanicheskii Zhurnal. Moscow & Leningrad 90: 1445-1462.

    Flora of Tropical East Africa

    • Harley & Reynolds (eds), Adv. Lab. Sci.: 149–161 (1992).
    • E.J. 28: 315 (1901)

    Sources

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2020. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2020. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0