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  1. Family: Lentibulariaceae Rich.
    1. Genus: Utricularia L.
      1. Utricularia reflexa Oliv.

        This species is accepted, and its native range is Tropical Africa to N. Botswana, Central Madagascar.

    [FTEA]

    Lentibulariaceae, P. Taylor. Flora of Tropical East Africa. 1973

    Habit
    Aquatic herb.
    Stolons
    Stolons filiform to relatively thick and fleshy, up to 50 cm. long or more, 0.3–3.0 mm. thick, glabrous or ± densely covered with simple or stellate hairs which extend also to the traps, basal part of the leaves and scapes; internodes 2–10(–15) mm.
    Rhizoids
    Rhizoids usually absent but when present 2–3 from the scape base, capillary, 3–5 mm. long.
    Leaves
    Leaves very numerous, digitately divided to the base into 2–5 primary segments, 3–30 mm. long; primary segments ovate in outline, repeatedly dichotomously divided; ultimate segments capillary, setulose.
    Traps
    Traps inserted in the angles between the primary segments and between subsequent dichotomies, very variable in number and size, ovoid, shortly stalked, 1–6 mm. long; mouth lateral; upper lip with 2 simple to copiously branched hairs; lower lip naked or with shorter simple hairs.
    Inflorescences
    Inflorescence lateral, 1.5–18 cm. high; flowers 1–3(4), distant, the lowermost sometimes very near the base of the scape; scape erect or rarely twining, filiform; scales absent; bracts basifixed, quadrate or orbicular, ± encircling the base of the pedicel, 1–3 mm. long; bracteoles absent; pedicels filiform, erect at anthesis, strongly reflexed or recurved in fruit, 2–35 mm. long.
    Calyx
    Calyx-lobes subequal, broadly ovate, 1–2 mm. long, scarcely accrescent, apex obtuse or rounded.
    Corolla
    Corolla pale to quite deep yellow, usually with brown or reddish nerves, ± densely covered externally with fine short hairs, rarely glabrous, 3–15 mm. long; upper lip broadly ovate or orbicular, apex truncate or rounded, entire or emarginate, 1.5–3 times as long as the upper calyx-lobe; lower lip orbicular to subreniform, apex ± emarginate or 2-lobed; palate raised, bigibbous; spur cylindrical, ± as long as the lower lip.
    Stamens
    Filaments linear; anther-thecae ± confluent.
    Pistil
    Ovary globose; style short or obsolete; stigma lower lip semi-orbicular, sometimes ciliate, upper very short or obsolete.
    Fruits
    Capsule globose, circumscissile, 3–4 mm. in diameter, few–many-seeded.
    Seeds
    Seeds lenticular, angular, 0.4–0.8 mm. wide, often narrowly winged on the angles; testa reticulate, cells distinct, isodiametric or ± elongate.
    [FWTA]

    Lentibulariaceae, P. Taylor. Flora of West Tropical Africa 2. 1963

    Habit
    Aquatic with long stolons bearing short lateral scapes
    Flowers
    1 to 4 yellow flowers, the pedicels elongating and reflexing in fruit.
    [FZ]

    Lentibulariaceae, P. Taylor. Flora Zambesiaca 8:3. 1988

    Habit
    Aquatic herb.
    Stolons
    Stolons filiform to relatively thick and fleshy, up to 50 cm. x 0.3–3.0 mm., or more or less densely covered with simple stellate hairs which extend also to the traps, basal part of the leaves and peduncle; internodes 2–10(15) mm.
    Rhizoids
    Rhizoids usually absent but when present 2–3 from the peduncle base, capillary, 3–5 mm. long.
    Leaves
    Leaves very numerous, digitately divided to the base into 2–5 primary segments, 3–1.30 mm. long, repeatedly dichotomously divided, the ultimate segments capillary, setulose.
    Traps
    Traps inserted in the angles between the primary segments and between subsequent dichotomies, very variable in number and size, ovoid, shortly stalked, 1–6 mm. long; mouth lateral with 2 dorsal simple to conspicuously branched hairs and sometimes with ventral shorter simple hairs.
    Inflorescences
    Inflorescence lateral, 1.5–18 cm. high; flowers 1–3(4), distant, the lowermost sometimes very near the base of the peduncle; peduncle erect or rarely twining, filiform; scales absent; bracts basifixed, quadrate or circular, more or less encircling the base of the pedicel, 1–3 mm. long; bracteoles absent; pedicels filiform, erect at anthesis, usually strongly deflexed or decurved in fruit, 2–35 mm. long.
    Calyx
    Calyx lobes subequal, broadly ovate, 1–2 mm. long, scarcely accrescent, with apex obtuse or rounded.
    Corolla
    Corolla pale to quite deep yellow, usually with brown or reddish nerves, more or less densely covered externally with fine short hairs, rarely glabrous, 3–15 mm. long; superior lip broadly ovate or circular, with apex truncate or rounded, entire or emarginate, 1.5–3 times as long as the upper calyx lobe; inferior lip circular to subreniform, with apex more or less emarginate or 2-lobed; palate raised, bigibbous; spur cylindrical, more or less as long as the inferior lip.
    Stamens
    Filaments linear; anther-thecae more or less confluent.
    Pistil
    Ovary globose; style short or obsolete; stigma inferior lip semi-circular, sometimes ciliate, superior very short or obsolete.
    Fruits
    Capsule globose, circumscissile, 3–4 mm. in diam., few-many-seeded.
    Seeds
    Seeds lenticular, angular, 0.4–0.8 mm. wide, often narrowly winged on the angles; testa reticulate, cells distinct, isodiametric or more or less elongate.

    Images

    Distribution

    Native to:

    Angola, Benin, Botswana, Burkina, Burundi, Cameroon, Central African Repu, Chad, Congo, Ethiopia, Gabon, Gambia, Ghana, Guinea, Guinea-Bissau, Ivory Coast, Kenya, Madagascar, Malawi, Mali, Niger, Nigeria, Senegal, Sierra Leone, Sudan, Tanzania, Togo, Uganda, Zambia, Zaïre, Zimbabwe

    Synonyms

    Other Data

    Utricularia reflexa Oliv. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Apr 4, 1904 Vogel, E. [94], Chad K000430260
    Apr 4, 1904 Welwitsch. [269], Angola K000430329 Unknown type material
    Richards, H.M. [18086], Zambia 22953.000
    Chevalier, A. [22297], Côte d'Ivoire K000430206 Unknown type material
    Hepper, F.N. [5475], Tanzania 34338.000
    Milne-Redhead, E. [4107], Angola K001009223
    Brummitt, R.K. [16798], Malawi 25694.139
    Schafer [22389] 39842.000
    Johnston, H.H. [s.n.], Angola K001009225
    Bamps, P. [4152], Angola K001009224
    Barter [890] K000430203 Unknown type material
    Barter [s.n.] K000430204
    Barter [s.n.], Nigeria K000430205 Unknown type material
    Kotschy [201] K000430254
    Kotschy [201] K000430255
    Fries, R.E. [977], Zambia K000430330 Unknown type material
    Speke [s.n.] K000430247
    Welwitsch. [271], Angola K000430328 Unknown type material
    Stopp, K. [BO219], Angola K001009221
    Stopp, K. [BO 230], Angola K001009222
    Hess-Wyss, H. [52/283], Angola K001009215
    Hess-Wyss, H. [52/672], Angola K001009216
    Hess-Wyss, H. [52/1670], Angola K001009217
    Hess-Wyss, H. [52/1670], Angola K001009218
    Hess-Wyss, H. [52/2036], Angola K001009219
    Hess-Wyss, H. [52/1780], Angola K001009220

    Bibliography

    First published in J. Linn. Soc., Bot. 9: 146 (1865)

    Accepted by

    • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
    • Thiombiano, A., Schmidt, M., Dressler, S., Ouédraogo, A., Hahn, K. & Zizka, G. (2012). Catalogue des plantes vasculaires du Burkina Faso Boissiera 65: 1-391.
    • Lisowski, S. (2009). Flore (Angiospermes) de la République de Guinée Scripta Botanica Belgica 41: 1-517.
    • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
    • Hedberg, I., Kelbessa, E., Edwards, S., Demissew, S. & Persson, E. (eds.) (2006). Flora of Ethiopia and Eritrea 5: 1-690. The National Herbarium, Addis Ababa University, Ethiopia & The Department of Systematic Botany, Upps.
    • Catarino, L., Martins, E.S., Diniz, M.A. & Pinto-Basto, M.F. (2006). Check-list da flora vascular do parque natural das Lagos de Cufada (Guiné-Bissau) Garcia de Orta, Série de Botânica 17: 97-141.
    • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
    • Sita, P. & Moutsambote, J.-M. (2005). Catalogue des plantes vasculaires du Congo, ed. sept. 2005: 1-158. ORSTOM, Centre de Brazzaville.
    • Jones, M. (1991). A checklist of Gambian plants: 1-33. Michael Jones, The Gambia College.
    • Taylor, P. (1989). The Genus Utricularia: a taxonomic monograph: 1-724. Her Majesty's stationery office, London.
    • Boudet, G., Lebrun, J.P. & Demange, R. (1986). Catalogue des plantes vasculaires du Mali: 1-465. Etudes d'Elevage et de Médecine Vétérinaire des Pays Tropicaux.
    • Brunel, J.F., Hiepo, P. & Scholz, H. (eds.) (1984). Flore Analytique du Togo Phanérogames: 1-751. GTZ, Eschborn.
    • Boulvert, Y. (1977). Catalogue de la Flore de Centrafrique 2(1): 1-85. ORSTOM, Bangui.
    • Peyre de Fabregues, B. & Lebrun, J.-P. (1976). Catalogue des Plantes Vascularies du Niger: 1-433. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
    • Berhaut, J. (1975). Flore illustrée du Sénégal 4: 1-625. Gouvernement du Sénégal, Ministère du développement rural direction des eaux et forêta, Dakar.
    • Lebrun, J.-P., Audru, J., Gaston, A. & Mosnier, M. (1972). Catalogue des Plantes Vasculaires du Tchad Méridional: 1-289. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
    • Lewalle, J. (1970). Liste floristique et répartition altitudinale de la flore du Burundi occidental: 1-84. Université officielle de Bujumbura.

    Literature

    Flora of West Tropical Africa

    • Pellegr. in Bull. Soc. Bot. Fr. 61: 19.
    • F.T.A. 4, 2: 492
    • in J. Linn. Soc. 9: 146 (1867)

    Kew Backbone Distributions

    • Thiombiano, A., Schmidt, M., Dressler, S., Ouédraogo, A., Hahn, K. & Zizka, G. (2012). Catalogue des plantes vasculaires du Burkina Faso Boissiera 65: 1-391.
    • Lisowski, S. (2009). Flore (Angiospermes) de la République de Guinée Scripta Botanica Belgica 41: 1-517.
    • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
    • Hedberg, I., Kelbessa, E., Edwards, S., Demissew, S. & Persson, E. (eds.) (2006). Flora of Ethiopia and Eritrea 5: 1-690. The National Herbarium, Addis Ababa University, Ethiopia & The Department of Systematic Botany, Upps.
    • Catarino, L., Martins, E.S., Diniz, M.A. & Pinto-Basto, M.F. (2006). Check-list da flora vascular do parque natural das Lagos de Cufada (Guiné-Bissau) Garcia de Orta, Série de Botânica 17: 97-141.
    • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
    • Sita, P. & Moutsambote, J.-M. (2005). Catalogue des plantes vasculaires du Congo, ed. sept. 2005: 1-158. ORSTOM, Centre de Brazzaville.
    • Jones, M. (1991). A checklist of Gambian plants: 1-33. Michael Jones, The Gambia College.
    • Taylor, P. (1989). The Genus Utricularia: a taxonomic monograph: 1-724. Her Majesty's stationery office, London.
    • Launert, E. (ed.) (1988). Flora Zambesiaca 8(3): 1-118. Royal Botanic Gardens, Kew.
    • Boudet, G., Lebrun, J.P. & Demange, R. (1986). Catalogue des plantes vasculaires du Mali: 1-465. Etudes d'Elevage et de Médecine Vétérinaire des Pays Tropicaux.
    • Brunel, J.F., Hiepo, P. & Scholz, H. (eds.) (1984). Flore Analytique du Togo Phanérogames: 1-751. GTZ, Eschborn.
    • Boulvert, Y. (1977). Catalogue de la Flore de Centrafrique 2(1): 1-85. ORSTOM, Bangui.
    • Peyre de Fabregues, B. & Lebrun, J.-P. (1976). Catalogue des Plantes Vascularies du Niger: 1-433. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
    • Berhaut, J. (1975). Flore illustrée du Sénégal 4: 1-625. Gouvernement du Sénégal, Ministère du développement rural direction des eaux et forêta, Dakar.
    • Lebrun, J.-P., Audru, J., Gaston, A. & Mosnier, M. (1972). Catalogue des Plantes Vasculaires du Tchad Méridional: 1-289. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
    • Lewalle, J. (1970). Liste floristique et répartition altitudinale de la flore du Burundi occidental: 1-84. Université officielle de Bujumbura.

    Flora of Tropical East Africa

    • P. Taylor in Fl. Afr. Centr., Lentib.: 45 (1972).
    • P. Taylor in Kew Bulletin 18: 163 (1964)
    • P. Taylor in Flora of West Tropical Africa, ed. 2, 2: 380 (1963)
    • Stapf in Flora of Tropical Africa 4 (2): 492 (1906).
    • Oliv. in Journal of the Linnean Society of London, Botany 9: 146 (1865).

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of West Tropical Africa
    Flora of West Tropical Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2020. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2020. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Plants and People Africa
    Common Names from Plants and People Africa http://www.plantsandpeopleafrica.com/
    © Plants and People Africa http://www.plantsandpeopleafrica.com http://creativecommons.org/licenses/by/3.0/