1. Family: Cannabaceae Martinov
    1. Genus: Celtis L.
      1. Celtis philippensis Blanco

        This species is accepted, and its native range is Tropical & Subtropical Asia to N. Australia.

    [FZ]

    Ulmaceae, C. M. Wilmot-Dear. Flora Zambesiaca 9:6. 1991

    Habit
    Evergreen much-branched tree 5–20 m. tall, monoecious; bole often with short buttresses, bark smooth grey, wood white; young stems and branches very sparsely white-pubescent, glabrescent.
    Leaves
    Leaves (5)6.5–12(17.5) x 2.5–4.5(8) cm., elliptic to elliptic-ovate or elliptic-oblong; apex with a wide (usually short) acumen and mucronate tip; base slightly asymmetrical and broadly cuneate to rounded or subcordate; margin entire or (rarely in Flora Zambesiaca region) coarsely dentate in the upper half; lamina thinly coriaceous, pale green, punctate but smooth and ± shiny above, glabrous; 3-nerved from the base, the basal lateral nerves extending almost to the apex, as strongly prominent beneath as the midrib, upper lateral nerves 1–2(3) on each side of the midrib, rather fine and inconspicuous; secondary venation fairly closely parallel, ± horizontal between the midrib and basal lateral nerves.
    Petiole
    Petiole 4–10(16) mm. long; stipules 3–7 mm. long, lanceolate, shortly produced below the point of attachment, ± pubescent.
    Inflorescences
    Inflorescence in leaf axils towards the ends of branches; lower inflorescences 5–30 mm. long, containing many crowded male (lowers, these sessile or with pedicels to 2 mm. long, and a few female and bisexual flowers at apices of branches of these inflorescences, their pedicels usually longer; upper inflorescences usually short with several bisexual flowers.
    Calyx
    Sepals 1.5–2.5 mm. long, pubescent.
    Pistil
    Ovary ± glabrous, with a basal ring of long hairs; styles very shortly bifid, (1)1.5–2(3.5) mm. long.
    Fruits
    fruits red, 9–12 x 7–10 mm., ovoid, glabrous; pedicels 3–7 mm. long.
    [FTEA]

    Ulmaceae, R. M. Polhill. Flora of Tropical East Africa. 1966

    Habit
    Much branched evergreen monoecious tree or shrub, 3–20 m. tall, often with short sharp buttresses; bark smooth, grey.
    Twigs
    Young twigs puberulous or subglabrous.
    Leaves
    Leaf-blades chartaceous to coriaceous, elliptic to elliptic-oblong, (5–)8.5–17.5 cm. long, 3.5–7.8 cm. wide, acuminate, broadly cuneate to rounded or subcordate at the base, ± a little unequal-sided, entire or coarsely toothed in the upper half, glabrous, 3-nerved from the base, with the basal lateral nerves extending practically to the apex and the upper prominent lateral nerves l–2(–3) on each side of the midrib; petiole 4–16 mm. long.
    Stipules
    Stipules lanceolate, 3–7 mm. long, shortly produced below the point of attachment, ± pubescent.
    Inflorescences
    Cymes 5–30 mm. long in flower, up to 40 mm. long in fruit; lower ones with many clustered ♂ flowers and a few ♀ and hermaphroditeflowers at the top; upper ones with several hermaphroditeflowers.
    Calyx
    Sepals 4–5, 1.5– 2.5 mm. long, pubescent.
    Pistil
    Ovary glabrous or subglabrous with a basal ring of long hairs; styles shortly bifurcate, 1–2–3.5 mm. long.
    Fruits
    Fruits on 3–7 mm. long pedicels, ovoid, shortly beaked, up to 12 mm. long, 10 mm. across, red, glabrous.
    Habitat
    Lowland rain, swamp and riverine forest; 30–1200 m.
    Distribution
    K7 T3 T6 T8 U2 U4 Z Ethiopia to West Africa and south to Angola and Mozambique, also Madagascar, Mascarene Is. and tropical Asia to Australia

    Images

    Distribution

    Native to:

    Andaman Is., Bismarck Archipelago, Borneo, China South-Central, China Southeast, Hainan, India, Jawa, Laos, Lesser Sunda Is., Malaya, Maluku, Myanmar, New Guinea, Northern Territory, Philippines, Queensland, Solomon Is., Sri Lanka, Sulawesi, Sumatera, Taiwan, Thailand, Vietnam

    Celtis philippensis Blanco appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Sep 1, 2015 Poilane, E. [10116], Vietnam K000852767
    Jan 1, 2012 I Made Maduarta [IMM 122], Lesser Sunda Is. K000733399
    Jan 25, 2008 Cunningham, A. [193], Western Australia K000852752
    Nov 1, 2003 Onana, J.-M. [999], Cameroon K000197966
    Jan 1, 1976 Collins, D.J. [241], Thailand K000852768 Unknown type material
    Jan 1, 1976 Collins, D.J. [230], Thailand K000852770 Unknown type material
    Mar 1, 1971 Gardner, G. [s.n.], Sri Lanka K000852776
    Mar 1, 1971 Wight, R. [85], India K000852771 syntype
    Mar 1, 1971 Wight, R. [41], Tamil Nadu K000852772 syntype
    Mar 1, 1971 Walker, G.W. [s.n.], Sri Lanka K000852775
    Mar 1, 1971 s.coll. [s.n.], Jawa K000852765
    Mar 1, 1971 s.coll. [214], Sri Lanka K000852773 syntype
    Mar 1, 1971 s.coll. [214], Sri Lanka K000852774 syntype
    Jan 1, 1971 Commerson [s.n.], Mauritius K000243045
    Jan 1, 1971 Elmer, A.D.E. [16028], Philippines K000852760
    Jan 1, 1971 Elmer, A.D.E. [10999], Philippines K000852761
    Jan 1, 1971 s.coll. [s.n.], Western Australia K000852750
    Jan 1, 1971 s.coll. [s.n.], Northern Territory K000852751 Unknown type material
    May 1, 1970 Collins, D.J. [241], Thailand K000852769 Unknown type material
    Jan 1, 1963 Lebrun [3074], Zaire Celtis wightii K000243034
    Jan 1, 1963 Welw. [6302], Angola Celtis wightii K000243042 Unknown type material
    Chevalier, A. [21.459], Ivory Coast Celtis wightii K000243030 Unknown type material
    s.coll. [Cat. no. 9056] Celtis wightii K001132149

    First published in Fl. Filip.: 197 (1837)

    Accepted by

    • Tange, S. & al. (2018). Flora of Nam Kading national protected area II: 30 new records of Angiosperms for Laos Edinburgh Journal of Botany 75: 107-116.
    • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
    • Guymer, G.P. (2013). Celtis strychnoides Planch. an earlier name for Celtis australiensis Sattarian (Ulmaceae) Austrobaileya 9: 146-147.
    • Kalema, J. & Beentje, H. (2012). Conservation checklist of the trees of Uganda: 1-235. Kew Publishing, Royal Botanic Gardens, Kew.
    • Pandey, R.P. & Dilwakar, P.G. (2008). An integrated check-list flora of Andaman and Nicobar islands, India Journal of Economic and Taxonomic Botany 32: 403-500.
    • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
    • Wu, Z. & Raven, P.H. (eds.) (2003). Flora of China 5: 1-505. Science Press (Beijing) & Missouri Botanical Garden Press (St. Louis).
    • Lê, T.C. (2003). Danh l?c các loài th?c v?t Vi?t Nam 2: 1-1203. Hà N?i : Nhà xu?t b?n Nông nghi?p.
    • Govaerts, R. (1999). World Checklist of Seed Plants 3(1, 2a & 2b): 1-1532. MIM, Deurne.
    • Turner, I.M. (1995 publ. 1997). A catalogue of the Vascular Plants of Malaya Gardens' Bulletin Singapore 47(2): 347-655.
    • Launert, E. & Cope, G.V. (eds.) (1991). Flora Zambesiaca 9(6): 1-135. Royal Botanic Gardens, Kew.
    • Hancock, I.R. & Henderson, C.P. (1988). Flora of the Solomon Islands Research Bulletin Dodo Creek Research Station 7: 1-203.

    Literature

    Kew Backbone Distributions
    • Tange, S. & al. (2018). Flora of Nam Kading national protected area II: 30 new records of Angiosperms for Laos Edinburgh Journal of Botany 75: 107-116.
    • Guymer, G.P. (2013). Celtis strychnoides Planch. an earlier name for Celtis australiensis Sattarian (Ulmaceae) Austrobaileya 9: 146-147.

    Sources

    Art and Illustrations in Digifolia
    Digital Image © Board of Trustees, RBG Kew

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0