1. Family: Amaranthaceae Juss.
    1. Genus: Cyathula Blume
      1. Cyathula prostrata (L.) Blume

        This species is accepted, and its native range is Tropics & Subtropics.

    [CPLC]

    Bernal, R., Gradstein, S.R. & Celis, M. (eds.). 2015. Catálogo de plantas y líquenes de Colombia. Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá. http://catalogoplantasdecolombia.unal.edu.co

    Distribution
    Naturalizada en Colombia; Alt. 0 - 1900 m.; Amazonia, Andes, Llanura del Caribe, Orinoquia, Pacífico, Sierra Nevada de Santa Marta, Valle del Cauca, Valle del Magdalena.
    Habit
    Hierba
    Conservation
    No Evaluada
    [FTEA]

    Amaranthaceae, C.C. Townsend. Flora of Tropical East Africa. 1985

    Habit
    Annual herb (? sometimes short-lived perennial).
    Stem
    Stems prostrate and rooting at the lower nodes to erect, 0.2–1.2 m., simple or usually branched up to about the middle, ± swollen at the nodes, lower branches divaricate, the upper more erect; stem and branches bluntly 4-angled to subterete, striate or sulcate, subglabrous to ± densely pilose (especially the lower internodes).
    Leaves
    Leaves mostly rhombic to rhombic-ovate, sometimes rhombic-elliptic to shortly oval or subcircular, 1.5–8 × 1–4.5 cm., occasionally with the margin outline distinctly excavate below and/or above the middle, shortly acuminate at the apex, acute to rather blunt (more rarely rounded), shortly cuneate to cuneate-attenuate at the base, subglabrous to moderately pilose with strigose hairs on both surfaces, subsessile or distinctly (up to 13 mm.) petiolate.
    Inflorescences
    Spikes terminal on the stem and branches, at first dense, soon considerably elongating to as much as 35 cm. with maturing lower flowers increasingly distant, 5–7 mm. wide, peduncle up to ± 10 cm., axis and peduncle thinly to ± densely pilose; bracts and bracteoles lanceolate-ovate, ± 1.5–2 mm., mucronate with the shortly excurrent midrib, ciliate; flowers in sessile or shortly pedunculate cymose clusters (peduncles to ± 2 mm.) of 2–3 hermaphrodite flowers, the 2 laterals subtended by 2 modified flowers, or the uppermost hermaphrodite flowers of the spike solitary, similarly subtended by modified flowers, bibracteolate.
    Tepal
    Tepals elliptic-oblong, 2.25–3 mm., 3-nerved, subglabrous to ± densely white-pilose; outer firmer with the lateral nerves more distinct and joining the shortly excurrent midrib just below the apex, usually more densely white-pilose than the inner; inner sometimes ± falcate, slightly shorter.
    Spines
    Spines of modified flowers sharply uncinate, numerous, glabrous, reddish, ± 2 mm., fasciculate, in fruit scarcely exceeding the tepals of the fertile flower; 2–3-flowered clusters ± globose, finally deflexed, falling as a unit to form a burr ± 5 cm. in diameter.
    Stamens
    Filaments very slender, ± 1.5 mm., the pseudostaminodes rectangular-cuneate with a truncate dentate or excavate apex.
    Ovary
    Ovary with a pileiform cap.
    Style
    Style slender, ± 0.6 mm., often slightly swollen towards the base.
    Fruits
    Capsule ovoid, membranous save for the flat firm apex, ± 1.5 mm.
    Seeds
    Seed ovoid, smooth, ± 1.5 mm., shining, brown.
    [FZ]

    Amaranthaceae, C. C. Townsend. Flora Zambesiaca 9:1. 1988

    Habit
    Annual herb (? sometimes short-lived perennial), stems prostrate and rooting at the lower nodes to erect, 0.2–1.2 m., simple or usually branched up to about the middle, more or less swollen at the nodes, lower branches divaricate, the upper more erect; stem and branches bluntly 4-angled to subterete, striate or sulcate, subglabrous to more or less densely pilose (especially the lower internodes).
    Leaves
    Leaves 1.5–8 × 1–5 cm., mostly rhomboid to rhomboid-ovate, sometimes rhomboid-elliptic to shortly oval or subcircular, occasionally with the margin outline distinctly excavate below and/or above the middle, shortly acuminate at the apex, acute to rather blunt (more rarely rounded), shortly cuneate to cuneate-attenuate at the base, subglabrous to moderately pilose with strigose hairs on both surfaces, subsessile or distinctly (up to 13 mm. long) petiolate.
    Inflorescences
    Spikes terminal on the stem and branches, at first dense, soon considerably elongating to as much as 25 cm. but mostly c. 6–15 cm. with maturing lower flowers increasingly distant, 5–7 mm. wide, peduncle up to c. 10 cm. long, axis and peduncle thinly to more or less densely pilose;bracts and bracteoles membranous, lanceolate-ovate, c. 1.5–2 mm. long, mucronate with the shortly excurrent midrib, glabrous or ciliate; flowers in sessile or shortly pedunculate clusters (peduncles to c. 2 mm. long), cymose, of 2–3 hermaphrodite flowers, the two laterals subtended by two modified flowers, or the uppermost hermaphrodite flowers of the spike solitary, similarly subtended by modified flowers, bibracteolate.
    Tepal
    Tepals 2.25–3 mm. long, elliptic-oblong, 3-nerved, subglabrous to more or less densely white-pilose; the outer firmer with the lateral nerves more distinct and joining the shortly excurrent midrib just below the apex, usually more densely white-pilose than the inner; the inner sometimes more or less falcate, slightly shorter.
    Spines
    Spines of modified flowers sharply uncinate, numerous, glabrous, reddish, c. 2 mm., fasciculate, in fruit scarcely exceeding the tepals of the fertile flower; 2- or 3-flowered clusters, more or less globose finally deflexed, falling as a unit to form a “burr” c. 5 mm in diam.
    Androecium
    Filaments very slender, c. 1.5 mm. long, the pseudostaminodes rectangular-cuneate with a truncate, dentate or excavate apex.
    Ovary
    Ovary with a pileiform cap.
    Style
    Style slender, c. 0.6 mm. long, often slightly swollen towards the base.
    Fruits
    Capsule ovoid, membranous save for the flat, firm apex, c. 1.5 mm. long.
    Seeds
    Seed ovoid, c. 1.5 mm. long, shining, brown, smooth.
    [FWTA]

    Amaranthaceae, Hutchinson and Dalziel. Flora of West Tropical Africa 1:1. 1954

    Habit
    More or less erect, much-branched weed, 1–3 ft. high
    Leaves
    Leaves green, sometimes tinged with red
    Flowers
    Flowers green, often galled
    Ecology
    In waste places and forest margins.

    Images

    Distribution

    Native to:

    Angola, Assam, Bangladesh, Benin, Borneo, Brazil North, Brazil Northeast, Brazil South, Brazil Southeast, Brazil West-Central, Burkina, Cambodia, Cameroon, Central African Repu, China South-Central, China Southeast, Colombia, Congo, East Himalaya, Ecuador, El Salvador, Ethiopia, Fiji, Gabon, Gambia, Ghana, Guinea, Guinea-Bissau, Gulf of Guinea Is., Hainan, Honduras, India, Ivory Coast, Jawa, Laos, Lesser Sunda Is., Liberia, Malawi, Malaya, Mali, Maluku, Myanmar, Nepal, New Guinea, Nicaragua, Nicobar Is., Nigeria, Panamá, Peru, Philippines, Queensland, Samoa, Senegal, Sierra Leone, Solomon Is., Sri Lanka, Sudan, Sulawesi, Sumatera, Taiwan, Tanzania, Thailand, Togo, Tonga, Uganda, Vanuatu, Venezuela, Vietnam, Zambia, Zaïre, Zimbabwe

    Introduced into:

    Bolivia, Cook Is., French Guiana, Guyana, Jamaica, Leeward Is., Marquesas, Mauritius, Puerto Rico, Réunion, Seychelles, Society Is., Suriname, Trinidad-Tobago, Tubuai Is., Venezuelan Antilles, Windward Is.

    Cyathula prostrata (L.) Blume appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Jan 1, 2008 Ellen, R.F. [185], Indonesia K000611703
    Jan 1, 1998 Ndam, N. [1095], Cameroon K000011856
    Jan 1, 1998 Fraser, P. [457], Cameroon K000011855
    Dec 1, 1994 Cheek, M. [5835], Cameroon K000086534
    Dec 1, 1994 Cheek, M. [5731], Cameroon K000086536
    Dec 1, 1994 Cable, S. [351], Cameroon K000086533
    Dec 1, 1994 Cable, S. [188], Cameroon K000086535
    Dec 1, 1994 Cable, S. [426], Cameroon K000086537
    Dec 1, 1994 Cable, S. [599], Cameroon K000086538
    Dec 1, 1993 Sidwell, K. [112], Cameroon K000381463
    Jan 1, 1972 Maas, P.J.M. [P12831], Brazil K001207223
    Feb 1, 1968 Meurillon, A. [CNAD 547], Cameroon K000025589
    Jan 1, 1968 Leeuwenberg, A.J.M. [6935], Cameroon K000025587
    Jan 1, 1953 Morton, J.K. [6752], Cameroon K000025585
    Jan 1, 1953 Morton, J.K. [7053], Cameroon K000025586
    Krapovickas, A. [40167], Brazil K001207221
    Hage, J.L. [759], Brazil K001207220
    Irwin, H.S. [47329], Brazil K001207212
    Eiten, G. [10144], Brazil K001207210
    Silva, L.A.M. [423], Brazil K001207219
    Archer, W.A. [7844], Brazil K001207206
    Archer, W.A. [7844], Brazil K001207211
    Mori, S. [10289], Brazil K001207215
    Pires, J.M. [51320], Brazil K001207213
    Belém, R.P. [1113], Brazil K001207224
    Santos, R.R. [1473], Brazil K001207207
    Prance, G.T. [1270], Brazil K001207214
    Prance, G.T. [5862], Bolivia K001207216
    Prance, G.T. [13722], Brazil K001207217
    Prance, G.T. [10278], Brazil K001207222
    Cavalcante, P. [2400], Brazil K001207218
    Strudwick, J.J. [4000], Brazil K001207208
    Costa, M.A.S. [579], Brazil K001207209
    Mildbraed, G. W. J. [10789], Cameroon K000025584
    Chuml, H. [220], Cameroon K000025582
    Chuml, H. [313], Cameroon K000025583
    Chuml, H. [125], Cameroon K000025581
    Burchell [9259], Brazil K001207204
    Burchell [9464], Brazil K001207205
    s.coll. [Cat. no. 6938], India Desmochaeta repens K001126300
    s.coll. [Cat. no. 6938], India Desmochaeta repens K001126301
    s.coll. [Cat. no. 6938] Desmochaeta repens K001126302

    First published in Bijdr. Fl. Ned. Ind.: 549 (1826)

    Accepted by

    • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
    • Velayos, M., Barberá, P., Cabezas, F.J., de la Estrella, M., Fero, M. & Aedo, C. (2014). Checklist of the vascular plants of Annobón (Equatorial Guinea) Phytotaxa 171: 1-78.
    • Jørgensen, P.M., Nee, M.H. & Beck., S.G. (eds.) (2013). Catálogo de las plantas vasculares de Bolivia Monographs in Systematic Botany from the Missouri Botanical Garden 127: 1-1741. Missouri Botanical Garden.
    • Thiombiano, A., Schmidt, M., Dressler, S., Ouédraogo, A., Hahn, K. & Zizka, G. (2012). Catalogue des plantes vasculaires du Burkina Faso Boissiera 65: 1-391.
    • Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies Smithsonian Contributions to Botany 98: 1-1192.
    • Ramos Senna, L., Giulietti, A.M. & Rapini, A. (2010). Flora da Bahia: Amaranthaceae - Amaranthoideae e Gomphrenoideae Sitientibus Sérrie Ciências Biológicas 10: 3-73.
    • Lejoy, J., Ndjele, M.-B. & Geerinck, D. (2010). Catalogue-flore des plantes vasculaires des districts de Kisangani et de la Tshopo (RD Congo) Taxonomania 30: 1-307.
    • Sarmah, K.K. & Borthakur, S.K. (2009). A checklist of angiospermic plants of Manas national park in Assam, India Pleione 3: 190-200.
    • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
    • Hokche, O., Berry, P.E. & Huber, O. (eds.) (2008). Nuevo Catálogo de la Flora Vascular de Venezuela: 1-859. Fundación Instituto Botánico de Venezuela.
    • Nelson Sutherland, C.H. (2008). Catálogo de las plantes vasculares de Honduras. Espermatofitas: 1-1576. SERNA/Guaymuras, Tegucigalpa, Honduras.
    • Newman, M., Ketphanh, S., Svengsuksa, B., Thomas, P., Sengdala, K., Lamxay, V. & Armstrong, K. (2007). A checklist of the vascular plants of Lao PDR: 1-394. Royal Botanic Gardens, Edinburgh.
    • Catarino, L., Martins, E.S., Diniz, M.A. & Pinto-Basto, M.F. (2006). Check-list da flora vascular do parque natural das Lagos de Cufada (Guiné-Bissau) Garcia de Orta, Série de Botânica 17: 97-141.
    • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
    • Catarino, L., Sampaio Martins, E., Pinto-Basto, M.F. & Diniz, M.A. (2006). Plantas Vasculares e Briófitos da Guiné-Bissau: 1-298. Instituto de investigação científica tropical, Instituto Português de apoio ao desenvolvimento.
    • Sita, P. & Moutsambote, J.-M. (2005). Catalogue des plantes vasculaires du Congo, ed. sept. 2005: 1-158. ORSTOM, Centre de Brazzaville.
    • Correa A., Mireya D. Galdames, Carmen Correa A., M. D., C. Galdames & M. S. de Stapf (2004). Catálogo de las Plantas Vasculares de Panamá: 1-599. Smithsonian Tropical Research Institute.
    • Kress, W.J., DeFilipps, R.A., Farr, E. & Kyi, D.Y.Y. (2003). A Checklist of the Trees, Shrubs, Herbs and Climbers of Myanmar Contributions from the United States National Herbarium 45: 1-590. Smithsonian Institution.
    • Lê, T.C. (2003). Danh l?c các loài th?c v?t Vi?t Nam 2: 1-1203. Hà N?i : Nhà xu?t b?n Nông nghi?p.
    • Edwards, S., Tadesse, M., Demissew, S. & Hedberg, I. (eds.) (2000). Flora of Ethiopia and Eritrea 2(1): 1-532. The National Herbarium, Addis Ababa University, Ethiopia & The Department of Systematic Botany, Upps.
    • Dy Phon, P. (2000). Dictionnaire des plantes utilisées au Cambodge: 1-915. chez l'auteur, Phnom Penh, Cambodia.
    • Govaerts, R. (1999). World Checklist of Seed Plants 3(1, 2a & 2b): 1-1532. MIM, Deurne.
    • Boggan, J. Funck, V. & Kelloff, C. (1997). Checklist of the Plants of the Guianas (Guyana, Surinam, Franch Guiana) ed. 2: 1-238. University of Guyana, Georgetown.
    • Turner, I.M. (1995). A catalogue of the Vascular Plants of Malaya Gardens' Bulletin Singapore 47(1): 1-346.
    • Bosser, J. & al. (eds.) (1994). Flore des Mascareignes 136-148: 1. IRD Éditions, MSIRI, RBG-Kew, Paris.
    • Launert, E. (ed.) (1988). Flora Zambesiaca 9(1): 1-179. Royal Botanic Gardens, Kew.
    • Hancock, I.R. & Henderson, C.P. (1988). Flora of the Solomon Islands Research Bulletin Dodo Creek Research Station 7: 1-203.
    • Boudet, G., Lebrun, J.P. & Demange, R. (1986). Catalogue des plantes vasculaires du Mali: 1-465. Etudes d'Elevage et de Médecine Vétérinaire des Pays Tropicaux.
    • Townsend, C.C. (1985). Flora of Tropical East Africa, Amaranthaceae: 1-136.
    • Grierson, A.J.C. & Long, D.G. (1984). Flora of Bhutan 1(2): 189-462. Royal Botanic Gardens, Edinburgh.
    • Boulvert, Y. (1977). Catalogue de la Flore de Centrafrique 2(1): 1-85. ORSTROM, Bangui.
    • Berhaut, J. (1971). Flore illustrée du Sénégal 1: 1-626. Gouvernement du Sénégal, Ministère du développement rural direction des eaux et forêta, Dakar.
    • Hutchinson, J., Dalziel, J.M. & Keay, R.W.J. (1954-1958). Flora of West Tropical Africa, ed. 2, 1: 1-828.
    • Christopherson, E. (1935). Flowering Plants of Samoa Bernice P. Bishop Museum Bulletin 128: 1-221.

    Literature

    Flora of West Tropical Africa
    • Chev. Bot. 529.
    • —F.T.A. 6, 1: 43
    Kew Backbone Distributions
    • Forzza, R.C., Zappi, D. & Souza, V.C. (2016-continuously updated). Flora do Brasil 2020 em construção http://reflora.jbrj.gov.br/reflora/listaBrasil/ConsultaPublicaUC/ResultadoDaConsultaNovaConsulta.do.
    • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
    • Jørgensen, P.M., Nee, M.H. & Beck., S.G. (eds.) (2013). Catálogo de las plantas vasculares de Bolivia Monographs in Systematic Botany from the Missouri Botanical Garden 127: 1-1741. Missouri Botanical Garden.
    • Thiombiano, A., Schmidt, M., Dressler, S., Ouédraogo, A., Hahn, K. & Zizka, G. (2012). Catalogue des plantes vasculaires du Burkina Faso Boissiera 65: 1-391.
    • Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies Smithsonian Contributions to Botany 98: 1-1192.
    • Idárraga-Piedrahita, A., Ortiz, R.D.C., Callejas Posada, R. & Merello, M. (eds.) (2011). Flora de Antioquia: Catálogo de las Plantas Vasculares 2: 1-939. Universidad de Antioquia, Medellín.
    • Ramos Senna, L., Giulietti, A.M. & Rapini, A. (2010). Flora da Bahia: Amaranthaceae - Amaranthoideae e Gomphrenoideae Sitientibus Sérrie Ciências Biológicas 10: 3-73.
    • GBIF (2008-continuously updated). Global Biodiversity Information Facility http://www.gbif.org/.
    • Pandey, R.P. & Dilwakar, P.G. (2008). An integrated check-list flora of Andaman and Nicobar islands, India Journal of Economic and Taxonomic Botany 32: 403-500.
    • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
    • Nelson Sutherland, C.H. (2008). Catálogo de las plantes vasculares de Honduras. Espermatofitas: 1-1576. SERNA/Guaymuras, Tegucigalpa, Honduras.
    • Catarino, L., Martins, E.S., Diniz, M.A. & Pinto-Basto, M.F. (2006). Check-list da flora vascular do parque natural das Lagos de Cufada (Guiné-Bissau) Garcia de Orta, Série de Botânica 17: 97-141.
    • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
    • Sita, P. & Moutsambote, J.-M. (2005). Catalogue des plantes vasculaires du Congo, ed. sept. 2005: 1-158. ORSTOM, Centre de Brazzaville.
    • Correa A., Mireya D. Galdames, Carmen Correa A., M. D., C. Galdames & M. S. de Stapf (2004). Catálogo de las Plantas Vasculares de Panamá: 1-599. Smithsonian Tropical Research Institute.
    • Dy Phon, P. (2000). Dictionnaire des plantes utilisées au Cambodge: 1-915. chez l'auteur, Phnom Penh, Cambodia.
    • Boggan, J. Funck, V. & Kelloff, C. (1997). Checklist of the Plants of the Guianas (Guyana, Surinam, Franch Guiana) ed. 2: 1-238. University of Guyana, Georgetown.
    • Bosser, J. & al. (eds.) (1994). Flore des Mascareignes 136-148: 1. IRD Éditions, MSIRI, RBG-Kew, Paris.
    • Hancock, I.R. & Henderson, C.P. (1988). Flora of the Solomon Islands Research Bulletin Dodo Creek Research Station 7: 1-203.
    • Boulvert, Y. (1977). Catalogue de la Flore de Centrafrique 2(1): 1-85. ORSTROM, Bangui.
    • Berhaut, J. (1971). Flore illustrée du Sénégal 1: 1-626. Gouvernement du Sénégal, Ministère du développement rural direction des eaux et forêta, Dakar.
    • Yuncker, T.G. (1959). Plants of Tonga Bernice P. Bishop Museum Bulletin 220: 1-283.
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    • J. G. Baker & C. B. Clarke in Flora of Tropical Africa 6(1): 43 (1909).
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    Sources

    Art and Illustrations in Digifolia
    Digital Image © Board of Trustees, RBG Kew

    Catálogo de Plantas y Líquenes de Colombia
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of West Tropical Africa
    Flora of West Tropical Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0