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This species is accepted, and its native range is Ecuador.

[KBu]

Santamaría-Aguilar, D. & Monro, A.K. (2019). Compendium of Freziera (Pentaphylacaceae) of South America including eleven new species and the typification of 22 names. Kew Bulletin 74: 14. https://doi.org/10.1007/s12225-018-9790-1

Conservation
Using the three documented gatherings of Freziera ecuadoriensis and IUCN (2001) criteria applied using GeoCAT (Bachman et al. 2011) we calculate the EOO to be 2698 km2 (EN, IUCN criterion B). A survey of the EOO using the Global Forest Watch (2014) interactive map suggests that there has been c. 2% deforestation between 2001 and 2015 (subcriteria b). However, a review of the three localities in GoogleEarth suggests that deforestation rates have been much higher. For Øllgaard & Balslev 9345 it is possible to estimate the collection locality as within a few hundred metres of 2°43'8.94"S, 78°37'11.79"W (decimal) using the altitude and directions to the collection site provided on the herbarium label. This locality is within a large valley where >90% of the forest at the elevation range where the gathering was made has been converted to agricultural land, presumably prior to 2001. Luteyn & Romoleroux 14544 was collected at a locality now within a km of a largely deforested landscape and where below 2400 m all forest has been converted to agriculture (Google Earth). Jørgensen et al. 1052 was collected at a locality which now falls within a largely agricultural landscape where <50% of the original forest remains (Google Earth). In addition, the collectors observed that the forest at the locality in 1994 when the gathering was made, was under heavy pressure from charcoal production. We therefore suggest that low estimates for active deforestation in the EOO by Global Forest Watch are due to most of the EOO having already been deforested by 2001, the date from which data has been compiled. We conclude that the EOO has been subject to profound forest loss and fragmentation (subcriteria a) since the time that the gatherings we cite here were made (1976 – 1994) and therefore assess F. ecuadoriensis as Endangered (EN) under Criteria B and subcriteria a & b.
Distribution
Freziera ecuadoriensis is endemic to Ecuador from where it is known from the provinces of Azuay, Loja and Zamora-Chinchipe provinces, at elevations of 2200 to 3360 m.
Ecology
Freziera ecuadoriensis is documented from montane primary and secondary forest, roadsides and open areas associated with Chusquea Kunth (Poaceae).
Morphology Branches
Mature branches terete to weakly angulate, solid, bark brown, greyish or blackish; leaf-bearing branches terete to weakly angulate, densely hirsute, the hairs 0.7 – 2 mm, golden or yellow, papillose or not, with or without lenticels, where present lenticels whitish, elliptical to orbicular
Morphology General Buds
Terminal bud conduplicate-involute 1 – 3.2 cm, densely hirsute, the hairs 0.3 – 1.6 mm, golden, yellow or red-brown
Morphology General Habit
Shrub or tree to 10 m; bark and cambium unknown
Morphology Leaves
Leaves petiolate, petiole (0.1 –) 0.3 – 0.6 cm, adaxially grooved, abaxially rounded, hirsute or glabrescent, where pubescent the hairs golden, yellow or red-brown, not winged, margins entire; colleters absent; lamina 3.8 – 7.7 × 1.3 – 3.8 cm, narrowly elliptic or ovate, usually drying brown; adaxial surface glabrous (pubescent in young leaves), not pustulate; abaxial surface hirsute or woolly, the hairs 0.3 – 2.7 mm, golden or yellow, papillose or not; midrib adaxial surface flattened or weakly grooved, glabrescent or densely pubescent, abaxial surface rounded, prominent, pustulate or not, densely pubescent; lateral veins 10 – 24 pairs (including minor ones), flattened and impressed on adaxial surface, raised on abaxial surface; higher order venation distinct on both surfaces, more conspicuous on the abaxial surface; base cuneate or subobtuse, frequently revolute, weakly asymmetrical; margin serrulate, with 19 – 46 teeth per side, each tooth with a black or red-brown, conical, caducous seta, setae frequently ringed by hairs; apex acuminate or acute, bearing a single curved black or brown caducous seta
Morphology Reproductive morphology Flowers
Flowers unisexual?: 2.5 – 3.0 mm diam. in bud; sepals 5, imbricate; outer sepals 3 – 5.5 × 2.4 – 4.5 mm, broadly ovate, abaxial surface tomentose, adaxial surface glabrous, margin entire and ciliate, apex obtuse; inner sepals 3 – 3.2 (4 –) × 2.2 – 2.8 mm, ovate to broadly ovate, abaxial surface glabrous or glabrescent, adaxial surface glabrous, margin entire and ciliate, apex obtuse; petals 5, 3 – 5 × 1.8 – 2.5 mm, white or pink-tinged, distinct, glabrous, ovate, margin entire, apex acute or obtuse Pistillate flowers: staminodes 16 – 22, 0.5 – 1.1 mm, free, flattened, linear, apex acute; gynoecium 3-locular, 1.7 – 3 × 1.2 – 1.5 mm, subglobose, glabrous; style simple; stigmatic lobes 3
Morphology Reproductive morphology Fruits
Fruits 6 – 8 × 4.5 – 6 mm, globose, glabrous, green or green turning dark brown; fruit walls 0.3 – 0.4 mm thick; seeds (16 –) 43 – 57 per fruit, red-brown to brown, shiny, 1 – 1.6 mm long, sub-reniform, foveolate.
Morphology Reproductive morphology Inflorescences
Inflorescences axillary, fasciculate, bearing 1, 2 (3) flowers per axil; pedicel c. 4 mm, terete, erect, hirtellous; bracts at or above pedicel base persistent or caducous, 3.2 – 5.4 × 1 – 3.8 mm (7 × 6 mm), narrowly triangulate or broadly ovate, keeled or not keeled, abaxial surface woolly, hirtellous or nearly glabrous, adaxial surface glabrous, occasionally sparsely pubescent, margin with setae, apex acute to acuminate with a caducous black or red-brown curved seta at apex; bracteoles at pedicel apex persistent, opposite, unequal, 2, 3.2 – 5.5 × 1.8 – 3.5 mm, weakly triangulate, keeled or not keeled, abaxial surface appressed pubescent to hirtellous, adaxial surface glabrous, margin entire, occasionally with a few setae, apex rounded or acute, occasionally bearing a seta
Note
This species is named after Ecuador where it is endemic. The collection B. Øllgaard & H. Balslev 9345 (MO, NY) from Azuay Province differs from the type and other collections from Zamora-Chinchipe Province in having relatively larger leaves with broader leaf laminae. Further research may conclude that this collection belongs to a distinct taxon. Freziera ecuadoriensis can be distinguished from other Freziera species by the densely pubescent young branches, relatively small leaves with relatively short petioles lacking associated colleters, conspicuous venation on both surfaces, dense pubescence on the abaxial surface, revolute weakly asymmetrical leaf base and serrulate margin, few-flowered inflorescences, pedicellate flowers, narrow floral buds (2.5 – 3 mm wide) and thin-walled fruits. Freziera ecuadoriensis is most similar to F. suberosa Tul. from which it can be distinguished by the pubescent abaxial surface of the outer sepals, smaller inner sepals and smaller fruits.
Phenology
Freziera ecuadoriensis has been collected in flower from July to December and in fruit from April to December.
Type
Ecuador, Zamora-Chinchipe Province, Loja-Zamora old road, 03°58'S 079°06'W, 2200 – 2745 m, 23 April 1992 (fr.), J. L. Luteyn & K. Romoleroux 14544 (holotype K [K001289261]; isotypes MO [MO1577147], NY [NY03091296]).

Native to:

Ecuador

Freziera ecuadoriensis D.Santam. & A.K.Monro appears in other Kew resources:

First published in Kew Bull. 74(1)-14: 15 (2019)

Accepted by

  • Govaerts, R., Nic Lughadha, E., Black, N., Turner, R. & Paton, A. (2021). The World Checklist of Vascular Plants, a continuously updated resource for exploring global plant diversity. https://doi.org/10.1038/s41597-021-00997-6 Scientific Data 8: 215.

Literature

Kew Bulletin

  • Angiosperm Phylogeny Group — APG (2016). An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants: APG IV. Bot. J. Linn. Soc. 181: 1 – 20.
  • Arroyo Padilla, L. (1993). Theaceae D. Don. In: T. J. Killeen, E. García Estigarribia & S. G. Beck (eds), Guía de Árboles de Bolivia, pp. 777 – 782. Herbario Nacional de Bolivia & Missouri Botanical Garden, Edit. Quipus srl., La Paz.
  • Bachman, S., Moat, J., Hill, A. W., de la Torre, J. & Scott, B. (2011). Supporting Red List threat assessments with GeoCAT: geospatial conservation assessment tool. In: V. Smith & L. Penev (eds), e-Infrastructures for data publishing in biodiversity science. ZooKeys 150: 117 – 126, and GeoCAT. Geospatial Conservation Assessment Tool. [Online]. Available from: http://geocat.kew.org/. Accessed 14 April 2017.
  • Baptiste, B., Pinedo-Vasquez, M., Gutierrez-Velez, V. H., Andrade, G. I., Vieira, P., Estupiñán-Suárez, L. M., Londoño, M. C., Laurance, W. & Lee, T. M. (2017). Greening peace in Colombia. Nat. Ecol. Evol. 1: 0102.
  • Berry, P. E. & Weitzman, A. L. (2005). Ternstroemiaceae. In: P. E. Berry, K. Yatskievych & B. K. Holst (eds), Flora of the Venezuelan Guayana 9: 300 – 308, Rutaceae – Zygophyllaceae. Missouri Botanical Garden Press, St. Louis.
  • Berry, P. E. & Weitzman, A. L. (2008). Ternstroemiaceae. In: O. Hokche, P. E. Berry & O. Huber (eds), Nuevo Catálogo de la Flora Vascular de Venezuela, pp. 638 – 640. Fundación Instituto Botánico de Venezuela, Caracas.
  • Blume, C. L. (1856). Museum botanicum Lugduno-Batavum 2: 1 – 256. E. J. Brill, Leiden.
  • Bonpland, A. (1805, published 1808). In: A. von Humboldt & A. Bonpland, Plantae Aequinoctiales, per Regnum Mexici, in Provinciis Caracarum et Nova Andalusiae, in Peruvianorum, Quitensium, Novae Granate Andibus, ad Orinoci, Fluvi Nigri, Fluminis Amazonum tipas nascentes. 1: 1 – 234. F. Schoell, Paris.
  • Britton, N. L. (1889). An enumeration of the plants collected by Dr. H. H. Rusby in South America, 1885 – 1886. Part XVI. Bull. Torrey Bot. Club 16: 61 – 64.
  • Choisy, J. D. (1855). Memoire sur les Familles des Ternstroemiacees et Camelliacees. Mém. Soc. Phys. Genève 14: 1 – 95.
  • Christenhusz, M. J. M. & Byng, J. W. (2018). Pentaphylacaceae. In: M. J. M. Christenhusz, M. Fay & J. W. Byng (eds), The Global Flora. A practical flora to vascular plant species of the world. GLOVAP nomenclature Part 1. p. 144. Plant Gateway.
  • Cuello, N. A. & Santamaría-Aguilar, D. (2015). A. new species of Freziera (Pentaphylacaceae) from the Venezuelan Andes. Harvard Pap. Bot. 20: 147 – 150.
  • Deng, J. Z., Starck, S. R. & Hecht, S. M. (2000). Pentacyclic triterpenoids from Freziera sp. that inhibit DNA polymerase. Bioorg. Med. Chem. Lett. 8, 247 – 250.
  • Dorr, L. J., Stergios, B., Smith, A. R. & Cuello, N. A. (2000). Catalogue of the Vascular Plants of Guaramacal National Park, Portuguesa and Trujillo states, Venezuela. Contr. U.S. Natl. Herb. 40: 1 – 155.
  • Forrest, J. L., Sanderson, E. W., Wallace, R., Siles, T. M. L., Gómez, L. H. C. & Coppolillo, P. (2008). Patterns of Land Cover Change in and Around Madidi National Park, Bolivia. Biotropica 40: 285 – 294.
  • Gentry, A. H. (1981). New species and a new combination in Palmae, Theaceae, Araliaceae, Apocynaceae, and Bignoniaceae from the Choco and Amazonian Peru. Ann. Missouri Bot. Gard. 68: 112 – 121.
  • Global Forest Watch (2014). Interactive Map. World Resources Institute. Available from: www.globalforestwatch.org. Accessed 09 June 2017.
  • Global Plants database (2017). Available from: plants.jstor.org. Accessed 11 November 2017.
  • Gradstein, S. R. (2017). Freziera. In: R. Bernal, S. R. Gradstein & M. Celis (eds), Catálogo de plantas y líquenes de Colombia. Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá. Available from: http://catalogoplantasdecolombia.unal.edu.co. Accessed 08 November 2017.
  • Hiepko, P. (1987). The collections of the Botanical Museum Berlin-Dahlem (B) and their history. Englera 7: 219 – 252.
  • Hieronymus, G. (1895). Plantae Lehmannianae in Guatemala, Costarica, Columbia et Ecuador collectae, additis quibusdam ab aliis collectoribus ex iisdem regionibus necnon e Venezuela et Peruvia allatis, quas determinavit et descripsit adjuvantibus aliis auctoribus. Bot. Jahrb. Syst. 20: 1 – 72.
  • Hieronymus, G. (1896). Plantae steubelianae novae, quas descripsit adjuvantibus aliis auctoribus. Bot. Jahrb. Syst. 21: 306 – 378.
  • Hirth, S. (2014). Pentaphylacaceae. In: P. M. Jørgensen, M. Nee & S. G. Beck, Catálogo de las Plantas Vasculares de Bolivia. 2 Vols, Pp. 1007 – 1008. Missouri Botanical Garden Press, St Louis.
  • Holtmeier, F. K. (2015). Carl Troll: His footprints in physical geography. Prog. Phys. Geogr. 39: 554 – 565.
  • IUCN Red List of threatened species (2017). Available from: www.iucnredlist.org. Accessed 11 November 2017.
  • IUCN Standards and Petitions Subcommittee. (2016). Guidelines for using the IUCN Red List Categories and Criteria. Version 12. Prepared by the Standards and Petitions Subcommitttee, 101 pp. Available from: http://www.iucnredlist.org/documents/RedListGuidelines.pdf. Accessed 29 April 2017.
  • IUCN [International Union for Conservation of Nature and Natural Resources Species Survival Commission]. (2001). IUCN Red List Categories: Version 3.1. International Union for Conservation of Nature and Natural Resources, Gland and Cambridge.
  • International Code of Nomenclature for algae, fungi, and plants (Melbourne Code) (ICN) (2011). Available from: http://www.iapt-taxon.org/nomen/main.php. Accessed 05 June 2017.
  • Jiménez, Q. & Santamaría, D. (2015). Freziera, Theaceae. In: B. E. Hammel, M. H. Grayum, C. Herrera & N. Zamora (eds), Manual de Plantas de Costa Rica. Vol. VIII. Monogr. Syst. Bot. Missouri Bot. Gard. 131: 393 – 406.
  • Knapp, S. (2008). Lectotypification of Ruiz and Pavón’s names in Solanum (Solanaceae). Anales Jard. Bot. Madrid 65: 307 – 329.
  • Kobuski, C. E. (1938). Studies in Theaceae. III. Eurya subgenera Euryodes and Penteurya. Ann. Missouri Bot. Gard. 25: 299 – 359.
  • Kobuski, C. E. (1941). Studies in the Theaceae, VIII. A synopsis of the genus Freziera. J. Arnold Arbor. 22: 457 – 496.
  • Kobuski, C. E. (1942). Studies in the Theaceae XII. Notes on the South American species of Ternstroemia. J. Arnold Arbor. 23: 298 – 343.
  • Kobuski, C. E. (1951). Studies in the Theaceae, XXII. Some new species of Theaceae in South America. J. Arnold Arbor. 32: 152 – 154.
  • Kobuski, C. E. (1952). Theaceae. In: Contributions to the Flora of Venezuela II (J. A Steyermark and Collaborators). Fieldiana, Bot. 28: 380 – 384.
  • Kobuski, C. E. (1953). Studies in the Theaceae, XXVII. Miscellaneous new species in Theaceae. J. Arnold. Arbor. 34: 125 – 137.
  • Korthals, P. W. (1841). Verhandelingen over de natuurlijke geschiedenis der Nederlandsche overseesche Bezittingen, Botanie 4: 115, t. 7. Leiden.
  • Kuntze, O. (1891). Revisio Generum Plantarum 1: 1 – 374. A. Felix, Leipzig.
  • Lanjouw, J. & Stafleu, F. A. (1954). Index herbariorum, part 2, 1. Regnum Veg. Utrecht.
  • Li, W., Fu R. & Dickinson R. E. (2006). Rainfall and its seasonality over the Amazon in the 21st century as assessed by the coupled models for the IPCC AR4. J. Geophys. Res. 111: D02111, DOI:10.1029/2005JD006355.
  • Lingelsheim, A. (1909). In: A. Lingelsheim, F. Pax & H. Winkler, Plantae novae bolivianae. II. Repert. Spec. Nov. Regni Veg. 7: 107 – 114.
  • Little, E. L. (1969). New tree species from Esmeraldas, Ecuador (continued). Phytologia 18: 457 – 472.
  • Macbride, J. F. (1956). Theaceae. Flora of Peru. Publ. Field Mus. Nat. Hist., Bot. Ser. 13: 726 – 741.
  • Melchior, H. (1925). Theaceae. In: A. Engler & K. Prantl (eds), Die natürlichen Pflanzenfamilien (2nd edn). pp. 109 – 154. Engelmann, Leipzig.
  • Merello, M. (2011). Pentaphylacaceae. In: A. Idárraga-Piedrahita, R. D. C. Ortiz, R. Callejas Posada & M. Merello (eds), Flora de Antioquia: Catálogo de las Plantas Vasculares, Listado de las plantas vasculares del departamento de Antioquia, pp. 745 – 746. Series Biodiversidad y Recursos Naturales. Universidad de Antioquia, Missouri Botanical Garden & Oficina de planeación departamental de la gobernación de Antioquia, Editorial D’Vinni, Bogotá.
  • Monteagudo Mendoza, A. L., Valenzuela Gamarra, L., Vásquez Martínez, R., del Pilar Rojas Gonzáles, R., Phillips, O. L. & Lopez-Gonzalez, G. (2014). Primer catálogo de los árboles y afines de la Reserva Comunal El Sira, Perú/ First Catalogue of Tree species of the Communal Reserve Sira, Peru. Arnaldoa 21: 127 – 164.
  • Nee, M. H. (2008). Flora de la Región del Parque Nacional Amboró, Bolivia, Dillenidae 3: 1 – 255. Fundación Amigos de la Naturaleza, Santa Cruz.
  • Pittier, H. (1922). New or noteworthy plants from Colombia and Central America 8/ Theaceae. A new species of Eurya from Panama. Contr. U.S. Natl. Herb. 20: 453 – 490.
  • Pool, A. (2001). Theaceae. In: W. D. Stevens, C. Ulloa, A. Pool & O. M. Montiel (eds), Flora de Nicaragua. Monogr. Syst. Bot. Missouri Bot. Gard. 85: 2443 – 2448.
  • Ruiz, H. & Pavón, J. (1794.) Florae Peruvianae, et Chilensis Prodromus 1 – 150. Imprenta de Sancha, Madrid.
  • Ruiz, H. & Pavón, J. (1958). Flora Peruviana, et Chilensis, sive descriptions, et icones plantarum Peruvianarum, et Chilensium, secundum systema Linnaeanum digestae, cum characteribus plurium generum evulgatorum reformatis. Anales Inst. Bot. Cavanilles 5: 355 – 462.
  • Rusby, H. H. (1893). On the collections of Mr. Miguel Bang in Bolivia. Mem. Torrey Bot. Club 3: 1 – 67.
  • Rusby, H. H. (1907). An enumeration of the plants collected in Bolivia by Miguel Bang. Part 4. With descriptions of new genera and species. Bull. New York Bot. Gard. 4: 309 – 470.
  • Sandwith, N. Y. (1943). Freziera microphylla Sandwith, Theaceae, Tribus Ternstroemieae. Hooker's Icon. Pl. 35, t. 3450.
  • Santamaría-Aguilar, D. & Lagomarsino, L. P. (2015). Cuatro nuevas especies y un nuevo registro de Freziera (Pentaphylacaceae) de Ecuador and Perú. J. Bot. Res. Inst. Texas 9: 89 – 102.
  • Santamaría-Aguilar, D. & Lagomarsino, L. P. (2017). Two new Pentaphylacaceae species from southern Central America. Brittonia 69: 100 – 108.
  • Santamaría-Aguilar, D. & Ortiz, R. D. C. (2016). Freziera dasycarpa (Pentaphylacaceae), a new species from the montane forest of Antioquia, Colombia. Phytotaxa 263: 279 – 285.
  • Santamaría-Aguilar, D. (2015). Validation of two names and the description of a new species of Freziera (Pentaphylacaceae) from the Peruvian Andes. Harvard Pap. Bot. 20: 69 – 76.
  • Santamaría-Aguilar, D., Fuentes, A. F. & Lagomarsino, L. P. (2018). Three new species of Freziera (Pentaphylacaceae, Freziereae) from Bolivia and Peru. Phytotaxa 349: 101 – 114.
  • Santamaría-Aguilar, D., Monro, A. K., Jiménez-Madrigal, Q. & Lagomarsino, L. P. (2014). Two new species of Freziera (Pentaphylacaceae) from Costa Rica. Harvard Pap. Bot. 19: 219 – 225.
  • Santamaría-Aguilar, D., Neill, D. A. & Lagomarsino, L. P. (2016). A new species of Freziera (Pentaphylacaceae) from Ecuador with large leaves and flowers. Edinburgh J. Bot. 73: 292 – 296.
  • Shorthouse, D. P. (2010). SimpleMappr, an online tool to produce publication-quality point maps. Retrieved from http://www.simplemappr.net. Accessed July 2018.
  • Standley, P. C. (1931). Studies of American plants V. Publ. Field Mus. Nat. Hist., Bot. Ser. 8: 295 – 398.
  • Swenson J. J., Carter C. E., Domec J-C. & Delgado C. I. (2011). Gold mining in the Peruvian Amazon: global prices, deforestation, and mercury imports. PloS ONE 6: e18875.
  • Szyszylowicz, I. V. (1893). Theaceae. In: H. G. A. Engler & K. Prantl (eds), Die Natürlichen Pflanzenfamilien 3 (6): 157 – 192. Engelmann, Leipzig.
  • Triana, J. & Planchon, J. E. (1862). Prodromus Florae Novo-Granaten ou énumeration des plantes de la Nouvelle-Grenade, avec description des espécies nouvelles. Ann. Sci. Nat., Bot., sér. 4, 18: 258 – 381.
  • Tropicos Database (2017). Available from: tropicos.org (last accessed: 11 Nov. 2017).
  • Tulasne, R. L. (1847). Flore de la Colombie. Plantes Nouvelles. Ann. Sci. Nat., Bot. sér. 3. 8: 326 – 343.
  • Ulloa Ulloa, C., Acevedo-Rodríguez, P., Beck, S., Belgrano, M. J., Bernal, R., Berry, P. E., Brako, L., Celis, M., Davidse, G., Forzza, R. C., Gradstein, S. R., Hokche, O., León, B., León-Yánez, S., Magill, R. E., Neill, D. A., Nee, M., Raven, P. H., Stimmel, H., Strong, M. T., Villaseñor, J. L., Zarucchi, J. L., Zuloaga, F. O. & Jørgensen, P. M. (2017). An integrated assessment of the vascular plant species of the Americas. Science 358: 1614 – 1617.
  • Urban, I. (1896a). Additamenta ad cognitionem Florae Indiae occidentalis. Particula III. Bot. Jahrb. Syst. 514 – 638.
  • Urban, I. (1896b). Patascoya, eine neue Ternstroemiaceen-Gattung. Ber. Deutsch. Bot. Ges. 14: 282 – 283.
  • Villaseñor Ríos, J. L. (2016). Checklist of the native vascular plants of Mexico/ Catálogo de las plantas vasculares nativas de México. Revista Mex. Biodivers. 87: 559 – 902.
  • Wawra von Fernsee, H. (1886). Ternstroemiaceae. In: C. F. P. von Martius (ed.), Flora Brasiliensis 12: 261 – 334. Monachii & Lipsiae [Munich & Leipzig]
  • Weitzman, A. L. (1987a). Systematics of Freziera Willd. (Theaceae). Harvard University, Cambridge, Massachusetts, Ph.D. Thesis.
  • Weitzman, A. L. (1987b). Taxonomic studies in Freziera (Theaceae), with notes on reproductive biology. J. Arnold Arbor. 68: 323 – 334.
  • Weitzman, A. L. (1999). Theaceae. In: P. M. Jørgensen & S. León-Yánez (eds), Catalogue of the vascular plants of Ecuador. Monogr. Syst. Bot. Missouri Bot. Gard. 75: 923 – 924.
  • Weitzman, A. L., Dressler, S. & Stevens, P. F. (2004). Ternstroemiaceae. In: K. Kubitzki (ed.), The Families and Genera of Vascular Plants. Flowering Plants. Dicotyledons. Vol. 6: 450 – 460. Springer-Verlag, Berlin.
  • Zorzanelli, J. P. F., Carrijo, T. T., Fiaschi, P., Jardim, J. G., Santamaría-Aguilar, D. & Amorim, A. M. (2015). A first record of Freziera (Pentaphylacaceae) from the Brazilian Atlantic Forest, with the description of a new species. Syst. Bot. 40: 1075 – 1080.
  • van der Werff, H. (1993). Theaceae. In: L. Brako & J. L. Zarucchi (eds), Catalogue of the flowering plants and Gymnosperms of Peru/Catálogo de las Angiospermas y Gimnospermas del Peru. Monogr. Syst. Bot. Missouri Bot. Gard. 45: 1144 – 1146.

Kew Backbone Distributions
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

Kew Bulletin
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Kew Names and Taxonomic Backbone
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0