1. Family: Piperaceae Giseke
    1. Genus: Piper L.
      1. Piper guineense Schumach. & Thonn.

        This species is accepted, and its native range is Tropical Africa, Madagascar.

    [FWTA]

    Piperaceae, Hutchinson and Dalziel. Flora of West Tropical Africa 1:1. 1954

    Habit
    A climber, 30–40 ft. long, on trees
    Fruits
    Fruit red or red-brown when ripe, black when dry.
    [FTEA]

    Piperaceae, Bernard Verdcourt. Flora of Tropical East Africa. 1996

    Habit
    Dioecious usually ± glabrous liane, 4–20(–23) m. tall, climbing into trees by means of adventitious roots near the nodes; stems glabrous or rarely pubescent; older stems with corky ridges.
    Leaves
    Leaves alternate, very variable according to position, round to elliptic, ovate or lanceolate, 4.5–20.5 cm. long, 2–13 cm. wide, acuminate at the apex, rounded to cordate at the base or cuneate, pubescent on nerves beneath or entirely glabrous, rarely pubescent all over; petiole 0.5–4 cm. long; nervation variable, the nerves palmate from at or near base, and usually additional pinnate ones.
    Flowers
    Flowers yellow or yellow-green in terminal or lateral leaf-opposed spikes, 2–3 cm. long, attaining 4–10 cm. (rarely more) in fruit; peduncle glabrous or slightly pubescent, 0.5–1(–2.5) cm. long; bracts round or obovate, peltate, ± 1 mm. in diameter, ciliate at margin, pubescent beneath.
    Stamens
    Stamens 2(–3).
    Pistil
    Ovary ovoid; stigmas 3–5.
    Fruits
    Fruits mostly red or reddish brown but also orange or yellow to buff, globose to obovoid, 3–6 mm. in diameter on pedicels 0.5–1 cm. long.
    Habitat
    Evergreen forest and forest edges, usually in wet places, gallery forest along rocky rivers; 750–1650 m.
    Distribution
    Mali and Guinea Bissau to Angola, Zaire, Zambia and Sudan K5 T1 T4 U2 U4

    Images

    Distribution

    Native to:

    Angola, Benin, Burundi, Cabinda, Cameroon, Central African Repu, Congo, Ethiopia, Gabon, Gambia, Ghana, Guinea, Guinea-Bissau, Gulf of Guinea Is., Ivory Coast, Kenya, Liberia, Madagascar, Mali, Nigeria, Sierra Leone, Sudan, Tanzania, Togo, Uganda, Zambia, Zaïre

    Piper guineense Schumach. & Thonn. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Oct 1, 2008 Onana, J.-M. [3601], Cameroon K000580040
    Jul 1, 2007 Tchiengue, B. [1895], Cameroon K000461038
    Mar 1, 2006 Cheek, M. [11106], Cameroon K000339021
    Mar 1, 2006 Etuge, M. [5152], Cameroon K000339020
    Jan 1, 2003 Lane, P. [299], Cameroon K000180218
    Jan 1, 2003 Wheatley, J.I. [423], Cameroon K000180279
    Jan 1, 2003 Cable, S. [3242], Cameroon K000180230
    Jan 1, 2003 Kenfack, D. [305], Cameroon K000180225
    Jan 1, 2003 Etuge, M. [4667], Cameroon K000180221
    Jan 1, 2003 Etuge, M. [2605], Cameroon K000180224
    Jan 1, 2003 Etuge, M. [4298], Cameroon K000180226
    Jan 1, 2003 Etuge, M. [2762], Cameroon K000180232
    Jan 1, 2003 Groves, M. [27], Cameroon K000180219
    Jan 1, 1994 Harris, D.J. [3850], Cameroon K000381421
    Jan 1, 1994 Ndam, N. [722], Cameroon K000381422
    Jan 1, 1994 Akogo, M. [24], Cameroon K000381423
    Jan 1, 1987 Etuge, M. [235], Cameroon K000180070
    May 1, 1976 Letouzey, R. [14843], Cameroon K000180072
    Jan 1, 1976 Letouzey, R. [14844], Cameroon K000180073
    Milne-Redhead, E. [4487], Zambia 18140.000
    Cheek, M. [7513], Cameroon K000180194
    Etuge, M. [1403], Cameroon K000180195
    Etuge, M. [4151], Cameroon K000180228
    illegible [s.n.], Sierra Leone K000350306
    Vogel, T. [97], Sierra Leone K000350307 Unknown type material
    Preuss, P.R. [417], Cameroon K000180074
    Thompson, S.A. [1367], Cameroon K000180069
    Ghogue, J.-P. [485], Cameroon K000180223

    First published in Beskr. Guin. Pl.: 19 (1827)

    Accepted by

    • Sosef, M.S.M. & al. (eds.) (2017). Flore du Gabon 50: 1-77. Muséum National d'Histoire Naturelle, Paris.
    • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
    • Figueiredo, E., Paiva, J., Stévart, T., Oliveira, F. & Smith, G.F. (2011). Annotated catalogue of the flowering plants of São Tomé and Príncipe Bothalia, A Journal of Botanical Research 41: 41-82.
    • Fischer, E., Rembold, K., Althof, A. & Obholzer, J. (2010). Annotated checklist of the vascular plants of Kakamega forest, Western province, Kenya Journal of East African Natural History 99: 129-226.
    • Lejoy, J., Ndjele, M.-B. & Geerinck, D. (2010). Catalogue-flore des plantes vasculaires des districts de Kisangani et de la Tshopo (RD Congo) Taxonomania 30: 1-307.
    • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
    • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
    • Edwards, S., Tadesse, M., Demissew, S. & Hedberg, I. (eds.) (2000). Flora of Ethiopia and Eritrea 2(1): 1-532. The National Herbarium, Addis Ababa University, Ethiopia & The Department of Systematic Botany, Upps.
    • Pope, G.V. (ed.) (1997). Flora Zambesiaca 9(2): 1-64. Royal Botanic Gardens, Kew.
    • Verdcourt, B. (1996). Flora of Tropical East Africa, Piperaceae: 1-24.
    • Jones, M. (1991). A checklist of Gambian plants: 1-33. Michael Jones, The Gambia College.
    • Brunel, J.F., Hiepo, P. & Scholz, H. (eds.) (1984). Flore Analytique du Togo Phanérogames: 1-751. GTZ, Eschborn.
    • Lewalle, J. (1970). Liste floristique et répartition altitudinale de la flore du Burundi occidental: 1-84. Université officielle de Bujumbura.
    • Robyns, W. & al. (eds.) (1948-1963). Flore du Congo Belge et du Ruanda-Urundi 1-10.

    Literature

    Flora of West Tropical Africa
    • Chev. Fl. Viv. 1: 139.
    • —F.T.A. 6, 1: 145
    Kew Backbone Distributions
    • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
    • Figueiredo, E., Paiva, J., Stévart, T., Oliveira, F. & Smith, G.F. (2011). Annotated catalogue of the flowering plants of São Tomé and Príncipe Bothalia, A Journal of Botanical Research 41: 41-82.
    • Fischer, E., Rembold, K., Althof, A. & Obholzer, J. (2010). Annotated checklist of the vascular plants of Kakamega forest, Western province, Kenya Journal of East African Natural History 99: 129-226.
    • Lejoy, J., Ndjele, M.-B. & Geerinck, D. (2010). Catalogue-flore des plantes vasculaires des districts de Kisangani et de la Tshopo (RD Congo) Taxonomania 30: 1-307.
    • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
    • Sita, P. & Moutsambote, J.-M. (2005). Catalogue des plantes vasculaires du Congo, ed. sept. 2005: 1-158. ORSTOM, Centre de Brazzaville.
    • Jones, M. (1991). A checklist of Gambian plants: 1-33. Michael Jones, The Gambia College.
    • Brunel, J.F., Hiepo, P. & Scholz, H. (eds.) (1984). Flore Analytique du Togo Phanérogames: 1-751. GTZ, Eschborn.
    • Lewalle, J. (1970). Liste floristique et répartition altitudinale de la flore du Burundi occidental: 1-84. Université officielle de Bujumbura.
    Flora of Tropical East Africa
    • Agnew, A.D.Q. & Agnew, S., Upland Kenya Wild Flowers, ed. 2: 48 (1994).
    • Beentje, Kenya Trees, Shrubs and Lianas p. 66 (1994).
    • Blundell, Wild Fl. E. Afr., reprint: 22 (1992).
    • Hepper, W. Afr. Herb. Isert & Thonning: 103 (1976).
    • F. White, Forest Flora of Northern Rhodesia p. 19 (1962).
    • Keay in Flora of West Tropical Africa, ed. 2, 1: 84 (1954).
    • F. W. Andr., The Flowering Plants of the Anglo-Egyptian Sudan 1: 26, fig. 20 (1950).
    • Balle in Flore du Congo Belge et du Ruanda-Urundi, 1: 17 (1948).
    • Balle in Bulletin du Jardin Botanique de l'État a Bruxelles 16: 375, t. 16 (1942) (much extra synonymy and information)
    • A. Engler & O. Drude, Die Vegetation Der Erde, IX, Pflanzenwelt Afrikas 3(1): 3, fig. 2A–D (1915).
    • Thonner, Fl. Pl. Afr.: 161, t. 27 (1915).
    • Engl. in Wissenschaftliche Ergebnisse der Deutschen Zentral-Afrika-Expedition 1907-1908, 2 (Botanik) p. 177 (1911).
    • Baker & C.H. Wright in Flora of Tropical Africa 6(1): 145 (1909).
    • Hiern, Cat. Afr. Pl. Welw. 1: 912 (1900).
    • C. DC. in DC., Prodr. 16(1): 343 (1869).
    • Thonn. in Schumach., Beskr. Guin. Pl.: 19 (1827).

    Sources

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of West Tropical Africa
    Flora of West Tropical Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0