1. Family: Rhamnaceae Juss.
    1. Genus: Gouania Jacq.
      1. Gouania longispicata Engl.

        This species is accepted, and its native range is Benin to Ethiopia and S. Tropical Africa.

    [FZ]

    Rhamnaceae, R. B. Drummond. Flora Zambesiaca 2:2. 1966

    Habit
    Climbing shrub or liane; branchlets with coiled tendrils; tendrils rusty-pubescent when young, glabrescent later.
    Leaves
    Leaf-lamina up to 8·5 × 7 cm., ovate, apex acute to mucronate, margin closely serrulate, base rounded to cordate, tomentellous to nearly glabrous above, persistently tomentose beneath, secondary nerves in 5–9 pairs impressed above and prominent below, tertiary venation prominent and regularly subparallel beneath; petiole tomentose, up to 2·5 cm. long; stipules up to 1 cm. long, subulate, tomentose, caducous.
    Inflorescences
    Inflorescence up to 50 cm. long, of elongated racemes up to 20 cm. long and composed of fascicles of up to 20 flowers; pedicels up to 5 mm. long, tomentellous; bracts up to 8 mm. long, triangular, tomentose.
    Receptacle
    Receptacle tomentose.
    Calyx
    Sepals 1·5–2 mm. long, deltate, tomentose outside, glabrous within.
    Corolla
    Petals 1·5 mm. long, obovate, cucullate, shortly unguiculate.
    Stamens
    Stamens 1·5 mm. long.
    Disc
    Disk 3 mm. in diam., lobed opposite the calyx-teeth.
    Pistil
    Ovary inferior: style 0–5 mm. long, 3-fid or very short and obscurely 3-lobed.
    Fruits
    Fruit 6 × 11 mm., 3-winged, pubescent, glabrescent with age; wings very loosely reticulate-veined.
    Seeds
    Seed 3 × 2 mm., plano-convex; testa shining brown.
    [FTEA]

    Rhamnaceae, Marshall C. Johnston (University of Texas Herbarium). Flora of Tropical East Africa. 1972

    Habit
    Climbing or sprawling shrubs or lianes several m. long.
    Branches
    Branchlets and tendrils with longitudinal pubescent stripes of antrorse brownish hairs 0.2–0.3 mm. long, glabrescent, very slightly zig-zag.
    Leaves
    Leaf-blades ovate, 4–8.5 cm. long, 2.5–4.5(–7) cm. wide, round at base, acute or acuminate (acumen 5–10 mm. long), serrulate or doubly serrulate, above dark and with sparse antrorse golden-brown hairs 0.2–0.4 mm. long, beneath with a dense persistent grey or brown tomentum, on each side of midrib with 6–7 secondary nerves; petioles 10–25 mm. long, ventrally pubescent.
    Stipules
    Stipules subulate, ± 3 mm. long (rarely to 10 mm.).
    Inflorescences
    Thyrses merely terminal or aggregated into loose panicles; cymes 3–10-flowered; peduncles of cymes 1 mm. long and pubescent or absent; pedicels 2–3 mm. long, pubescent.
    Flowers
    Flowers whitish or greenish white, fragrant and nectariferous.
    Hypanthium
    Cup ± 2 mm. wide, externally pubescent.
    Calyx
    Sepals ± 1 mm. long.
    Corolla
    Petals ± 1 mm. long.
    Style
    Style ± 0.5 mm. long.
    Fruits
    Fruit 6–8 mm. long, 8–11 mm. wide.
    Figures
    Fig. 4.
    Habitat
    Forests, particularly at margins and in disturbed places, riverine thickets and wooded grassland; 300–2400 m.
    Distribution
    K3 K4 K5 K7 T2 T3 T4 T6 T7 T8 U1 U2 U3 U4
    [FWTA]

    Rhamnaceae, Hutchinson and Dalziel. Flora of West Tropical Africa 1:2. 1958

    Habit
    A scandent shrub or liane
    Leaves
    Leaves greyish beneath
    Flowers
    Flowers greenish-yellow
    Ecology
    In montane forest.

    Distribution

    Native to:

    Benin, Burundi, Cameroon, Ethiopia, Kenya, Malawi, Mozambique, Nigeria, Rwanda, Sudan, Tanzania, Uganda, Zaïre, Zimbabwe

    Gouania longispicata Engl. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Oct 1, 2008 Corcoran, M.R. [9], Cameroon K000580210
    Jan 1, 2004 Tadjouteu, F. [411], Cameroon K000198491
    Jul 1, 2002 Cheek, M. [9616], Cameroon K000027201
    Jul 1, 2002 Ghogue, J.-P. [26], Cameroon K000027200
    Jul 1, 2002 Barlow, L. [15], Cameroon K000027199
    Dec 1, 2000 Etuge, M. [3462], Cameroon K000337646
    Dec 1, 2000 Etuge, M. [2339], Cameroon K000337651
    Dec 1, 2000 Buzgo, M. [697], Cameroon K000337648
    Dec 1, 2000 Buzgo, M. [697], Cameroon K000337649
    Dec 1, 2000 Kamundi, D. [619], Cameroon K000337645
    Feb 1, 1999 Onana, J.-M. [527 A], Cameroon K000337647
    Mar 1, 1997 Munyenyembe, P. [806], Cameroon K000337650

    First published in Pflanzenw. Ost-Afrikas, C: 256 (1895)

    Accepted by

    • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
    • Fischer, E., Rembold, K., Althof, A. & Obholzer, J. (2010). Annotated checklist of the vascular plants of Kakamega forest, Western province, Kenya Journal of East African Natural History 99: 129-226.
    • Timberlake, J.R., Bayliss, J., Alves, T., Francisco, J., Harris, T., Nangoma, D. & de Sousa, C. (2009). Biodiversity and Conservation of Mchese Mountain, Malawi. Report produced under the Darwin Initiative Award 15/036: 1-71. Royal Botanic Gardens, Kew.
    • Govaerts, R. (2003). World Checklist of Seed Plants Database in ACCESS G: 1-40325.
    • Hedberg, I. & Edwards, S. (eds.) (1989 publ. 1990). Flora of Ethiopia and Eritrea 3: 1-659. The National Herbarium, Addis Ababa University, Ethiopia & The Department of Systematic Botany, Upps.
    • Johnston, M.C. (1972). Flora of Tropical East Africa, Rhamnaceae: 1-40.
    • Exell, A.W., Fernandes, A. & Wild, H. (eds.) (1966). Flora Zambesiaca 2(2): 353-653. Royal Botanic Gardens, Kew.
    • Hutchinson, J., Dalziel, J.M. & Keay, R.W.J. (1954-1958). Flora of West Tropical Africa, ed. 2, 1: 1-828.

    Literature

    Flora of West Tropical Africa
    • Staner in Bull. Jard. Bot. Brux. 15: 421, fig. 60.
    • M. L. Green in Kew Bull. 1916: 198
    • Pflanzenw. Ost-Afr. C: 256 (1895)
    Kew Backbone Distributions
    • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
    • Fischer, E., Rembold, K., Althof, A. & Obholzer, J. (2010). Annotated checklist of the vascular plants of Kakamega forest, Western province, Kenya Journal of East African Natural History 99: 129-226.
    Flora of Tropical East Africa
    • R.B. Drummond in Flora Zambesiaca 2: 434, t. 88/C (1966).
    • Cufod., Enumeratio Plantarum Aethiopiae Spermatophyta (Supplement in Bull. Jard. Bot. Brux.) p. 502 (1960).
    • Evrard in Flore du Congo Belge et du Ruanda-Urundi, 9: 449 (1960).
    • Keay in Flora of West Tropical Africa, ed. 2, 1: 670 (1958).
    • Verdc. in Bulletin du Jardin Botanique de l'État a Bruxelles 27: 359 (1957).
    • Brenan, in Mem. N.Y. Bot. Card. 8: 239 (1953).
    • Suesseng. in A. Engler & K. Prantl, Die Natürlichen Pflanzenfamilien. ed. 2, 20d: 168 (1953).
    • J.P.M. Brenan, Check-lists of the Forest Trees and Shrubs of the British Empire no. 5, part II, Tanganyika Territory p. 466 (1949)
    • Eyles in Trans. Roy. Soc. S. Afr. 5: 407 (1916).
    • M.L. Green in Bulletin of Miscellaneous Information, Kew 1916: 198 (1916).
    • Bak. f. in Journal of the Linnean Society of London, Botany 40: 45 (1911).
    • Engl., Die Pflanzenwelt Ost-Afrikas und der Nachbargebiete, Theile C: 256 (1895).

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of West Tropical Africa
    Flora of West Tropical Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2020. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2020. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0