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  1. Family: Rubiaceae Juss.
    1. Genus: Spermacoce L.
      1. Spermacoce pusilla Wall.

        This species is accepted, and its native range is Indian Subcontinent to S. China and Philippines.

    [FTEA]

    Rubiaceae, B. Verdcourt. Flora of Tropical East Africa. 1976

    Habit
    Erect or rarely prostrate annual usually branched herb (2–)7.5–60 cm. tall, with grooved ± glabrous or slightly papillate-puberulous often reddish stems.
    Leaves
    Leaf-blades linear-lanceolate to narrowly lanceolate, 1–5.3 cm. long, 2–5.5 mm. wide, acute at the apex, narrowed to the base, minutely scabrid above, glabrous beneath; true petiole absent; stipule-sheath 1.5–2 mm. long, glabrous or pubescent, 1.5–2 mm. long, bearing 7 setae 2–4 mm.
    Flowers
    Flowers in dense very compact spherical clusters at most nodes, 0.6–1(–1.5) cm. in diameter; bracteoles filiform, numerous, 2 mm. long.
    Calyx
    Calyx-tube ovoid, 1 mm. long, pubescent; teeth 4, equal or slightly unequal, subulate, 0.6–1.2 mm. long, scabrid.
    Corolla
    Corolla white or pink; tube narrowly funnel-shaped, 1.3 mm. long; lobes 0.8–1.1 mm. long, 0.4 mm. wide, with some long flattened hairs at the apex.
    Filaments
    Filaments exserted ± 1 mm.
    Style
    Style exserted 0.5 mm.
    Fruits
    Capsule ellipsoid, 1.5 mm. long, glabrous or ± sparsely pubescent.
    Seeds
    Seeds chestnut-brown, shiny, oblong-ellipsoid or narrowly oblong, 1.3 mm. long, 0.55 mm. wide, 0.4 mm. thick, with a wide ventral groove.
    Figures
    Fig. 50/13, p. 342.
    Habitat
    Grassland, grassland with scattered trees, woodland, roadsides, stony areas, rocky places, also as a weed of cultivation; (150–)450–1920 m.
    Distribution
    widespread in tropical Africa, Madagascar and tropical Asia to Japan and Philippines K3 K4 K5 K6 T1 T3 T4 T6 T7 T8 U1 U2 U3 U4
    [FZ]

    Rubiaceae, B. Verdcourt. Flora Zambesiaca 5:1. 1989

    Habit
    Erect or rarely prostrate annual usually branched herb (2)7.5–60 cm. tall, with grooved ± glabrous or slightly papillate-puberulous often reddish stems.
    Leaves
    Leaf blades 1–5.3 cm. x 2–5.5 mm., linear-lanceolate to narrowly lanceolate, acute at the apex, narrowed to the base, minutely scabrid above, glabrous beneath; true petiole absent; stipule sheath 1.5–2 mm. long, glabrous or pubescent, bearing 7 setae 2–4 mm. long.
    Flowers
    Flowers in dense very compact spherical clusters at most nodes, 0.6–1(1.5) cm. in diam.; bracteoles filiform, numerous, 2 mm. long.
    Calyx
    Calyx tube 1 mm. long, ovoid, pubescent; teeth 4, equal or slightly unequal, 0.6–1.2 mm. long, subulate, scabrid.
    Corolla
    Corolla white or pink; tube 1.3 mm. long, narrowly funnel-shaped; lobes 0.8–1.1 x 0.4 mm. with some long flattened hairs at the apex.
    Filaments
    Filaments exserted c. 1 mm.
    Style
    Style exserted 0.5 mm.
    Fruits
    Capsule 1.5 mm. long, ellipsoid; glabrous or ± sparsely pubescent.
    Seeds
    Seeds chestnut-brown, shiny, 1.3 x 0.55 x 0.4 mm., oblong-ellipsoid or narrowly oblong, with a wide ventral groove.

    Images

    Distribution

    Doubtfully present in:

    Borneo, Sumatera

    Native to:

    China South-Central, China Southeast, East Himalaya, Hainan, India, Malaya, Myanmar, Nepal, Nicobar Is., Pakistan, Philippines, Sri Lanka, Taiwan, Thailand, Vietnam, West Himalaya

    Introduced into:

    Angola, Benin, Burkina, Burundi, Cameroon, Central African Repu, Chad, Congo, Ethiopia, Gabon, Gambia, Guinea, Guinea-Bissau, Ivory Coast, Kenya, Liberia, Madagascar, Nigeria, Rwanda, Senegal, Sudan, Tanzania, Uganda, Zambia, Zaïre

    Synonyms

    Other Data

    Spermacoce pusilla Wall. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Feb 1, 2004 Etuge, M. [4731], Cameroon K000746452
    Feb 1, 2004 Pollard, B.J. [703], Cameroon K000746453
    Feb 1, 2004 Onana, J.-M. [1882], Cameroon K000746451
    May 1, 2002 Cheek, M. [9421], Cameroon K000026347
    Jan 1, 2000 Munyenyembe, P. [903], Cameroon K000338737
    Bidgood, S. [5167], Tanzania K000190395
    Teijsmann, J.E. [s.n.], Sumatera K000772200
    Wallich, N. [Cat. no. 823], Nepal K001111984
    Wallich, N. [Cat. no. 823], Nepal K001111985

    Bibliography

    First published in W.Roxburgh, Fl. Ind. 1: 379 (1820)

    Accepted by

    • Balkrishna, A. (2018). Flora of Morni Hills (Research & Possibilities): 1-581. Divya Yoga Mandir Trust.
    • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
    • Thiombiano, A., Schmidt, M., Dressler, S., Ouédraogo, A., Hahn, K. & Zizka, G. (2012). Catalogue des plantes vasculaires du Burkina Faso Boissiera 65: 1-391.
    • Kuo, M.L. (ed.) (2012). Flora of Taiwan, ed. 2, Suppl.: 1-414. Editorial Committee of the Flora of Taiwan, Second Edition, National Taiwan University.
    • Fischer, E., Rembold, K., Althof, A. & Obholzer, J. (2010). Annotated checklist of the vascular plants of Kakamega forest, Western province, Kenya Journal of East African Natural History 99: 129-226.
    • Lisowski, S. (2009). Flore (Angiospermes) de la République de Guinée Scripta Botanica Belgica 41: 1-517.
    • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
    • Sosef, M.S.M. & al. (2006). Check-list des plantes vasculaires du Gabon Scripta Botanica Belgica 35: 1-438.
    • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
    • Catarino, L., Sampaio Martins, E., Pinto-Basto, M.F. & Diniz, M.A. (2006). Plantas Vasculares e Briófitos da Guiné-Bissau: 1-298. Instituto de investigação científica tropical, Instituto Português de apoio ao desenvolvimento.
    • Govaerts, R. (2003). World Checklist of Selected Plant Families Database in ACCESS: 1-216203. The Board of Trustees of the Royal Botanic Gardens, Kew.

    Literature

    Kew Backbone Distributions

    • Jongkind, C. (2014). Fauna & Flora of Liberia, flowering plant species list www.liberianfaunaflora.org.
    • Thiombiano, A., Schmidt, M., Dressler, S., Ouédraogo, A., Hahn, K. & Zizka, G. (2012). Catalogue des plantes vasculaires du Burkina Faso Boissiera 65: 1-391.
    • Wu, Z., Raven, P.H. & Hong, D. (eds.) (2011). Flora of China 19: 1-884. Missouri Botanical Garden Press, St. Louis.
    • Fischer, E., Rembold, K., Althof, A. & Obholzer, J. (2010). Annotated checklist of the vascular plants of Kakamega forest, Western province, Kenya Journal of East African Natural History 99: 129-226.
    • Lisowski, S. (2009). Flore (Angiospermes) de la République de Guinée Scripta Botanica Belgica 41: 1-517.
    • Pandey, R.P. & Dilwakar, P.G. (2008). An integrated check-list flora of Andaman and Nicobar islands, India Journal of Economic and Taxonomic Botany 32: 403-500.
    • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
    • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
    • Catarino, L., Sampaio Martins, E., Pinto-Basto, M.F. & Diniz, M.A. (2006). Plantas Vasculares e Briófitos da Guiné-Bissau: 1-298. Instituto de investigação científica tropical, Instituto Português de apoio ao desenvolvimento.
    • Ehrendorfer, F., Schönbeck-Temesy, E., Puff, C. & Rechinger, W. (2005). Rubiaceae Flora Iranica 176: 1-287. Naturhistorisches Museums Wien.
    • Ruhsam, M in Ruhsam, M. (2005). Survey of all the Rubiaceae specimens at the Royal Botanic Gardens, Kew.
    • Sita, P. & Moutsambote, J.-M. (2005). Catalogue des plantes vasculaires du Congo, ed. sept. 2005: 1-158. ORSTOM, Centre de Brazzaville.
    • H?, P.-H. (2000). Câyc? Vi?tnam. An Illustrated flora of Vietnam, ed. 2, 3: 1-1020. Pham-hoang Ho, Montréal.
    • Jones, M. (1991). A checklist of Gambian plants: 1-33. Michael Jones, The Gambia College.
    • Lebrun, J.P. (1973). Énumération des plantes vasculaires du Sénégal: 1-209. Maisons Alfort: Institut d'élevage et de médecine vétérinaire des pays tropicaux.
    • Lebrun, J.-P., Audru, J., Gaston, A. & Mosnier, M. (1972). Catalogue des Plantes Vasculaires du Tchad Méridional: 1-289. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.

    Flora of Tropical East Africa

    • Wall. in Roxb., Fl. Indica, ed. Carey & Wall. 1: 379 (1820).

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2020. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2020. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0