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This species is accepted, and its native range is Bolivia to Brazil and Paraguay.


J. Prado. (2005). A New Species and Hybrid of Adiantum (Pteridaceae) from South America. Kew Bulletin, 60(1), 117-121. Retrieved from

Bolivia, known only from Pando. In Brazil, wide-ranging from north to south, occurring in the states of Acre, Amazonas, Pará, Mato Grosso, Pernambuco, Alagoas, Minas Gerais, Bahia, Espírito Santo and São Paulo.
Wet forests, 0 - 600 m.
Morphology General Habit
Plants terrestrial.
Morphology General Scales
Scales of two kinds, both appressed throughout and with apex spreading, concolorous, varying from brown or light brown to reddish brown, scales narrowly triangular with a filiform apex, strongly ciliate proximally and sparsely denticulate at margins, scales arachnoid, basally stellate, process narrow
Morphology Leaves
Fronds monomorphic, 2- pinnate, 22 - 45 cm long, the laminae (9.5 -)14 - 27 cm wide
Morphology Leaves Pinnae
Pinnae oblong-triangular, slightly decreasing at the base, tapering at the apex, (5 -)9 - 16 x 2 - 4.5 cm wide, the lateral pinnae 1 - 3 pairs, oblique, alternate, the terminal pinna similiar, 1 - 1.5 times longer than the subtending pinnae, 0.7 - 1 times as long as the median pinnae; indument of the costae like that of the stipes and rachises
Morphology Leaves Pinnules
Pinnules (5 -) 14 - 16(- 20) pairs, 2 - 2.5 times longer than wide, chartaceous, continuous to the rachis (not articulate), free-veined, with a more or less evident midrib, the proximal pairs reduced, somewhat rounded or triangular, the medial pairs dimidiate, oblong to somewhat quadrangular, the acroscopic base truncate, the sterile apex obtuse to acute, the sterile margins regularly serrate, the fertile apex angular, the distal pinnules c. 1/2 as long as the medial pinnules, the adaxial surface of the pinnules glabrous, the veins slightly prominulous, idioblasts present among the veins, the abaxial surface of the pinnules glaucous and with sparse light brown to reddish brown hairs, borne on the veins, the hairs simple, septate, 0.05 - 1 mm long, the veins slightly prominulous, the idioblasts among veins inconspicuous
Morphology Leaves Rachis
Rachises similar to the stipes and their indument similar but more congested
Morphology Leaves Stipes
Stipes 0.5 - 1 cm apart, 2/3 - 1/2 the length of the frond, black, adaxially sulcate, scaly
Morphology Reproductive morphology Sori
Sori arcuate to oblong, varying from (2 -)7 - 10 (- 14) per pinnule
Morphology Reproductive morphology Sori Indusium
Indusium glabrous, entire to erose
Morphology Reproductive morphology Spores
Spores trilete, 30 - 40 μm in equatorial diameter, tetrahedral-globose
Adiantum windischii belongs to the group of A. latifolium Lam. because it has long-creeping rhizomes, 2-pinnate laminae, and 1 - 3(4) pairs of lateral pinnae. It differs, however, by having two kind of scales (narrowly triangular with a filiform apex, and scales arachnoid, basally stellate, on the stipe and rachis (vs. mostly one kind of scales on A. latifolium - scales narrowly triangular), and pinnules abaxially with simple, septate, light brown to reddish brown hairs (vs. pinnules abaxially glabrate). The two species related to Adiantum windischii are A. argutum Splitg. and A. incertum Lindm. Adiantum argutum differs in having acuminate pinnules, abaxially with sparse, pectinate, minute setiform scales (Prado & Lellinger 2002). Adiantum incertum has pinnules with hairlike scales abaxially and with a few basal processes. The species epithet honours Dr Paulo G. Windisch, pteridologist at Universidade do Vale do Rio dos Sinos (UNISINOS).
Brazil. Espírito Santo: Mun. Linhares, 8 km W of Linhares on road to Colatina (on N side of Rio Doce), 19°22'S, 40°07'W, 22 Feb. 1988, W. W Thomas et al. 6091 (holotypus: SP; isotypi: NY n.v., SPF).
Vegetative Multiplication Rhizomes
Rhizomes long-creeping, 2 - 3 mm wide, scaly, the scales 1 - 2 mm long, somewhat shiny, essentially concolorous, appressed, medium or dark brown to reddish brown, narrowly triangular, sparsely denticulate at margins

Native to:

Bolivia, Brazil North, Brazil Northeast, Brazil Southeast, Brazil West-Central, Paraguay

Adiantum windischii J.Prado appears in other Kew resources:

First published in Kew Bull. 60: 119 (2005)

Accepted by

  • Roskov Y. & al. (eds.) (2018). World Ferns: Checklist of Ferns and Lycophytes of the World Species 2000 & ITIS Catalogue of Life Naturalis, Leiden, the Netherlands.


Kew Bulletin

  • Lellinger, D. B. (1989). The ferns and fern-allies of Costa Rica, Panama, and the Chocó (part 1: Psilotaceae through Dicksoniaceae). Pteridologia 2A: 1 - 364.
  • Mickel, J. T. & Beitel, J. M. (1988). Pteridophyte flora of Oaxaca, Mexico. Mem. New York Bot. Gard. 46: 1 - 568.
  • Moran, R. C. & Watkins, J. E. Jr. (2002). The occurrence and morphology of Adiantum x variopinnatum (Pteridaceae). Brittonia 54 (1): 57 - 60.
  • Prado, J. & Lellinger, D. B. (2002). Adiantum argutum, an unrecognized species of the A. latifolium group. Amer. Fern J. 92(1): 23 - 29.
  • Proctor, G. R. (1985). Ferns of Jamaica. British Museum (Natural History), London.
  • Smith, A. R. (1981). Pteridophytes. In: D. E. Breedlove (ed.), Flora of Chiapas. California Academy of Sciences, San Francisco.
  • Tryon, R. M. & Stolze, R. G. (1989). Pteridophyta of Peru, Part II. 13. Pteridaceae- 15. Dennstaedtiaceae. Fieldiana, Bot., n.s. 22: 1 - 128.
  • Walker, T. G. & Jermy, A. C. (1985). Descriptions of new species and hybrids and a new combination. Bull. Brit. Mus. (Nat. Hist.) 13: 251 - 276.
  • Zimmer, B. (1995). Grupo de Adiantum petiolatum. In: G. Davidse, M. Sousa & S. Knapp (eds.), Flora Mesoamericana, Universidad Nacional Autónoma de México, Ciudad de México, v. 1, pp. 110-113.

Kew Backbone Distributions
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at and
© Copyright 2017 World Checklist of Selected Plant Families.

Kew Bulletin
Kew Bulletin

Kew Names and Taxonomic Backbone
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at and
© Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families.