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  1. Family: Papaveraceae Juss.
    1. Genus: Corydalis DC.
      1. Corydalis devendrae Pusalkar

        This species is accepted, and its native range is W. Himalaya.


    Pusalkar, P.K. Kew Bull (2011) 66: 545.

    Perennial herb, 5 – 12 cm high (aerial part); roots tuberous; tubers two, vertical, unequal, larger (parental) up to 5.5 cm long, up to 6 mm thick; main stem usually present, sometimes absent, if present, slender, elongated with length depending on years of perennation, covered throughout with remains of sheathing petiole bases or naked in lower part; stem(s) 1 – few, simple or branched from the base, arising from the axis of basal leaves, glabrous or glaucous
    Cauline leaves similar, 2 – 3, smaller, narrower upwards, petioled below, sessile above, opposite or alternate Foliage somewhat spongy, greyish or bluish-grey, glaucous, erect or spreading; basal leaves long-petiolate, 8 – 20 cm long; petiole 5 – 15 cm long, glabrous; base broadened, sheathing; lamina 3-pinnatisect, oblong or obovate in outline, 3 – 5 × 2 – 5 cm, 3 – 7-jugate with terminal pinna; pinnae ovate to obovate-cuneate in outline, opposite to alternate, petiolulate below, sessile above, 4.5 – 15 × 3.5 – 20 mm, pinnatisect; ultimate segments oblanceolate to oblong-oblanceolate or obovate, 2 – 10 × 1.5 – 4 mm, apex acute to obtuse, obtusely mucronate, margins entire, surfaces glaucous
    Flowers 15 – 18 mm long, brownish-yellow or yellow; outer petal yellow with green hood; lateral wings yellow; spur yellow; lower petal colour similar to upper; inner petals yellow or pale white, green-tipped with pale yellow wings
    Inflorescence terminal, simple to compound, 7 – 15 (– 27)-flowered, 2 – 6.5 cm long corymbose racemes, slightly elongated, lax after anthesis, ± equalling the foliage, raceme branches spreading; lateral corymb(s) 2 – 6 (– 8)-flowered, on 1.5 – 5 cm long peduncle; bracts foliaceous, 2 – 3-pinnatisect, sessile or subsessile, up to 5 cm long, up to 3.2 cm broad; bracteoles foliaceous, flabellately segmented, up to 2 cm long and broad, smaller, less segmented upwards; pedicels glabrous or glaucous, equal or shorter than bracteoles; fruiting pedicels apically hooked, lower often becoming longer than bracteoles
    Sepals 2 (a pair of opposite sepals), one on either side, ovate to cordate, 1.5 – 2 × 1.75 – 2.2 mm, white, membranous, with fimbriate-dentate margins
    Lower petal 8 – 11 mm long, spreading from the base or not, but not deflexed in the middle; base 4 – 6 mm long, not saccate; lamina 3 – 5.2 mm long, apex subacute to obtuse, concave in the middle with spreading lateral wings, dorsally wingless/not crestate Inner petals 2, flat, cohering at the tip, covering stamens and carpel, 8 – 10 mm long; lamina 4 – 5 mm long, 1.8 – 2.2 mm broad, dorsally winged, inner petal apex green-tipped, lamina base clawed with 0.3 – 1 mm long curved or hooked claw; inner petal base narrowed Upper petal 15 – 18 mm long; lamina 10 – 12 mm long, 2 – 2.5 mm broad in the middle, dorsally wingless or non crestate, apex obtuse or subacute, basely spurred; spur 5 – 6 mm long (nearly half as long as the lamina of upper petal or nearly ⅓ as long as flower), straight or bending slightly downward, moderately broad, sometimes slightly narrowed towards the tip, apex obtuse, base nectariferous; nectary internal, linear, slightly shorter than the spur, fused with the lower wall of the spur for ⅘ the the length of nectary with 0.7 – 1.2 mm long, free, distal part with clavate head located 1 – 2 mm behind the tip of spur
    Stamens 6, in 2 bundles, each bundle with 1 dithecous and 2 monothecous anthers; phalange forming white, membranous covering around carpel; anthers yellow, oblong, 0.5 – 1 mm long
    Carpels 2; ovary linear-elliptic to oblanceolate-elliptic, flat, 3.5 – 4.2 mm long, glabrous; style 3 – 4 mm long, glabrous upcurved at almost right angles; stigma dilated, flattened, quadrate to sub orbicular-ovate in outline, 0.5 – 1 × 0.7 – 1.2 mm, with terminal, lateral and basal papillate lobes showing confluent pappillate zones; fruiting stigma without pappillae
    Capsule ellipsoid or elliptic-obovoid, green turning brown on drying, 6 – 10 (– 12) mm long, 2.6 – 4 mm broad, glaucous, with persistent style and stigma; seeds 3 – 5 (– 6), subbiseriate, black, lenticular, 1.5 – 2 mm wide, glossy, base with attached caruncle.
    India: Western Himalaya. Probably endemic.
    Locally common, amidst boulders on open slope; alt. 3600 – 5000 m.
    Flowering July – September; fruiting July (late) – September (late).
    Data Deficient (DD). It is suggested that, considering the very small distributional range of the species in the northern trans-Himalayan regions in Chamoli district of Uttarakhand state in India, it should be treated as vulnerable, pending discovery of additional localities.
    The species is named as a token of respect to a teacher of the author and the Additional Director, Botanical Survey of India, Kolkata, Dr Devendra Kumar Singh. Corydalis devendrae, with its characters of tuberous roots, yellow flowers and dorsally non- crestate/wingless upper and lower petals is impossible to confuse with any other species. No other species in sect. Latiflorae has this set of diagnostic characters. Of sect. Latiflorae, C. nana appears to be the most similar to C. devendrae, as they grow in the same type of habitat (amidst boulders and scree), have similar habit, greyish or bluish-grey foliage and yellow flowers, but both are easily distinguished by the characters of the roots and petals, along with other characters as given in the keys. Comparison of important distinguishing characters of these species is given in Table 1.


    Native to:

    West Himalaya

    Other Data

    Corydalis devendrae Pusalkar appears in other Kew resources:


    First published in Kew Bull. 66: 552 (2012)


    Kew Bulletin

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    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2020. Published on the Internet at and
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families.