1. Family: Apiaceae Lindl.
    1. Genus: Heracleum L.
      1. Heracleum elgonense (H.Wolff) Bullock

        This species is accepted, and its native range is Ethiopia to E. Tropical African Mountains.

    [FTEA]

    Umbelliferae, C.C. Townsend. Flora of Tropical East Africa. 1989

    Habit
    Juicy perennial, 0.6–1.8 m. tall, with stout but easily broken, fistular, sulcate-striate, densely finely pilose stems; stems 1.25–1.8 cm. wide below, simple or with 1–2 ascending branches with very reduced leaves.
    Leaves
    Basal leaves several in a loose rosette, narrowly oblong in outline, ± 35–60 × 8–13 cm., simply pinnate with usually 6–7 pairs of roundish or bluntly deltoid, mostly ± 5 × 5–7 × 7 cm., irregularly pinnatifid leaflets (the upper often overlapping), the ultimate teeth obtuse, mucronate; indumentum ± densely pilose over the entire surface and especially the lower surface of the veins; sheaths tapering, broad, striate, ± 7.5–14 × 3_4 cm.; petiole ± 12–17 cm.; median stem leaves shortly petiolate or sessile on the sheaths, similar but usually with fewer and smaller pinnae at shorter distances apart; sheaths very broad, clasping the stem, ± 4.5–9 × 2.5–4.5 cm.; uppermost leaves commonly with the leaflets more congested, longer and narrower, with sharper teeth, these sheaths also broad and inflated.
    Inflorescences
    Umbels several, on sulcate peduncles 3–24 cm. long; rays 12–20, sulcate, densely pilose, 1.5–6.5 cm.; involucre absent or of 1–few linear to broadly oblong bracts up to ± 1 cm. long; partial umbels ± 20–26-flowered, pedicels 3–15 mm., densely furnished with rather shining hairs of diverse lengths; involucel of ± 4–14 linear bracteoles 6–13 mm. long, at least some exceeding the pedicels in young fruit.
    Calyx
    Calyx-teeth narrowly to broadly triangular, ± 0.5–1 mm., early-deciduous.
    Flowers
    Flowers white, the outermost petals of the outer partial umbels very strongly radiate, deeply cordate, (7–)8–ll mm.; inner petals elliptic-oblong, ± 2 mm., with an incurved tip.
    Fruits
    Fruit broadly elliptic, ± 6–10 × 3.5–6 mm., pilose especially on the margins; stylopodia conical, exceeding the apical notch of the fruit; styles arcuate-recurved, slender, ± 2 mm.; vittae 4 dorsally, 2 on the commissure, all reaching 3/4 of the way to the base.
    Figures
    Fig. 41/1–6.
    Habitat
    From shade of Hagenia-Rapania woodland to open moorland and tree Senecio communities, frequently in damp grassy places by streams; 2820–3790 m.
    Distribution
    K3 K5 U3 known only on Elgon
    [FTEA]

    Umbelliferae, C.C. Townsend. Flora of Tropical East Africa. 1989

    Habit
    Juicy perennial, 0.15–0.9 m. tall, with a stout rootstock; stems stout but easily broken, fistular, sulcate-striate, densely finely pilose, 0.6–1.5 cm. wide below, simple or with a single ascending branch with one very reduced leaf.
    Leaves
    Basal leaves several in a rosette, narrowly oblong in outline, ± 15–40 × 2.5–15 cm., simply pinnate with usually 4–8 pairs of roundish to broadly, bluntly or acutely deltoid, 1–8.5 × 1.25–6 cm., dentate-incised or pinnatifid leaflets (the upper frequently overlapping), the ultimate teeth obtuse to acute, mucronate; indumentum ± dense along the primary and secondary nerves and margins, or sometimes more generally pilose on the upper surface; sheaths broad, striate, 5–14 × 2–3 cm., tapering or rounded above; petiole ± 5–10 cm.; median stem leaves shortly petiolate or sessile on the sheaths, similar but gradually reducing in size and number of pinnae; sheaths very broad, clasping the stem, ± 5–10 × 2–3 cm.; uppermost leaves commonly shorter and more condensed, leaflets similar but more sharply toothed, or if narrow then entire or remotely dentate, the sheaths of these also broad and inflated.
    Inflorescences
    Umbels several, on sulcate peduncles 6–18 cm. long; rays ± 9–15, sulcate, densely pilose, 1–4 cm.; involucre of 1–9 narrow or broad and blunt bracts up to 1 cm. long; partial umbels ± 10–20-flowered, pedicels 3–10 mm., densely furnished with rather shining hairs of different lengths; involucel of ± 7–10 linear bracteoles 3–8 mm. in length, at least some exceeding the pedicels in young fruit.
    Calyx
    Calyx-teeth broadly triangular, ± 0.5 mm., early-deciduous.
    Flowers
    Flowers white, the outermost petals of the outer partial umbels radiate, deeply cordate, (3–)4–6.5(–8 in Ethiopia) mm.; inner petals elliptic-oblong, ± 1.25–1.5 mm., with an incurved tip.
    Fruits
    Fruit broadly elliptic, ± 6–12 × 4.5–6 mm., sparsely pilose over the entire surface or on the margins only; stylopodia shortly conical, not exceeding the apical notch of the fruit; styles deflexed, ± 1–1.5 mm.; vittae 4 dorsally, 2 on the commissure, all reaching ± 3/4 of the way to the base or the commissural short and less than 1/3 from the apex.
    Habitat
    A mountain species occurring on boggy moorland, in Senecio groves, on loamy soil at the foot of rock outcrops and in moorland on a rocky peak; 3360–4130 m.
    Distribution
    K4 T2

    Distribution

    Native to:

    Ethiopia, Kenya, Sudan, Tanzania, Uganda

    Heracleum elgonense (H.Wolff) Bullock appears in other Kew resources:

    First published in Bull. Misc. Inform. Kew 1932: 496 (1932)

    Accepted by

    • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
    • Van Wyk, B.-E., Tilney, P.M. & Magee, A.R. (2013). African Apiaceae: a synopsis of the Apiaceae/Umbelliferae of Sub-Saharan Africa and Madagascar: 1- 317. Briza Academic Books, Pretoria.
    • Hedberg, I., Edwards, S. & Nemomissa, S. (eds.) (2003). Flora of Ethiopia and Eritrea 4(1): 1-352. The National Herbarium, Addis Ababa University, Ethiopia & The Department of Systematic Botany, Upps.
    • Townsend, C.C. (1989). Flora of Tropical East Africa, Umbelliferae: 1-127.

    Literature

    Kew Backbone Distributions
    • Van Wyk, B.-E., Tilney, P.M. & Magee, A.R. (2013). African Apiaceae: a synopsis of the Apiaceae/Umbelliferae of Sub-Saharan Africa and Madagascar: 1- 317. Briza Academic Books, Pretoria.
    • Townsend, C.C. (1989). Flora of Tropical East Africa, Umbelliferae: 1-127.
    Flora of Tropical East Africa
    • A.D.Q. Agnew, Upland Kenya Wild Flowers p. 361 (1974).
    • F. W. Andr., The Flowering Plants of the Anglo-Egyptian Sudan 2: 362 (1952).
    • Bullock in Bulletin of Miscellaneous Information, Kew 1932: 496 (1932).

    Sources

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2020. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2020. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0