1. Family: Lamiaceae Martinov
    1. Genus: Vitex L.
      1. Vitex amaniensis W.Piep.

        This species is accepted, and its native range is Tanzania to Zimbabwe.

    [FZ]

    Flora Zambesiaca. Vol 8, Pt 7. Avicenniaceae, R. Fernandes. Nesogenaceae, M.A. Diniz. Verbenaceae, R. Fernandes. Lamiaceae, R. Fernandes. 2005.

    Type
    Type from Tanzania.
    Habit
    Tree c. 15–18 m tall; bark white or pale brown, smooth; older branches dark red-brown; young branches crisply pilose Tree c. 15–18 m tall; bark white or pale brown, smooth; older branches dark red-brown; young branches crisply pilose.
    Leaves
    Leaves drying dark, 3–5-foliolate; petioles 9–15 cm long, terete; median petiolules 0.5–2.5 cm long; leaflets 10–20 × 5–6.5 cm, median leaflet largest, elliptic to broad obovate, rounded or apiculate to shortly acuminate at the apex, cuneate to ± attenuate at the base, entire to remotely serrulate, glabrous on both surfaces with a few short hairs on the nerves beneath, nerves ± raised on both surfaces Leaves drying dark, 3–5-foliolate; petioles 9–15 cm long, terete; median petiolules 0.5–2.5 cm long; leaflets 10–20 × 5–6.5 cm, median leaflet largest, elliptic to broad obovate, rounded or apiculate to shortly acuminate at the apex, cuneate to ± attenuate at the base, entire to remotely serrulate, glabrous on both surfaces with a few short hairs on the nerves beneath, nerves ± raised on both surfaces.
    Ovary
    Ovary obovoid, sparsely gland-dotted at apex only, glabrous, truncate at apex; style 5 mm long
    Ecology
    In the Flora Zambesiaca area known only from the evergreen rain forest at Chirinda in Zimbabwe; c. 1100 m.
    Note
    Only four gatherings of this species from the Flora Zambesiaca region have been seen by this author – all from the Chirinda Forest in Zimbabwe.  This rain forest species was apparently first collected there in 1948 (by N.C. Chase 435, Jan 1948, Chirinda Forest) and wrongly determined as V. lokundjensis W. Piep. by Moldenke (loc. cit. 1956).  V. amaniensis and V. lokundjensis show some superficial similarities (both are tall rain forest trees with leaves drying dark, leaflets petiolulate (to 2.5 cm long) and flowers in lax few-flowered axillary compound dichasia).  However, this mainly Cameroon species differs in many characters from our plant: in the terete petiole, longer bracteoles, lanate calyx, glabrous and gland-dotted ovary and the eglandular hairs on filaments.  Our plant shows great uniformity and differs from many of the Tanzanian plants in the non-gland-dotted leaves, obsolete calyx teeth and median lower lobe of the corolla larger and yellow.  Some Tanzanian specimens (e.g. Ulugurue, Tanana, Morogoro, E.M. Bruce 774 (K)) approach our plants, being clearly intermediates of a range of variation.  Although there is a major disjunction between the Tanzanian and the Zimbabwean plants, both localities occur in similar montane habitats with a similar floristic composition.
    Distribution
    Zimbabwe District code: ZIM E, ZIM B, ZIM E, ZIM S, MOZ N, MOZ Z, MOZ T, MOZ MS, MOZ GI, MOZ M. Mozambique Also in Tanzania. Zambia
    Inflorescences
    Inflorescences axillary, 7–11 cm long, of ± slender, few-flowered compound dichasia, shortly crisped-pubescent; bracteoles 0.4–0.5 cm long, linear. Inflorescences axillary, 7–11 cm long, of ± slender, few-flowered compound dichasia, shortly crisped-pubescent; bracteoles 0.4–0.5 cm long, linear
    Calyx
    Calyx obconical, 5-toothed, not gland-dotted, densely pubescent, accrescent in fruit; tube c. 2 mm long; teeth c. 0.3 mm, erect. Calyx obconical, 5-toothed, not gland-dotted, densely pubescent, accrescent in fruit; tube c. 2 mm long; teeth c. 0.3 mm, erect
    Corolla
    Corolla pale yellow, 10–11 mm long, with scattered gland-dots, densely pubescent; tube c. 5 mm long, straight; lobes at obtuse angle to tube, middle lower lobe 5.5 × 4.5 cm. Corolla pale yellow, 10–11 mm long, with scattered gland-dots, densely pubescent; tube c. 5 mm long, straight; lobes at obtuse angle to tube, middle lower lobe 5.5 × 4.5 cm
    Stamens
    Stamens scarcely exserted from tube; filaments pubescent at base, hairs eglandular. Stamens scarcely exserted from tube; filaments pubescent at base, hairs eglandular
    Pistil
    Ovary obovoid, sparsely gland-dotted at apex only, glabrous, truncate at apex; style 5 mm long.
    Fruits
    Drupe c. 1.4 × 1 cm, obovoid (?immature), rounded at apex, ± enclosed in the enlarged calyx; fruiting calyx cup-shaped, pubescent, not gland-dotted. Drupe c. 1.4 × 1 cm, obovoid (?immature), rounded at apex, ± enclosed in the enlarged calyx; fruiting calyx cup-shaped, pubescent, not gland-dotted.
    [FTEA]

    Verbenaceae, B. Verdcourt. Flora of Tropical East Africa. 1992

    Habit
    Tree 12–20 m. tall, with smooth white bark or pale grey-brown ± corky bark on twigs; branchlets crisply pilose.
    Leaves
    Leaves drying dark, 5-foliolate; leaflets elliptic to obovate-elliptic, 6.5–18 cm. long, 3.2–9(–11) cm. wide, acuminate, cuneate at the base, glabrous above, pubescent beneath only on the nerves which are rather close and parallel, glandular-punctate, entire to remotely serrulate with upwardly directed teeth; petioles 6–12 cm. long; petiolules 0.5–2 cm. long.
    Inflorescences
    Inflorescences axillary, 8–25 cm. long, ± few-flowered dichasial cymes, the axes ferruginous tomentose or shortly crisped-pilose; bracteoles linear, 0.5–1 cm. long; peduncles 3–11.5(–13) cm. long; pedicels of central flowers up to 5 mm. long but others much shorter.
    Calyx
    Calyx campanulate, 3–4 mm. long, somewhat 2-lipped, densely grey crisped-hairy, irregularly toothed, the teeth ± 1 mm. long.
    Corolla
    Corolla lilac-white or pale blue with deeper blue lips and mauve lines; tube 3.5 mm. long, ferruginous tomentose outside; larger lip 3-fid, the main lobe 2.75 × 3 mm., the others much smaller; smaller lip with 3 ovate-triangular lobes 2.2 × 1–2 mm.
    Ovary
    Ovary glandular at apex.
    Fruits
    Fruit black, ellipsoid-oblong, 0.9–1.1 cm. long, 6.5 mm. wide, glabrous; calycine cup 0.7–1 cm. wide, pale brown pubescent.
    Habitat
    Rain-forest, forest edges and relicts; 900–1400 m.
    Distribution
    not known elsewhere T3 T6 T7

    Images

    Distribution

    Native to:

    Tanzania, Zimbabwe

    Vitex amaniensis W.Piep. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Braun [1054], Tanzania K000192778

    First published in Beibl. Bot. Jahrb. Syst. 141: 60 (1928)

    Accepted by

    • Fernandes, R. & Diniz, M.A. (2005). Avicenniaceae, Nesogenaceae, Verbenaceae and Lamiaceae (subfams, Viticoideae and Ajugoideae) Flora Zambesiaca 8(7): 1-161. Royal Botanic Gardens, Kew.
    • Govaerts, R. (2003). World Checklist of Selected Plant Families Database in ACCESS: 1-216203. The Board of Trustees of the Royal Botanic Gardens, Kew.

    Literature

    Flora Zambesiaca
    • F.T.E.A., Verbenaceae: 58, 66 (1992).
    • Brenan, Check-list For. Trees Shrubs Tang. Terr: 643 (1949).
    Kew Backbone Distributions
    • Fernandes, R. & Diniz, M.A. (2005). Avicenniaceae, Nesogenaceae, Verbenaceae and Lamiaceae (subfams, Viticoideae and Ajugoideae) Flora Zambesiaca 8(7): 1-161. Royal Botanic Gardens, Kew.
    Flora of Tropical East Africa
    • J.P.M. Brenan, Check-lists of the Forest Trees and Shrubs of the British Empire no. 5, part II, Tanganyika Territory p. 643 (1949).
    • Pieper in F. Fedde, Repertorium Speciorum Novarum Regni Vegetabilis 26: 164 (1929).
    • Pieper in A. Engler, Botanische Jahrbücher für Systematik, Pflanzengeschichte und Pflanzengeographie 62, Beibl. 141: 46, 60 (1928) (in key)

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0